Description
Key Characters:
Growth Form:
Shrubs or trees.
Stems:
Roots:
Leaves:
Leaves simple.
Opposite.
Surfaces glabrous to pubescent, membranous to coriaceous.
Margins entire to revolute.
Stipules valvate, distinct or connate, usually firm and appressed, not forming a calyptra, caducous; stipules.
Flowers:
Flowers in inflorescences terminal, paniculate, usually with 1 or more orders of verticillate branching, ultimate branches usually terminated by a 3–flowered cymule, or inflorescences trichotomous from base with 1–2 orders of opposite divergent branching, inflorescences rarely capitate or globose.
Flowers insect-pollinated, bisexual (perfect) and usually heterostylous, in Hawaiian species functionally unisexual.
Calyx 4–6-lobed, glabrous or pubescent, the tube usually shorter than hypanthium, truncate or lobes short, acute or obtuse.
Corolla white, 4–6-lobed, the tube somewhat expanded toward apex, glabrous or sometimes pubescent at the throat, the lobes reflexed at anthesis.
Stamens as many as and alternate with the corolla lobes; filaments usually inserted on the corolla tube or throat; anthers exserted at anthesis in functionally staminate flowers, dithecal, opening by longitudinal slits.
Ovary inferior, occasionally partly so, or very rarely superior, 2–celled, placentation basal; ovules 1 per cell, erect, anatropous; style 1 or as many as carpels, terminal, slender; stigmas 2, lobed or capitate, divergent in functionally pistillate flowers, appressed in functionally staminate flowers, dry or occasionally wet.
Fruit:
Fruit drupaceous; red; orange (Hawaiian species); or sometimes purplish black; globose; pyriform; fusiform; or cylindrical; pyrenes 1–2(–4); triangular or semicircular in cross section; sometimes with secondary ridges developed; endocarp usually thick and hard.
Seeds 1 per pyrene; usually conforming to the form of the pyrene; seed coat thin and in some Hawaiian taxa invaginated on the ventral or inner face into a T-shape in cross section; endosperm otherwise not ruminate; carnose; the embryo small with flat; thin cotyledons; seeds with well-developed oily endosperm; or endosperm occasionally scanty or absent.
Ploidy:
Habitat:
Widespread throughout the tropics.
Elevation Range: