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Hymenasplenium unilaterale

(Lam.) Hayata (1927)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Aspleniaceae Genus: Hymenasplenium

pāmoho [pamoho]

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Extirpated
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Bot. Mag. (Tokyo) 41: 712 (1927)

Other References

Palmer 2003:58 (K, O, Mo, L, M, H [as Asplenium excisum]) 74 (K, O, Mo, L, M, H [as Asplenium unilaterale]); Pteridophyte Phylogeny Group 2016:582 (Syn. A. unilaterale = Hymenasplenium unilaterale); Palmer & Thomas in press (A. excisum misapplied = Hymenasplenium unilaterale); Degener 30155 BISH (L†, 1963)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Hymenasplenium unilaterale Mountains near Koele Preserved_Specimen Forbes, C.N. 26 Lanai BISH 1913-06-01
2 Hymenasplenium unilaterale unknown Preserved_Specimen 115 BISH
3 Hymenasplenium unilaterale unknown Preserved_Specimen Hillebrand, W.B. BISH
4 Hymenasplenium unilaterale East Maui, Lower Shelf of Kipahulu Valley above Palikea Camp Preserved_Specimen Welton, P. 2005 Maui BISH 1997-10-07
5 Hymenasplenium unilaterale Punaluu Mt Preserved_Specimen Rock, J.F.C. 15 Oahu BISH 1908-08-01
6 Hymenasplenium unilaterale Glenwood. Preserved_Specimen Lyon, H.L. 1099 Hawaii BISH 1908-03-20
7 Hymenasplenium unilaterale Mt. Kaala, E. side Preserved_Specimen Forbes, C.N. Oahu BISH 1908-08-06
8 Hymenasplenium unilaterale East Maui, Upper Shelf of Kipahulu Valley, off of Central Pali trail in stream bed below waterfall Preserved_Specimen Welton, P. 2390 Maui BISH 2017-06-15
9 Hymenasplenium unilaterale W Maui, Hanaula, upper Pohakea Ravine Preserved_Specimen Hobdy, R.W. 243 Maui BISH 1971-06-26
10 Hymenasplenium unilaterale Kilauea; N.E. Ledge 19.4N 155.3W Preserved_Specimen Krajina, V.J. 611226084 Hawaii BISH 1961-12-26
11 Hymenasplenium unilaterale Akaka Falls, Honomu Preserved_Specimen Forbes, C.N. 623 Hawaii BISH 1915-05-03
12 Hymenasplenium unilaterale Palolo. Preserved_Specimen Lyon, H.L. 1029 Oahu BISH 1907-11-24
13 Hymenasplenium unilaterale head of Kahana Valley, Hauula FR Preserved_Specimen St.John, H. 13413 Oahu BISH 1933-12-10
14 Hymenasplenium unilaterale R fork Wailupe Valley Preserved_Specimen Garber, D.W. 156 Oahu BISH 1920-06-12
15 Hymenasplenium unilaterale East Maui, Nahiku. Preserved_Specimen Lyon, H.L. 1029 Maui BISH 1909-01-15
16 Hymenasplenium unilaterale East Maui, Upper Shelf Kipahulu Valley Central Pali Trail Preserved_Specimen Welton, P. 1777001 Maui BISH 1993-10-19
17 Hymenasplenium unilaterale Kaluaaha Valley Preserved_Specimen Forbes, C.N. 376 Molokai BISH 1912-08-01
18 Hymenasplenium unilaterale Kaholuamanu, behind Waimea Preserved_Specimen Forbes, C.N. 439 Kauai BISH 1909-09-01
19 Hymenasplenium unilaterale Sacred Falls Preserved_Specimen Topping, D.L. 2758 Oahu BISH 1924-03-16
20 Hymenasplenium unilaterale Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2844 Kauai BISH 1895-09-10
21 Hymenasplenium unilaterale Kaluaaha Valley Preserved_Specimen Forbes, C.N. 376 Molokai BISH 1912-08-01
22 Hymenasplenium unilaterale Kaiholena Preserved_Specimen Munro, G.C. 1 Lanai BISH 1913-01-01
23 Hymenasplenium unilaterale Makiki Preserved_Specimen Mann, H. 168 Oahu BISH 1864-01-01
24 Hymenasplenium unilaterale R fork Niu Valley Preserved_Specimen Garber, D.W. 457 Oahu BISH 1920-06-29
25 Hymenasplenium unilaterale Pahole Gulch; drainage #1 Preserved_Specimen Welton, P. 587 Oahu BISH 1990-10-06
26 Hymenasplenium unilaterale Waioli Valley Preserved_Specimen Forbes, C.N. 94 Kauai BISH 1909-07-29
27 Hymenasplenium unilaterale Kaiholena Preserved_Specimen Munro, G.C. 178 Lanai BISH 1913-01-01
28 Hymenasplenium unilaterale nw of Olaa, Hiulani forest Preserved_Specimen Skottsberg, C.J.F. 443 Hawaii BISH 1922-09-08
29 Hymenasplenium unilaterale Kaholuamanoa, above Waimea Preserved_Specimen Forbes, C.N. 439 Kauai BISH 1909-09-01
30 Hymenasplenium unilaterale Waialae stream near Ranger Station Preserved_Specimen Cranwell, L.M. 3080 Kauai BISH 1938-08-22
31 Hymenasplenium unilaterale Fire Break Trail Preserved_Specimen Topping, D.L. 2741 Oahu BISH 1924-01-27
32 Hymenasplenium unilaterale Kalae Preserved_Specimen Forbes, C.N. 38 Molokai BISH 1912-06-01
33 Hymenasplenium unilaterale Waianae Mts., Makaleha Valley, ridge up Kaala Preserved_Specimen Meebold, A. 21330 Oahu BISH 1935-12-01
34 Hymenasplenium unilaterale Haupu Range, left side of Kipu Kai Gap Preserved_Specimen Forbes, C.N. 729 Kauai BISH 1916-11-01
35 Hymenasplenium unilaterale Waime Drainage Basin, west side Preserved_Specimen Forbes, C.N. 937 Kauai BISH 1917-07-03
