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Bolboschoenus maritimus subsp. paludosus

(A.Nelson) T.Koyama (1980)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Cyperaceae Genus: Bolboschoenus

Description

Key Characters:

Growth Form: Robust perennials, with horizontal, long–creeping rhizomes, also bearing tubers.

Stems: Rhizomes repent, clothed with brown scales, forming ligneous tubers at culm bases and nodes, culms erect, with several nodes, leafy, trigonous, smooth, 30–150 cm tall.

Roots:

Leaves: Leaves simple. Alternate. Blades linear, 3–8 mm wide. Apex acuminate. Surfaces scabrous on distal margins and midrib toward apex on lower surface, otherwise smooth. Margins entire. Sheathing; basal sheaths brown. Stipules absent.

Flowers: Flowers in inflorescences moderately dense, anthelate, consisting of terminal, congested, head-like clusters of 3 to numerous spikelets, +/- with 1–4 rays, each terminated by a headlike cluster of spikelets, the rays up to 5 cm long; involucral bracts 2–5, the lower ones much surpassing the inflorescence, the longest up to 30 cm long; spikelets pale brown, ellipsoid, ovoid–ellipsoid, or subcylindrical, terete (cylindrical), 1–2.5 cm long, 0.6–1 cm in diameter, apex subobtuse; glumes pale brown with pale margins, thin, membranous, narrowly ovate, 5–6.5 mm long, ca. 3 mm wide, tapering to an acute, bifid apex, pubescent externally, 1(3)– nerved, the midnerve prolonged ca. 2 mm beyond the apex. Flowers bisexual (perfect). Perianth bristles 2–6, up to ca. ½ as long as the achene, caducous. Corolla (petals) absent. Stamens 3. Ovary superior.

Fruit: Achenes brown; glossy; obovate; 3–4 mm long; 1.8–2.2 mm wide; lenticular; puncticulate; apex apiculate. Seeds 1; seed coat thin; free from pericarp; embryo basal; endosperm abundant.

Ploidy: 2n = 40; 76–77; 80; 86; 90; 96; 104; 106; 110; 114

Habitat: Primarily in coastal sites.

Elevation Range: 0–6 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Ni'ihau(Incl. Lehua) Indigenous
Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Pollinators

Bibliography

Name Published In: Acta Phytotax. Geobot. 31: 148 (1980)