36 Hymenasplenium unilaterale Niu Valley. Preserved_Specimen Lyon, H.L. Oahu BISH 1909-07-11
37 Hymenasplenium unilaterale East of Puu Kolekole on Bishop Estate land. Preserved_Specimen Corn, C. Molokai BISH 1983-09-13
38 Hymenasplenium unilaterale Kauai. Preserved_Specimen Brodie, A. Kauai BISH 1904-10-03
39 Hymenasplenium unilaterale Koolau Mts. between Punaluu and Kaipapau Preserved_Specimen Forbes, C.N. Oahu BISH 1908-11-14
40 Hymenasplenium unilaterale Kokee Preserved_Specimen Smith, F.G. Kauai BISH 1929-05-01
41 Hymenasplenium unilaterale Niu Valley. Preserved_Specimen Lyon, H.L. Oahu BISH 1909-07-11
42 Hymenasplenium unilaterale East Maui, upper trail, bank of stream. Preserved_Specimen Lyon, H.L. Maui BISH 1908-12-11
43 Hymenasplenium unilaterale Upper Hamakua Ditch Trail. Preserved_Specimen Lyon, H.L. Hawaii BISH 1919-07-07
44 Hymenasplenium unilaterale Hapapa, Waianae Preserved_Specimen Russ, G.W. Oahu BISH 1930-05-01
45 Hymenasplenium unilaterale Koiawe branch, Waipio Valley, upper ditch trail. Preserved_Specimen Lyon, H.L. Hawaii BISH 1919-09-03
46 Hymenasplenium unilaterale Kauai. Preserved_Specimen Brodie, A. Kauai BISH 1904-07-17
47 Hymenasplenium unilaterale Kilauea, Thurston lava tube Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
48 Hymenasplenium unilaterale Oahu Preserved_Specimen Whitney, J.M. Oahu BISH
49 Hymenasplenium unilaterale Hapapa, Waianae Preserved_Specimen Russ, G.W. Oahu BISH 1930-05-01
50 Hymenasplenium unilaterale East Maui, manawainui Planeze, trail to cable ridge from kekuewa Preserved_Specimen Welton, P. Maui BISH 2003-08-26
51 Hymenasplenium unilaterale East Maui, Upper Shelf Kipahulu Valley Central Pali Trail Central Pali Trail near Charlie Camp. Preserved_Specimen Haus, B. Maui BISH 1993-09-29
52 Hymenasplenium unilaterale ridge W of Kalihi Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1908-10-03
53 Hymenasplenium unilaterale East maui, lower shelf of Kipahulu near Palikea Camp Preserved_Specimen Welton, P. Maui BISH 2000-07-13
54 Hymenasplenium unilaterale Lanai Preserved_Specimen Munro, G.C. Lanai BISH 1915-01-01
55 Hymenasplenium unilaterale Kilauea, in Twin Crater Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-01
56 Hymenasplenium unilaterale Wahiawa gulch Preserved_Specimen Rock, J.F.C. Oahu BISH 1908-07-01
57 Hymenasplenium unilaterale East Maui, Kula pipeline. Preserved_Specimen Lyon, H.L. Maui BISH 1926-10-21
58 Hymenasplenium unilaterale Kauai mt Preserved_Specimen Rock, J.F.C. Kauai BISH 1908-08-01
59 Hymenasplenium unilaterale Manoa, Aihualama. Preserved_Specimen Lyon, H.L. Oahu BISH 1926-10-03
60 Hymenasplenium unilaterale unknown Preserved_Specimen Judd, J.R. 76 BISH 1988-01-01
61 Hymenasplenium unilaterale Palikea in side gully. Waianae Mountains In gulch with Cryptomeria japonica, Metrosideros polymorpha, Blechnum appendiculatum, Diplazium sanwichianum, Ceodes brunoniana, Microlepia strigosa, Athyrium microphyllum, Dryopteris fuscoatra, Vesicularia perviridis, Racopilum cuspidigerum, Distichophyllum freycinetii, and various outplanted natives. Preserved_Specimen Blades large on long stipes. Plants terrestrial. Rhizomes more or less thick with short creeping rhizomes. Thomas, M.K. 128 Oahu BISH 2021-09-15
62 Hymenasplenium unilaterale Oahu or Hawaii (Kona or Kilauea) Preserved_Specimen Lack, F. BISH 1880-01-01
63 Hymenasplenium unilaterale unknown Preserved_Specimen U.S. Exploring Expedition BISH 1838-01-01
64 Hymenasplenium unilaterale Hillebrands Glen Preserved_Specimen Forbes, C.N. Oahu BISH
65 Hymenasplenium unilaterale Honokahau Drainage Basin Preserved_Specimen Forbes, C.N. 402 Maui BISH 1917-09-25
66 Hymenasplenium unilaterale Kula Pipe Line, Waikamoi Preserved_Specimen Forbes, C.N. 1267 Maui BISH 1919-09-05
67 Hymenasplenium unilaterale Kapohaku Preserved_Specimen Munro, G.C. 189 Lanai BISH 1927-08-01
68 Hymenasplenium unilaterale Palolo. Preserved_Specimen Lyon, H.L. 1007 Oahu BISH 1907-11-17
69 Hymenasplenium unilaterale Slopes of Puu Hapapa. Preserved_Specimen Topping, D.L. 3356 Oahu BISH 1930-03-16
70 Hymenasplenium unilaterale unknown Preserved_Specimen BISH
71 Hymenasplenium unilaterale Kula Pipe Line N walls on sides of streams Preserved_Specimen creeping Munro, G.C. 755 Maui BISH 1927-11-25
72 Hymenasplenium unilaterale East rim of Ukumehame Gulch Wet Metrosideros forest Preserved_Specimen St.John, H. 25719 Maui BISH 1955-12-28
73 Hymenasplenium unilaterale Nahiku mossy wooded bank of stream Preserved_Specimen St.John, H. 17957 Maui BISH 1937-01-02
74 Hymenasplenium unilaterale W Maui, Iao Valley on a steep ridge on the east side Unusual habitat on a ridgetop in rich soil under a dense canopy of Aleurites and Pisonia. Preserved_Specimen Creeping rootstock, fronds erect to 0.6m tall, growing alone on bare soil. Hobdy, R.W. 1861 Maui BISH 1983-10-15