Other References

Wagner et al. 1990:1384 (Ni, K, O, Mo, M); Staples et al. 2002:9 (H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama unknown PreservedSpecimen Collector Number: s.n. BISH Specimen
2 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama salt lake Water's edge. PRESERVED_SPECIMEN Oahu BISH 12/13/1903
3 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pond's southern end PreservedSpecimen Stokes, J.F.G. Collector Number: s.n. Niihau BISH Specimen
4 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pearl Harbor National Wildlife Refuge, Oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN Crago, L.M. Imada, C.; Silbernagle, M. Collector Number: 2005-065 Oahu BISH 2/23/2005
5 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kahuku, by ditch 300 yds from ocean, leading from sugar mill to ocean PRESERVED_SPECIMEN pearsall, g.a. collector number: 470 Maui BISH 9/12/1959
6 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama ewa beach PRESERVED_SPECIMEN woolford, h.a. Oahu BISH 6/26/1956
7 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama SE Molokai In Batis-covered depression back of ocean. PRESERVED_SPECIMEN Herbst, D.R. 1896 Molokai BISH 8/8/1970
8 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pupuo? Pupao? Pupoo? PRESERVED_SPECIMEN Forbes, C.N. 408.Mo Molokai BISH 8/1/1912
9 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kailua PRESERVED_SPECIMEN st.john, h. collector number: 10044 Oahu BISH 11/11/1929
10 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama honolulu near ala moana Marshes. PRESERVED_SPECIMEN neal, m.c. collector number: 1192 Oahu BISH 12/2/1939
11 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama koko head One chief species on the edge of a pool of mud by a spring on the N.W. corner of saline mud flats near Koko Head. PRESERVED_SPECIMEN harris, j.a. Oahu BISH 8/26/1924
12 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama backwater between highway and beach near kaunakakai PRESERVED_SPECIMEN herbst, d.r. collector number: 1896 Molokai BISH 8/8/1970
13 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Heeia Bridge, sea level Margin of tidal marsh. PRESERVED_SPECIMEN Egler, F.E. 37-137 Oahu BISH 7/4/1937
14 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Oahu PRESERVED_SPECIMEN Mann, H. 230 Oahu BISH 2/6/1905
15 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama heeia bridge, sea level Margin of tidal marsh. PRESERVED_SPECIMEN egler, f.e. collector number: 37-137 Oahu BISH 7/4/1937
16 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapiolani Park, Honolulu PRESERVED_SPECIMEN Munro, G.C. 1 Oahu BISH 11/22/1948
17 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kaelepulu stream near kailua, o'ahu PRESERVED_SPECIMEN stokes, j.f.g. Oahu BISH 1/2/1920
18 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama james campbell national wildlife refuge, kahuku. in vicinity of bir-viewing gazebo Growing in standing water with Bacopa monnieri, Brachiaria mutica, Paspalum vaginatum, Echinochloa colona. PRESERVED_SPECIMEN imada, c.t. engilis, a.; duk, t. collector number: 92-26 Oahu BISH 4/5/1992
19 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama betw lono harbor and kaunakakai PRESERVED_SPECIMEN corn, c. Molokai BISH 5/18/1982
20 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kailua PRESERVED_SPECIMEN St. John, H. 10044 Oahu BISH 11/11/1929
21 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapiolani Park, Honolulu, sea level Brackish water, wet places. PRESERVED_SPECIMEN Wilder, G.P. 60 Oahu BISH 10/25/1923
22 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Halalii Lake, S of edge of dry lake bed PRESERVED_SPECIMEN St. John, H. 22756 Niihau BISH 8/12/1947
23 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pond's southern end at base of Metrosideros tree PRESERVED_SPECIMEN stokes, j.f.g. Maui BISH 1/8/1912
24 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama oahu PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 230 Oahu BISH
25 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapiolani park, honolulu, sea level Brackish water, wet places. PRESERVED_SPECIMEN wilder, g.p. collector number: 60 Oahu BISH 10/25/1923
26 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pearl harbor national wildlife refuge, oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN crago, l.m. imada, c.; silbernagle, m. collector number: 2005-065 Oahu BISH 2/23/2005