75 Hymenasplenium unilaterale Waipio, Kipapa Gulch, second forth fork. In moist gully. Preserved_Specimen Hosaka, E.Y. 1030 Oahu BISH 1933-05-14
76 Hymenasplenium unilaterale Waimea District. Waialae Stream and side streamlets. Metrosideros-Cheirodendron montane wet forest with Dicranopteris-Diplopterygium-Sticherus matting ferns with Leptecophylla tameiameiae, Vaccinium calycinum, V. dentatum, Broussaisia arguta, Kadua affinis, Peperomia hesperomannii, Gahnia vitiensis kauaiensis, G. beecheyi, Melicope clusiifolia, M. kauaiensi Preserved_Specimen Fern, terrestrial along stream bank, rhizome long-creeping, fertile, common Wood, K.R. 18271 Kauai BISH 2019-08-07
77 Hymenasplenium unilaterale East Maui, Olinda, Kula flume line stream gorge, mossy rocks Preserved_Specimen St.John, H. 10328 Maui BISH 1930-02-12
78 Hymenasplenium unilaterale Waimea District. Upper headwaters of Mohihi south of Waiakoali flats, Na Pali Kona Forest Reserve. on saturated stream walls. Metrosideros-Cheirodendron montane wet forest w/ Syzygium sandwicensis, Kadua affinis, Pittosporum gayanum, Ilex anomala, Melicope anisata, M. clusiifolia, Myrsine dentata, M. punctata, Psychotria hexandra, P. mariniana, Coprosma kauaiensis, Vacc Preserved_Specimen Epilithic fern, rhizome long creeping, few scales, apparently rheophytic, fertile, common. Wood, K.R. 17936 Kauai BISH 2018-09-20
79 Hymenasplenium unilaterale Kilauea Lava tube. Preserved_Specimen Fullaway, D. 1720 Hawaii BISH
80 Hymenasplenium unilaterale Kokee; end of Camp 10 Road, on trail to Circle Bog, but drop down North into tributary of Waiakoali, along steam on bank Metrosideros-Dicranopteris-Cheirodendron Wet Forest Psychotria, Coprosma, Saddleria, Broussaisia arguta, Elaphoglossum crassicaule, Vaccinium, Cyrtandra, Pittosporum, Platydesma, Peperomia kokeana, Seaginella, Deparia, Dryopteris, Callistopteris, Conio Preserved_Specimen Common on wet walls and streamside, teresstrial, upright, stipes black and shinny w/ ridges. pinna turned out from base, thin, no proliferations Tangalin, N. 2435 Kauai BISH 2010-12-08
81 Hymenasplenium unilaterale Wahiawa District. Summit of Mt. Kaala, northeast facing slopes. Lat/Long: 21.508319855, -158.1438446. Metrosideros wet forest with Cheirodendron, Broussaisia arguta, Diplazium sandwichianum, Mysine lessertiana, Cibotium, Antidesma platyphylla, Rubus argutus, Pittosporum, Freycinetia arborea, Ilex anomala, Melicope clusifoli. Preserved_Specimen Fern on waterfall face, 6-12 inches hight, fertile. Perlman, S. 23234 Oahu BISH 2013-01-16
82 Hymenasplenium unilaterale 10.5 mi up Kulani Road, Waiakea, S. Hilo moist shady ground in open lava tube Preserved_Specimen St.John, H. 22334 Hawaii BISH 1946-12-24
83 Hymenasplenium unilaterale East Maui, Hana District, Haleakala, Kipahulu Valley closed Acacia koa and Metrosideros forest Preserved_Specimen Higashino, P.K. 10031 Maui BISH 1983-07-17
84 Hymenasplenium unilaterale Waimea Forest. Preserved_Specimen Lyons, A.B. Hawaii BISH 1860-01-01
85 Hymenasplenium unilaterale Waimea District. Koaie headwaters near Koaie camp. along stream-banks. Metrosideros-Cheirodendron-Dicranopteris mixed wet forest w/ Melicope clusiifolia, Coprosma kauensis, Polyscias waialealae, Broussaisia arguta, Vaccinium calycinum. V. dentatum, Syzygium sandwicensis, Astrelia argyrocoma, Leptecophylla tameiameiae, Smilax mel Preserved_Specimen Terrestrial fern, rhizome creeping, occasional. Wood, K.R. 18129 Kauai BISH 2019-03-07
86 Hymenasplenium unilaterale Kaala summit in gully above FAA road near last hairpin turn. On small waterfall in native wet forest. Growing with Metrosideros polymorpha, Melicope clusiifolia, Melicope christophersenii, Schiedea trinerve, Vaccinium calycinum, Anoectochilus sandvicensis, Cheirodendron platyphyllum, Cheirodendron trigynum, Asplenium contiguum, Aspenium acuminatum, Phylostegia grandiflora, Cibotium menziesii, Cibotium chamissoi, Sadleria pallida, Dicranopteris linearis, and many others. Preserved_Specimen Lanceolate fronds with translucent pinnae. Rhizome mostly short creeping, dorsiventral. Pinnae mostly dimidiate. Thomas, M.K. 473 Oahu BISH 2022-10-27
87 Hymenasplenium unilaterale Waimea Canyon State Park; mile 11.3 on Hwy 550 Waimea Canyon in remnant forest on steep, dry slope. In shaded ravine with Pteris excelsa, P. xhillebrandii, Microlepia strigosa and Lantana camara Preserved_Specimen Flynn, T.W. 2211 Kauai BISH 1987-05-21
88 Hymenasplenium unilaterale Hanalei District; Namolokama Plateau On wet cliff. Preserved_Specimen Hobdy, R.W. 3369 Kauai BISH 1991-06-19
89 Hymenasplenium unilaterale Below Kamanu Ridge, headwaters of Waikoko, S of Wailua's N fork and above Wailua ditch. Metrosideros wet forest with wet diff communities. W/ Cheirodendron, Pipturus spp., Dubautia, Cyrtandra, Kadua centranthoides, K. elatior, K. foggiana, Psychotria, Melicope, Machaerina, Isachne, with ferns of Microlepia, Asplenium, Christella, Deparia. Preserved_Specimen Rhizome creeping, on wet cliff, pinnae turning distally upward, occasional. Wood, K.R. 12766 Kauai BISH 2008-01-12
90 Hymenasplenium unilaterale Mokuleia woods Preserved_Specimen Degener, O. 26874 Oahu BISH 1966-03-04
91 Hymenasplenium unilaterale Waipio, Kipapa Gulch, north ridge. Moist gully. Preserved_Specimen Hosaka, E.Y. 585 Oahu BISH 1932-05-29
92 Hymenasplenium unilaterale Kahaluu Rare in gulch streambed on rock. Associated vegetation: Syzygium mallacense, Pandanus tectorius, Cecropia obtusifolia, Psidium guajava, Aleurites moluccana, Clidemia hirta, Christella parasitica, Deparia petersenii, Lepisorus thunbergianus, Paederia foetida, Dioscorea bulbiflora, Oplismenus hirtellus Preserved_Specimen Kennedy, B. 20073 Oahu BISH 2007-04-16
93 Hymenasplenium unilaterale Kamakou Preserve; west fork of stream just east of Kawela Ridge, along upper reaches of gulch and ridge leading to Puu O Kaeha Low, mixed wet forest with Metrosideros dominant, also Ilex, Xylosma, Psychotria, and Syzygium. Preserved_Specimen Fern on wet rock walls of shady grotto by waterfall. Localised population. Lorence, D.H. 5654 Molokai BISH 1987-10-22
94 Hymenasplenium unilaterale Kalawahine trail, Koolau Mountains On cindery bank in shade in side gulch on the side of the trail. Many plants seen, growing with Dumortiera hirsuta, Distichophyllum freycinetii, Selaginella arbuscula, Adiantum raddianum, Begonia hirtella, Tectaria gaudichaudii, and Peperomia latifolia. Preserved_Specimen Lamina thin, nerly transparent. Stipes thin, rhizome thin and long creeping over rocky substrate. Thomas, M.K. 129 Oahu BISH 2021-09-17
95 Hymenasplenium unilaterale East Maui, Kailua, above upper ditch trail. Under falls on rocks. Preserved_Specimen Lyon, H.L. Maui BISH 1908-11-16
96 Hymenasplenium unilaterale Waimea Valley, ride gulch on NW side, 3/4 mile ipstream from intake Under fern in dense moist forest. Preserved_Specimen St.John, H. 13953 Kauai BISH 1934-01-01
97 Hymenasplenium unilaterale East Maui, Hana District, Kipahulu Valley NE of central ridge on rotting stump Preserved_Specimen DeWreede, R.E. 22 Maui BISH 1967-08-16
98 Hymenasplenium unilaterale East Maui, Olinda Flume, Waikamoi moist, vertical wall, side of creek, upper end Preserved_Specimen Crosby, M.R. 1791 Maui BISH 1964-07-16
99 Hymenasplenium unilaterale Waianae Mts., first gully below Peacock Flats, in small, dead-end side gully on left (as ascending) ca. 1 mi from beginning of trail Shaded, somewhat moist; associated with Urera, Cyrtandra, Planchonella Preserved_Specimen Herbst, D.R. 337 Oahu BISH 1967-02-25
100 Hymenasplenium unilaterale W Maui, Waikapu Canyon at very back in good native forest. Preserved_Specimen Plants growing terrestrially in soil. Hobdy, R.W. 2232 Maui BISH 1984-12-16
101 Hymenasplenium unilaterale East Maui, Haiku Uka, near Waikamoi Flume Road within Koolau Forest Reserve 5 m `Ohi`a Montane Wet Forest along a steep rocky streambed. Assoc. veg.: Broussaisia arguta, Pilea peploides, Peperomia sp., Carex alligata, Deschampsia nubigena, Athyrium microphyllum, Selaginella arbuscula. Preserved_Specimen Oppenheimer, H.L. 292 Maui BISH 1998-02-16
102 Hymenasplenium unilaterale Makaha Valley-Waianae Range Rain forest; on ground among Vandenboschia Preserved_Specimen Wilson, K.A. 203 Oahu BISH 1952-11-30
103 Hymenasplenium unilaterale Manuka Mauka, Kau Dist on tree trunk Preserved_Specimen St.John, H. 11285 Hawaii BISH 1931-12-24
104 Hymenasplenium unilaterale Waimea District. Napali-Kona Forest Reserve. Makaha Valley, ca. 1.3 miles west of Hwy. 550, just north of Kokio Keokeo picnic area Mixed mesophytic forest dominated by acadia and metrosideros.Terrestrial in mocerate shade.With Dryopteris unidentata, Tectaria gaudichaudii and Pteris species. Localized. Preserved_Specimen Terrestrial. Flynn, T.W. 2354 Kauai BISH 1987-08-28
105 Hymenasplenium unilaterale Waianae Mts., Makaha-Waianae Kai small cliff in wet forest Preserved_Specimen pendent Fosberg, F.R. 10858 Oahu BISH 1935-03-31
106 Hymenasplenium unilaterale Nuuanu in side gulch east of Pali lookout. Sothern Koolau mountains. Growing on talus slope on side of gully near waterfall in forest dominated by Cinnamomum, Psidium, and Citharexylum. Also growing with Touchardia latifolia, Hibiscus punaluuensis, Deparia prolifera, Crepidomanes draytonianum, and others. Preserved_Specimen Rhizome mostly short creeping. Fronds truncate, pinnae falcate-dimidate with coarse teeth. Some fronds more linear with less falcate pinnae. Thomas, M.K. 270 Oahu BISH 2022-02-21