27 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ewa Beach Marina Collected from low, wet places in Batis swamp. Colonies are surrounded by Batis mats. No other vegetation in the near area. PRESERVED_SPECIMEN Funk, E.J. 604 Oahu BISH 5/1/1990
28 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kanaha Bird Reserve, near Kahului Abundant along margins of Kanaha Bird Reserve, with S. validus. PRESERVED_SPECIMEN Henrickson, J.S. 3734 Maui BISH 6/26/1969
29 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Backwater between highway and beach near Kaunakakai PRESERVED_SPECIMEN Herbst, D.R. 1896 Molokai BISH 8/8/1970
30 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kanaha bird reserve, near kahului Abundant along margins of Kanaha Bird Reserve, with S. validus. PRESERVED_SPECIMEN henrickson, j.s. collector number: 3734 Maui BISH 6/26/1969
31 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Between Lono Harbor and Kaunakakai PRESERVED_SPECIMEN Corn, C. Not on sheet Molokai BISH 5/18/1982
32 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama North Kona District, Makalawena, on west edge of Opaeula Pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van Dyke, P. Collector Number: s.n. Hawaii BISH 11/8/2000
33 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kamalo Marshy areas near road. Associated with Batis and Pluchea indica. PRESERVED_SPECIMEN Nagata, K.M. 1762 Molokai BISH 8/9/1978
34 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waialae PRESERVED_SPECIMEN Caum, E.L. s.n. Oahu BISH 5/24/1929
35 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Koloa District, Mahaulepu. PRESERVED_SPECIMEN T. W. Flynn, W. L. Wagner & D. Mix 7133 Kauai US 10/6/2004
36 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama koloa district, mahaulepu. PRESERVED_SPECIMEN t. w. flynn, w. l. wagner & d. mix 7133 Kauai US 10/6/2004
37 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waialae PRESERVED_SPECIMEN caum, e.l. Oahu BISH 5/24/1929
38 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama central maui, southern coast of isthmus, kealia pond national wildlife refuge. along mokulele hwy access road to refuge main off PRESERVED_SPECIMEN c. imada 98-17 Maui US 10/6/1998
39 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kalaupapa; airport Common on trees and boulders. PRESERVED_SPECIMEN wysong, m.l. collector number: 611 Oahu BISH 2/8/2005
40 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pearl Harbor National Wildlife Refuge, Oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN Crago, L.M. 2005-065 Oahu BISH 2/23/2005
41 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ponds southern end PRESERVED_SPECIMEN Stokes, J.F.G. s.n. Niihau BISH 1/1/1912
42 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waikiki, Honolulu, near the Moana Hotel PRESERVED_SPECIMEN Wilder, G.P. not on sheet Oahu BISH 10/27/1930
43 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waikiki swamp PRESERVED_SPECIMEN hitchcock, a.s. collector number: 13808 Oahu BISH 6/19/1916
44 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama in ditch alond kihei rd., at junction with lipoa st In brackish water in ditch along road. Found with Cyperus, Spirodela. PRESERVED_SPECIMEN ishikawa, s. collector number: 363 Maui BISH 12/3/1973
45 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ewa Beach PRESERVED_SPECIMEN Woolford, H.A. Not on sheet Oahu BISH 6/26/1956
46 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapaa, kawaihau, sea level Rare on marshy flat. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1646 Kauai BISH 10/29/1936
47 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Central Maui, southern coast of isthmus, Kealia Pond National Wildlife Refuge. Along Mokulele Hwy access road to refuge main office Common sedge forming solid patches roadside fronting Batis herbland. Fertile. PRESERVED_SPECIMEN Imada, C.T. 98-17 Maui BISH 10/6/1998
48 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama north kona district, makalawena, on west edge of opaeula pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van dyke, p. collector number: s.n. Hawaii BISH 11/8/2000
49 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kalaupapa; airport seasonal wetland PRESERVED_SPECIMEN Wysong, M.L. 611 Molokai BISH 2/8/2005