107 Hymenasplenium unilaterale Gulch W of Borge Ridge Rd, Makawao Forest Reserve On wet shady banks near gulch bottom. Preserved_Specimen Hobdy, R.W. 1083 Maui BISH 1981-05-12
108 Hymenasplenium unilaterale Waianae range, Pahole gulch, R hand branch on rock; near Tectaria, Athyrium Preserved_Specimen Herat, T. 472 Oahu BISH 1973-07-23
109 Hymenasplenium unilaterale Manoa cliffs trail. Koolau Mountains, Oahu. On cinder deposit in shaded understory. Growing with Peperomia latifolia, Vandenboschia cyrthoteca, Tectaria gaudichaudii, Deparia petersenii, Deparia prolifera, Bischofia javanica, Cestrum nocturnum, and many bryophytes. Preserved_Specimen Thomas, M.K. 150 Oahu BISH 2021-10-25
110 Hymenasplenium unilaterale Kau, Manuka Dense shaded & damp fern forest Preserved_Specimen Terrestrial erect. Height: 18 inches Russ, G.W. 54 Hawaii BISH 1927-09-01
111 Hymenasplenium unilaterale W Maui, Hanaula Springs Trail Terrestrial in shady understory. Preserved_Specimen Hobdy, R.W. 1071 Maui BISH 1981-05-06
112 Hymenasplenium unilaterale W Maui, Iao Valley on a steep ridge on the east side Unusual habitat on a ridgetop in rich soil under a dense canopy of Aleurites and Pisonia. Preserved_Specimen Creeping rootstock, fronds erect to 0.6m tall, growing alone on bare soil. Hobdy, R.W. 1861 Maui BISH 1983-10-15
113 Hymenasplenium unilaterale Lihue District. Waiahi, upper southern headwaters. Metrosideros-Cheirodendron mixed wet forest with dissecting drainages and matting ferns of Diplopteryglum and Dicranopteris with Broussaisia arguta, Perrottettia sandwicensis, Touchardia latifolia, Pipturus albidus, P. ruber, Psychotria marinana, P. hexandra, Antidesma platyphylla, Polyscias oahuensis, Kadua affinis, Melico Preserved_Specimen Fern, epilithic on stream banks, rhizome long creeping, fronds erect, common. Wood, K.R. 17974 Kauai BISH 2018-11-15
114 Hymenasplenium unilaterale Kaala, by stream just outside NAR boundary fence, leeward side of bog reduced light Preserved_Specimen DLNR Staff Oahu BISH 1991-09-17
115 Hymenasplenium unilaterale Pauoa flats in gully near streamlet. In Psidium forest with some Metrosideros tremuloides, M. polymorpha, Acacia koa, Cibotium chamissoi, Miconia crenata, Ardisia elliptica, A. crenata, Citharexylum caudatum, Leucobryum gracile, Pyrrhobryum spiniforme, Vesicularia perveridis, and others. Preserved_Specimen Thomas, M.K. 151 Oahu BISH 2021-10-25
116 Hymenasplenium unilaterale ridge between Hanalilolilo and Pepeopae, Waikolu Valley, Kawela Moist walls of deep gorge. Preserved_Specimen St.John, H. 12553 Molokai BISH 1932-12-25
117 Hymenasplenium unilaterale East Maui, Hana, Koolau FR, Wailua Nui region, headwaters above Waiokamilo and Kano streams Metrosideros-Cheirodendron-Dicranopteris montane wet forest, with Clermontia arborea, Coprosma spp., Kadua affinis, Perrottetia, Peperomia spp., Cyrtandra spp., Myrsine spp., Pipturus albidus, with rich moss and fern understory Preserved_Specimen Terrestrial, rhizome long-creeping, rachis black brown Wood, K.R. 12596 Maui BISH 2007-10-23
118 Hymenasplenium unilaterale East Maui, above Manawainui canyon in spring fed cliff face in Healani stream, just above trail crossing Preserved_Specimen Hobdy, R.W. 338 Maui BISH 1978-09-28
119 Hymenasplenium unilaterale Kalauao, Aiea valley bottom Preserved_Specimen fern Hume, E.P. 143 Oahu BISH 1931-03-29
120 Hymenasplenium unilaterale 2nd gulch E of Puu Kaupakuhale, Mokuleia, Puu Kaala, Waianae Mts wet bottom Preserved_Specimen Fosberg, F.R. 8902 Oahu BISH 1932-10-23
121 Hymenasplenium unilaterale W Maui, Lahaina District, Lihau Montane Ohia Mixed Wet Forest at 4000 ft in a narrow drainage under a Metrosideros canopy with Coprosma, Broussaisia, Ilex, Cyrtandra spp. and a diversity of native ferns Preserved_Specimen Welton, P. 1288 Maui BISH 1991-08-27
122 Hymenasplenium unilaterale West Maui, in a narrow, steep drainage on the west wall of Kaluanui Stream. Site is east of Puu Kukui Watershed transect station 2.2100 Narrow, steep drainage. Specimen situated on a streambank in heavy shade of 5-6 meter canopy within Ohia Montane Wet Forest. Associated vegetation: Metrosideros polymorpha, Cheirodendron trigynum, Broussaisia arguta, Hedyotis hillebrandii, Melicope clusiifolia, Cibotium menziesii, Thelypteris stegnogrammoides, Diplazium sandwichianum, Nothoperanema rubiginosum, Vandenboschia davallioides, Sadleria pallida, Callistopteris baueriana, Deparia petersenii. Preserved_Specimen Meidell, J.S. 258 Maui BISH 1998-01-08