50 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pupuo? pupao? pupoo? PRESERVED_SPECIMEN forbes, c.n. collector number: 408.mo Molokai BISH 8/8/1912
51 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waialua PRESERVED_SPECIMEN Rock, J.F.C. 8758 Oahu BISH 5/21/2011
52 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kailua PRESERVED_SPECIMEN St.John, H. Collector Number: 10044 Oahu BISH 11/11/1929
53 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama halalii lake, s of edge of dry lake bed PRESERVED_SPECIMEN st.john, h. tuthill, l.d. collector number: 22756 Niihau BISH 8/12/1947
54 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kamalo PRESERVED_SPECIMEN nagata, k.m. kimura, b. collector number: 1762 Kauai BISH 8/9/1978
55 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waipio peninsula; waipio, pouhala marsh between kapakahi & waikele streams On margin of open water & growing in assoc. w/Bacopa, Eleocharis, Cyperus laevigatus, Batis. Occasional. PRESERVED_SPECIMEN imada, c.t. nishida, g. collector number: 96-5 Oahu BISH 12/18/1996
56 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama central maui, southern coast of isthmus, kealia pond national wildlife refuge. along mokulele hwy access road to refuge main office Common sedge forming solid patches roadside fronting Batis herbland. Fertile. PRESERVED_SPECIMEN imada, c.t. collector number: 98-17 Maui BISH 10/6/1998
57 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Honolulu PRESERVED_SPECIMEN Stokes, J.F.G. Oahu BISH 5/1/1919
58 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapaa, Kawaihau, sea level Rare on marshy flat. PRESERVED_SPECIMEN Hosaka, E.Y. 1646 Kauai BISH 10/29/1936
59 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama In ditch alond Kihei Rd., at junction with Lipoa St In brackish water in ditch along road. Found with Cyperus, Spirodela. PRESERVED_SPECIMEN Ishikawa, S. 363 Maui BISH 12/3/1973
60 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kahuku, by ditch 300 yds from ocean, leading from sugar mill to ocean On ditch bank, brackish water. PRESERVED_SPECIMEN Pearsall, G.A. 470 Oahu BISH 9/12/1959
61 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Salt Lake Water's edge. PRESERVED_SPECIMEN s.n. Oahu BISH 12/13/1903
62 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama James Campbell National Wildlife Refuge, Kahuku. In vicinity of bir-viewing gazebo Growing in standing water with Bacopa monnieri, Brachiaria mutica, Paspalum vaginatum, Echinochloa colona. PRESERVED_SPECIMEN Imada, C.T. 92-26 Oahu BISH 4/5/1992
63 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waikiki, honolulu, near the moana hotel PRESERVED_SPECIMEN wilder, g.p. Oahu BISH 10/27/1930
64 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Koko Head One chief species on the edge of a pool of mud by a spring on the N.W. corner of saline mud flats near Koko Head. PRESERVED_SPECIMEN Harris, J.A. s.n. Oahu BISH 8/26/1924
65 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kipapa Gulch, Waipahu, near beach Mud flat. PRESERVED_SPECIMEN Hosaka, E.Y. 759 Oahu BISH 9/25/1932
66 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama hamakua marsh, kailua, mauka (mountainside) of hamakua drive Pond dominated by Schoenoplectus mucronatus (70% cover), with Ludwigia palustris, Cuphea carthagenensis, Cyperus halpan, Juncus effusus, Utricularia gibba. PRESERVED_SPECIMEN imada, c.t. engilis, a.; yuen, a.; dr.duk collector number: 92-10 Hawaii BISH 4/4/1992
67 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama wailua golf course, fourth hole water hazard PRESERVED_SPECIMEN flynn, t. collector number: 1249 Kauai BISH 10/3/1985
68 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive Growing in muck and in open water; common. PRESERVED_SPECIMEN Imada, C.T. 92-10 Oahu BISH 4/4/1992
69 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama honolulu PRESERVED_SPECIMEN stokes, j.f.g. Oahu BISH 5/1/1919
70 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama North Kona District, Makalawena, on west edge of Opaeula Pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van Dyke, P. s.n. Hawaii BISH 11/8/2000
71 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama tantalus PRESERVED_SPECIMEN pearsall, g.a. Hawaii US 7/24/1953
72 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapiolani park, honolulu PRESERVED_SPECIMEN munro, g.c. collector number: 1 Oahu BISH 11/22/1948