123 Hymenasplenium unilaterale Kau FR, Wood Valley wet shady gulch Preserved_Specimen 18 erect fern Russ, G.W. 55 Hawaii BISH 1927-07-15
124 Hymenasplenium unilaterale Lihue District. Blue Hole, headwaters of Wailua River, below Waialeale and Kawaikini. Metrosideros lowland wet forest and shrubland surrounded by ca. 1000 m cliffs w/ Psychotria mariniana, P. hexandra, Antidesma platyphyllum var. hillebrandii, Pipturus albidus, P. ruber, Urera glabra, Kadua centranthoides, Touchardia latifolia, Dianella sandwicensis, Polyscias o Preserved_Specimen Fern, terrestrial w/ long creeping rhizome, fertile, common in saturated waterfall and talus stream bank habitats. Wood, K.R. 18632 Kauai BISH 2021-01-28
125 Hymenasplenium unilaterale Powerline Trail Rocks in stream. Preserved_Specimen Forbes, C.N. 160 Kauai BISH 1909-08-06
126 Hymenasplenium unilaterale Ekahanui alluvial rubble, densely shaded streambed Preserved_Specimen clumped terrestrial Takeuchi, W.N. 3519 Oahu BISH 1987-04-25
127 Hymenasplenium unilaterale Punaula Valley Mossy rocks by stream. Preserved_Specimen St.John, H. 12791 Molokai BISH 1932-12-28
128 Hymenasplenium unilaterale Puu Ohia in restoration area. Growing under Psychotria kaduana, Hibiscus arnottianus subsp. punaluuensis, Metrosideros polymorpha, and Cinammomum burmanii. Growing amongst Reholttumia hudsoniana, Oplismenus hirtellus, and Cibotium chamissoi. Preserved_Specimen Thomas, M.K. 52 Oahu BISH 2021-01-06
129 Hymenasplenium unilaterale Wailupe. Southern Koolau Mountains. Growing along stream bank in gulch. Area dominated by Dicranopteris linearis, Miconia crenata, Rubus rosifolius, Christella parasitica, Peperomia membranacea, and Selaginella arbuscula. Also growing with Deparia prolifera, Tectaria gaudichaudii, Metrosideros polymorpha, Antidesma platyphyllum, Bobea elatior, Kadua affinis, Cibotium chamissoi, Sadleria pallida, and others. Preserved_Specimen Fronds both narrowly ovate and ovate. Some plants with pinna less falcate than others. Rhizome creeping. Thomas, M.K. 643 Oahu BISH 2024-04-09
130 Hymenasplenium unilaterale East Maui, Hana District, Waihoi Valley under a large bank of Dicranopteris linearis, with Peperomia Preserved_Specimen Terrestrial. Harrison, B. 11 Maui BISH 1972-09-22
131 Hymenasplenium unilaterale S.Kohala District; Kohala Mts.; Kohakohou Stream; Below diversion dam; deeply carved drainage Metrosideros-Cheirodendron Montane Wet Forest w/ Clermontia drepanomorpha, Asplenium excisum, Carex alligata, Cyanea tritomantha, Dryopteris spp., Athyrium microphyllum, Diplazium sandwichianum, bryophytes common. Preserved_Specimen Rhizome brown, creeping in saturated talus; stipe and rachis black-brown; lamina filmy green. Wood, K.R. 4698 Hawaii BISH 1995-10-26
132 Hymenasplenium unilaterale Puna, Olaa FR, ~10.8 km from jct. of Stainback Hwy. and Hwy. 17 towards Kulani Prison closed Metrosideros, Cibotium forest Preserved_Specimen Herat, T. 588 Hawaii BISH 1974-06-15
133 Hymenasplenium unilaterale Kilauea, Thurston lava tube rain forest Preserved_Specimen Morley, H. 115 Hawaii BISH 1934-08-06
134 Hymenasplenium unilaterale Kaiholena Unit, TNC Kau Preserve, Old Plantation Springs area Closed to open Metrosideros forest 50-70% cover @ 8-15 m with Cheirodendron trigynum 10-30% cover @ 5-8 m, over secondary canopy of native shrubs with Cibotium chamissoi 50-80% @ 3-5 m, Sadleria 10-20% @ 1-3 m, and Broussaisia arguta 10-40% @ 1.5-4 m, dissected by riparian habitat Preserved_Specimen Rhizome creeping on saturated basalt walls of drainage Wood, K.R. 12856 Hawaii BISH 2008-03-11
135 Hymenasplenium unilaterale Kaluaa Gulch, Central Waianae Mountains Ceodes - Planchonella forest with Psidium and Schinus. Growing on talus slope with Microlepia strigosa, Oplismenus hirtellus, Diplazium sandwichianum, Miconia crenata, and Rubus rosifolius. Preserved_Specimen Thomas, M.K. 135 Oahu BISH 2021-10-06
136 Hymenasplenium unilaterale Wailele Stream, Olokui, Wailau Moist shaded bank. Preserved_Specimen St.John, H. 23273 Molokai BISH 1948-02-05
137 Hymenasplenium unilaterale Kaneohu-nui shady gulch Preserved_Specimen Topping, D.L. 3678 Oahu BISH 1933-10-01
138 Hymenasplenium unilaterale Waianae Mountains, head of Keekee Gulch below Hakaloa, Kuaokala Forest Reserve On earth bank in moist botom of gulch Preserved_Specimen Fronds isolated, arching Fosberg, F.R. 12867 Oahu BISH 1936-02-02