73 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kipapa gulch, waipahu, near beach Mud flat. PRESERVED_SPECIMEN hosaka, e.y. collector number: 759 Oahu BISH 9/25/1932
74 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Honolulu near Ala Moana Marshes. PRESERVED_SPECIMEN Neal, M.C. 1192 Oahu BISH 12/2/1939
75 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama ewa beach marina Collected from low, wet places in Batis swamp. Colonies are surrounded by Batis mats. No other vegetation in the near area. PRESERVED_SPECIMEN funk, e.j. lee, m.; b.ching collector number: 604 Oahu BISH 5/1/1990
76 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waikiki swamp PRESERVED_SPECIMEN Hitchcock, A.S. 13808 Oahu BISH 6/19/1916
77 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waipio Peninsula; Waipio, Pouhala Marsh between Kapakahi & Waikele Streams On margin of open water & growing in assoc. w/Bacopa, Eleocharis, Cyperus laevigatus, Batis. Occasional. PRESERVED_SPECIMEN Imada, C.T. 96-5 Oahu BISH 12/18/1996
78 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama s.e. molokai In Batis-covered depression back of ocean. PRESERVED_SPECIMEN herbst, d.r. collector number: 1896 Molokai BISH 8/8/1970
79 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Wailua Golf Course, Fourth hole water hazard PRESERVED_SPECIMEN Flynn, T. 1249 Kauai BISH 10/3/1985
80 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waialua PRESERVED_SPECIMEN rock, j.f.c. collector number: 8758 Oahu BISH 5/21/2011
81 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Halalii Lake, S of edge of dry lake bed PreservedSpecimen St. John, H. Tuthill, L.D. Collector Number: 22756 Niihau BISH Specimen 8/12/1947
82 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama unknown PreservedSpecimen Collector Number: s.n. BISH Specimen
83 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama salt lake Water's edge. PRESERVED_SPECIMEN Oahu BISH 12/13/1903
84 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pond's southern end PreservedSpecimen Stokes, J.F.G. Collector Number: s.n. Niihau BISH Specimen
85 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pearl Harbor National Wildlife Refuge, Oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN Crago, L.M. Imada, C.; Silbernagle, M. Collector Number: 2005-065 Oahu BISH 2/23/2005
86 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kahuku, by ditch 300 yds from ocean, leading from sugar mill to ocean PRESERVED_SPECIMEN pearsall, g.a. collector number: 470 Maui BISH 9/12/1959
87 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama ewa beach PRESERVED_SPECIMEN woolford, h.a. Oahu BISH 6/26/1956
88 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama SE Molokai In Batis-covered depression back of ocean. PRESERVED_SPECIMEN Herbst, D.R. 1896 Molokai BISH 8/8/1970
89 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pupuo? Pupao? Pupoo? PRESERVED_SPECIMEN Forbes, C.N. 408.Mo Molokai BISH 8/1/1912
90 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kailua PRESERVED_SPECIMEN st.john, h. collector number: 10044 Oahu BISH 11/11/1929
91 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama honolulu near ala moana Marshes. PRESERVED_SPECIMEN neal, m.c. collector number: 1192 Oahu BISH 12/2/1939
92 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama koko head One chief species on the edge of a pool of mud by a spring on the N.W. corner of saline mud flats near Koko Head. PRESERVED_SPECIMEN harris, j.a. Oahu BISH 8/26/1924
93 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama backwater between highway and beach near kaunakakai PRESERVED_SPECIMEN herbst, d.r. collector number: 1896 Molokai BISH 8/8/1970
94 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Heeia Bridge, sea level Margin of tidal marsh. PRESERVED_SPECIMEN Egler, F.E. 37-137 Oahu BISH 7/4/1937
95 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Oahu PRESERVED_SPECIMEN Mann, H. 230 Oahu BISH 2/6/1905
96 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama heeia bridge, sea level Margin of tidal marsh. PRESERVED_SPECIMEN egler, f.e. collector number: 37-137 Oahu BISH 7/4/1937
97 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapiolani Park, Honolulu PRESERVED_SPECIMEN Munro, G.C. 1 Oahu BISH 11/22/1948
98 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kaelepulu stream near kailua, o'ahu PRESERVED_SPECIMEN stokes, j.f.g. Oahu BISH 1/2/1920