139 Hymenasplenium unilaterale West Maui, in the upper headwaters of Honolua Stream. Site is east of of Puu Kukui Watershed transect station 3.450-3.700 Specimen situated on a streambank in heavy shade of 5-6 meter canopy within Ohia Montane Wet Forest. Associated vegetation: Metrosideros polymorpha, Cheirodendron trigynum, Psychotria kaduana, Broussaisia arguta, Cyanea macrostegia ssp. macrostegia, Cyrtandra grayana, Phyllostegia ambigua, Cibotium glaucum, Selaginella arbuscula. Preserved_Specimen Meidell, J.S. 263 Maui BISH 1997-12-09
140 Hymenasplenium unilaterale West Maui, Lahaina District, West Maui Natural Area Reserve-Panaewa Section, Halona Gulch Terrestrial, among rocks in intermittent stream in dense shade. Vegetation: Metrosideros, Cheirodendron, Ilex, Coprosma, Myrsine, Dodonaea, Pipturus, Touchardia, Urera, Neraudia, Cibotium, Sadleria, Athyrium, Cyrtandra, Clermontia, Cyanea, Diplazium, Dryopteris, Ctenitis, Blechnum, Tibouchina, Rubus, Peperomia. Preserved_Specimen Oppenheimer, H.L. 92126 Maui BISH 2021-09-27
141 Hymenasplenium unilaterale Waipio, Kipapa Gulch. In moist gully. Preserved_Specimen Hosaka, E.Y. 1048 Oahu BISH 1933-05-28
142 Hymenasplenium unilaterale Lihue District Waiahi south branch, north of Kamooloa, headwaters. Open wind- swept shrubland ridges dissected by drainages and patches of low forest 4-7 (-10) m tall, 90-95% cover. Metrosideros ca. 60% with 20% Cheirodendron interpersed with Psychotria, Broussaisia, Tetraplasandra, Kadua, Melicope, Perrottetia, ferns os Dicranopteris, Sticherus, Sphenomeris, Selaginella, Pneuma Preserved_Specimen terrestrial, rhizome short, sub-erect, common Wood, K.R. 14421 Kauai BISH 2010-12-17
143 Hymenasplenium unilaterale 2nd gulch, e. of Puu Kaupakuhale. n.e. slope of Puu Kaala, Waianae Mts. Mokuleia moist woods Preserved_Specimen St.John, H. 13172 Oahu BISH 1933-05-14
144 Hymenasplenium unilaterale Waimea District; Kawaiiki Ridge, above Koaie twin falls Metrosideros-Acacia mesic forest. W/ Dodonaea, Santalum freycinetianum, Kadua affinis, Xylosma hawaiiensis, Pouteria sandwicensis, Antidesma playphylla, Zanthoxylum dipetalum, Melicope ovata, M. barbigera, Poa sandwicensis, Gahnia be Preserved_Specimen terrestrial, with Pteris irregularis, on steep slope with brown granular soil Wood, K.R. 13877 Kauai BISH 2009-08-01
145 Hymenasplenium unilaterale Kapuna Gulch. In releve 23 Wet drainage with Pouteria sandwicensis; Hibiscus arnottianus; Aleurites moluccana; Claoxylon sandwicense; Diplazium sandwichianum. Preserved_Specimen Welton, P. 589 Oahu BISH 1991-10-10
146 Hymenasplenium unilaterale Lihue-koloa Forest Reserve; NW of Wahiawa bog, along tributary of Wahiawa stream, NW of stream and SE of Hulua Wet forest dominated by Metrosideros, Antidesma, Cyrtandra sp., and Athyrium. common, in wet, seeping areas. Preserved_Specimen Epiphytic on rocks or teerrestrial. Flynn, T.W. 2806 Kauai BISH 1988-04-07
147 Hymenasplenium unilaterale Kamokou Preserve, Kawela intake, Moloka`i. Montane wet forest of gulch. Preserved_Specimen Terrestrial fern. Pratt, L.W. 1237 Molokai BISH 1982-05-29
148 Hymenasplenium unilaterale Puukaneoa Trail Puukapapa Central Gulch behind Kunia, Waianae Mts Growing in stream bed in shade. Common. Preserved_Specimen Yee, R.S.N. 17 Oahu BISH 1971-08-17
149 Hymenasplenium unilaterale N slope of Haleakala on a waterfall Preserved_Specimen Forbes, C.N. 1212 Maui BISH 1919-08-21
150 Hymenasplenium unilaterale Waianae range, Kaluaa gulch, start of steep climb to summit Very shady, covered by Athyrium Preserved_Specimen Herat, T. 462 Oahu BISH 1973-07-15
151 Hymenasplenium unilaterale Iliiliula, Kauai. Ken Wood's promise land Stream bank growing with Metrosideros polymorpha var glaberrima, Cheirodendron faurei, Coprosma wimeae, Polyscias oahuensis, Polyscias kavaiensis, Coprosma kauaiensis, Kadua tryblium, Perottetia sandwicensis, Syzygium sandwicense, Hydrangea arguta, Psychotria hexandra subsp. hexandra, Labordia lydgatei, Melicope wawrae, Kadua affinis, Freycinetia arborea, Adenophorus periens, Cyanea undulata, Cyanea recta, Cyanea spathulata, Cyrtandra kealiae, Cyrtandra longifolia, Cyrtandra paludosa, Cyrtandra oenobarba, Cyanea fissa, Peperomia spp., Sadleria spp., Dubautia spp., Diplopterygium pinnatum, Dicranopteris, Amauropelta globulifera. Preserved_Specimen KPEPP 12022103 Kauai BISH 2021-11-10
152 Hymenasplenium unilaterale East Maui, Haiku Uka, near Waikamoi Flume Road within Koolau Forest Reserve 5 m `Ohi`a Montane Wet Forest along a steep rocky streambed. Assoc. veg.: Cheirodendron trigynum, Broussaisia arguta, Cibotium glaucum, Sadleria pallida, S. squarrosa, Athyrium microphyllum, Diplazium sandwichianum, Callistopteris baueriana, Asplenium insiticium. Preserved_Specimen Oppenheimer, H.L. 293 Maui BISH 1998-02-16
153 Hymenasplenium unilaterale Kahanahaiki Valley (a small valley between Makua and Pahole Valleys in the Waianae Mountains.) Unusually rich in fern numbers and species, including Marattia douglasii, Asplenium kaulfussii and a large population of Diellia x lauii (D. falcatum x unisora) covering 1/2 hectare in a manner that made it difficult to walk without stepping on them. Preserved_Specimen Palmer, D.D. 2893 Oahu BISH 1998-01-18
154 Hymenasplenium unilaterale Pauoa Valley damp places Preserved_Specimen Rock, J.F.C. Oahu BISH 1908-10-29
155 Hymenasplenium unilaterale West Maui, above the south tributary of Kahana Iki Stream Along a steep, rocky bank. Specimen was situated terrestrially and shaded beneath 5-6 meter within Ohia/Lama Lowland Mesic Forest. Associated vegetation: Touchardia latifolia, Cyrtandra grayi, Cyrtandra hawaiensis, Selaginella arbuscula, Blechnum occidentale, Deparia petersenii, Thelypteris parasitica, Thelypteris stegnogrammoides, Tectaria circutaria ssp. gaudichaudii, Nephrolepis exaltata. Preserved_Specimen Meidell, J.S. 271 Maui BISH 1997-12-03
156 Hymenasplenium unilaterale Hulopoe Gulch, head of Dark, rich gulch Preserved_Specimen Degener, O. 30155 Lanai BISH 1963-08-11
157 Hymenasplenium unilaterale Ukumehame Gulch dense, wet Metrosideros forest Preserved_Specimen St.John, H. 25719 Maui BISH 1955-12-28
158 Hymenasplenium unilaterale Wainiha Valley stream below Hono o Na Pali on ground in Hibiscus tiliaceus thicket along stream. Preserved_Specimen Wilson, K.A. 223 Kauai BISH 1952-12-31
159 Hymenasplenium unilaterale Kahanahaiki Valley (a small valley between Makua and Pahole Valleys in the Waianae Mountains.) Unusually rich in fern numbers and species, including Marattia douglasii, Asplenium kaulfussii and a large population of Diellia x lauii (D. falcatum x unisora) covering 1/2 hectare in a manner that made it difficult to walk without stepping on them. Preserved_Specimen Palmer, D.D. 2893 Oahu BISH 1998-01-18
160 Hymenasplenium unilaterale Waiahole-Waikane trail Preserved_Specimen Topping, D.L. 2707 Oahu BISH 1923-07-29
161 Hymenasplenium unilaterale East Maui, Makawao Forest Reserve, wet gully west of Borge Ridge Road Preserved_Specimen Hobdy, R.W. 3771 Maui BISH 1995-03-27
162 Hymenasplenium unilaterale Akaka Falls, Honomu Preserved_Specimen Forbes, C.N. 624 Hawaii BISH 1915-05-23
163 Hymenasplenium unilaterale 17 mi from Kohala toward Waimea Preserved_Specimen Degener, O. 30434 Hawaii BISH 1976-08-14
164 Hymenasplenium unilaterale Kaluaa-Hapapa. field no. Waianae 562 a terrestrial on bare soil, rubiaceous forest with e.g. Coprosma fauriei, Morinda trimera, and Psychotria mariniana Preserved_Specimen Takeuchi, W.N. 3057 Oahu BISH 1986-12-07
165 Hymenasplenium unilaterale N side of ridge on trail to Kaala moist woods Preserved_Specimen Bergman, H.F. Oahu BISH 1928-02-11
166 Hymenasplenium unilaterale near Hilo forest Preserved_Specimen Newell, Brother M. Hawaii BISH 1910-01-01
167 Hymenasplenium unilaterale East Maui, Hana District, Haleakala, Kipahulu Valley closed Acacia koa and Metrosideros forest Preserved_Specimen Higashino, P.K. 10031 Maui BISH 1983-07-17
168 Hymenasplenium unilaterale Large subgulch system in SW section of main gulch. Pahole Gulch, Waianae Mts., Waialua, Oahu On gulch floor in company with Athyrium. Aleurites-Pisonia-Psidium canopy Preserved_Specimen Nagata, K.M. 1335 Oahu BISH 1976-06-23
169 Hymenasplenium unilaterale unknown Preserved_Specimen BISH
170 Hymenasplenium unilaterale Pualaia Preserved_Specimen Forbes, C.N. 1755 Maui BISH 1919-12-05
171 Hymenasplenium unilaterale East Maui, Kula Pipe Line between Waikamoi Stream and Puohokamoa Stream Preserved_Specimen Krajina, V.J. 620417050 Maui BISH 1962-04-17
172 Hymenasplenium unilaterale Kohala Mts., Waimea Preserved_Specimen Forbes, C.N. 505 Hawaii BISH 1911-09-01
173 Hymenasplenium unilaterale Mountains near Koele Preserved_Specimen Forbes, C.N. 26 Lanai BISH 1913-06-01
174 Hymenasplenium unilaterale Honokahau Drainage Basin Preserved_Specimen Forbes, C.N. 402 Maui BISH 1917-09-25
175 Hymenasplenium unilaterale Molokai Preserved_Specimen Russell, C.A. Molokai BISH 1987-10-22
176 Hymenasplenium unilaterale East Maui, Upper Shelf headwalls and drainages of Kipahulu Valley Preserved_Specimen Welton, P. 1975 Maui BISH 1997-09-25
177 Hymenasplenium unilaterale East Maui, lower flume, Waikamoi Preserved_Specimen Hobdy, R.W. 593 Maui BISH 1979-05-20
178 Hymenasplenium unilaterale Mapulehu Valley Preserved_Specimen Forbes, C.N. 303 Molokai BISH 1912-07-01
179 Hymenasplenium unilaterale Kipahulu Valley, Haleakala, NPS Expedition III, W camp, transect 4 Preserved_Specimen Russell, C.A. 547 Maui BISH 1984-03-02
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