99 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama james campbell national wildlife refuge, kahuku. in vicinity of bir-viewing gazebo Growing in standing water with Bacopa monnieri, Brachiaria mutica, Paspalum vaginatum, Echinochloa colona. PRESERVED_SPECIMEN imada, c.t. engilis, a.; duk, t. collector number: 92-26 Oahu BISH 4/5/1992
100 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama betw lono harbor and kaunakakai PRESERVED_SPECIMEN corn, c. Molokai BISH 5/18/1982
101 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kailua PRESERVED_SPECIMEN St. John, H. 10044 Oahu BISH 11/11/1929
102 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapiolani Park, Honolulu, sea level Brackish water, wet places. PRESERVED_SPECIMEN Wilder, G.P. 60 Oahu BISH 10/25/1923
103 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Halalii Lake, S of edge of dry lake bed PRESERVED_SPECIMEN St. John, H. 22756 Niihau BISH 8/12/1947
104 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pond's southern end at base of Metrosideros tree PRESERVED_SPECIMEN stokes, j.f.g. Maui BISH 1/8/1912
105 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama oahu PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 230 Oahu BISH
106 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapiolani park, honolulu, sea level Brackish water, wet places. PRESERVED_SPECIMEN wilder, g.p. collector number: 60 Oahu BISH 10/25/1923
107 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pearl harbor national wildlife refuge, oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN crago, l.m. imada, c.; silbernagle, m. collector number: 2005-065 Oahu BISH 2/23/2005
108 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ewa Beach Marina Collected from low, wet places in Batis swamp. Colonies are surrounded by Batis mats. No other vegetation in the near area. PRESERVED_SPECIMEN Funk, E.J. 604 Oahu BISH 5/1/1990
109 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kanaha Bird Reserve, near Kahului Abundant along margins of Kanaha Bird Reserve, with S. validus. PRESERVED_SPECIMEN Henrickson, J.S. 3734 Maui BISH 6/26/1969
110 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Backwater between highway and beach near Kaunakakai PRESERVED_SPECIMEN Herbst, D.R. 1896 Molokai BISH 8/8/1970
111 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kanaha bird reserve, near kahului Abundant along margins of Kanaha Bird Reserve, with S. validus. PRESERVED_SPECIMEN henrickson, j.s. collector number: 3734 Maui BISH 6/26/1969
112 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Between Lono Harbor and Kaunakakai PRESERVED_SPECIMEN Corn, C. Not on sheet Molokai BISH 5/18/1982
113 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama North Kona District, Makalawena, on west edge of Opaeula Pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van Dyke, P. Collector Number: s.n. Hawaii BISH 11/8/2000
114 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kamalo Marshy areas near road. Associated with Batis and Pluchea indica. PRESERVED_SPECIMEN Nagata, K.M. 1762 Molokai BISH 8/9/1978
115 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waialae PRESERVED_SPECIMEN Caum, E.L. s.n. Oahu BISH 5/24/1929
116 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Koloa District, Mahaulepu. PRESERVED_SPECIMEN T. W. Flynn, W. L. Wagner & D. Mix 7133 Kauai US 10/6/2004
117 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama koloa district, mahaulepu. PRESERVED_SPECIMEN t. w. flynn, w. l. wagner & d. mix 7133 Kauai US 10/6/2004
118 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waialae PRESERVED_SPECIMEN caum, e.l. Oahu BISH 5/24/1929
119 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama central maui, southern coast of isthmus, kealia pond national wildlife refuge. along mokulele hwy access road to refuge main off PRESERVED_SPECIMEN c. imada 98-17 Maui US 10/6/1998
120 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kalaupapa; airport Common on trees and boulders. PRESERVED_SPECIMEN wysong, m.l. collector number: 611 Oahu BISH 2/8/2005
121 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pearl Harbor National Wildlife Refuge, Oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN Crago, L.M. 2005-065 Oahu BISH 2/23/2005
122 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ponds southern end PRESERVED_SPECIMEN Stokes, J.F.G. s.n. Niihau BISH 1/1/1912
123 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waikiki, Honolulu, near the Moana Hotel PRESERVED_SPECIMEN Wilder, G.P. not on sheet Oahu BISH 10/27/1930
124 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waikiki swamp PRESERVED_SPECIMEN hitchcock, a.s. collector number: 13808 Oahu BISH 6/19/1916
125 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama in ditch alond kihei rd., at junction with lipoa st In brackish water in ditch along road. Found with Cyperus, Spirodela. PRESERVED_SPECIMEN ishikawa, s. collector number: 363 Maui BISH 12/3/1973
126 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ewa Beach PRESERVED_SPECIMEN Woolford, H.A. Not on sheet Oahu BISH 6/26/1956
127 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapaa, kawaihau, sea level Rare on marshy flat. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1646 Kauai BISH 10/29/1936
128 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Central Maui, southern coast of isthmus, Kealia Pond National Wildlife Refuge. Along Mokulele Hwy access road to refuge main office Common sedge forming solid patches roadside fronting Batis herbland. Fertile. PRESERVED_SPECIMEN Imada, C.T. 98-17 Maui BISH 10/6/1998
129 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama north kona district, makalawena, on west edge of opaeula pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van dyke, p. collector number: s.n. Hawaii BISH 11/8/2000
130 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kalaupapa; airport seasonal wetland PRESERVED_SPECIMEN Wysong, M.L. 611 Molokai BISH 2/8/2005
131 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pupuo? pupao? pupoo? PRESERVED_SPECIMEN forbes, c.n. collector number: 408.mo Molokai BISH 8/8/1912
132 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waialua PRESERVED_SPECIMEN Rock, J.F.C. 8758 Oahu BISH 5/21/2011
133 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kailua PRESERVED_SPECIMEN St.John, H. Collector Number: 10044 Oahu BISH 11/11/1929
134 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama halalii lake, s of edge of dry lake bed PRESERVED_SPECIMEN st.john, h. tuthill, l.d. collector number: 22756 Niihau BISH 8/12/1947
135 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kamalo PRESERVED_SPECIMEN nagata, k.m. kimura, b. collector number: 1762 Kauai BISH 8/9/1978
136 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waipio peninsula; waipio, pouhala marsh between kapakahi & waikele streams On margin of open water & growing in assoc. w/Bacopa, Eleocharis, Cyperus laevigatus, Batis. Occasional. PRESERVED_SPECIMEN imada, c.t. nishida, g. collector number: 96-5 Oahu BISH 12/18/1996
137 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama central maui, southern coast of isthmus, kealia pond national wildlife refuge. along mokulele hwy access road to refuge main office Common sedge forming solid patches roadside fronting Batis herbland. Fertile. PRESERVED_SPECIMEN imada, c.t. collector number: 98-17 Maui BISH 10/6/1998
138 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Honolulu PRESERVED_SPECIMEN Stokes, J.F.G. Oahu BISH 5/1/1919
139 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapaa, Kawaihau, sea level Rare on marshy flat. PRESERVED_SPECIMEN Hosaka, E.Y. 1646 Kauai BISH 10/29/1936
140 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama In ditch alond Kihei Rd., at junction with Lipoa St In brackish water in ditch along road. Found with Cyperus, Spirodela. PRESERVED_SPECIMEN Ishikawa, S. 363 Maui BISH 12/3/1973
141 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kahuku, by ditch 300 yds from ocean, leading from sugar mill to ocean On ditch bank, brackish water. PRESERVED_SPECIMEN Pearsall, G.A. 470 Oahu BISH 9/12/1959
142 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Salt Lake Water's edge. PRESERVED_SPECIMEN s.n. Oahu BISH 12/13/1903
143 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama James Campbell National Wildlife Refuge, Kahuku. In vicinity of bir-viewing gazebo Growing in standing water with Bacopa monnieri, Brachiaria mutica, Paspalum vaginatum, Echinochloa colona. PRESERVED_SPECIMEN Imada, C.T. 92-26 Oahu BISH 4/5/1992
144 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waikiki, honolulu, near the moana hotel PRESERVED_SPECIMEN wilder, g.p. Oahu BISH 10/27/1930
145 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Koko Head One chief species on the edge of a pool of mud by a spring on the N.W. corner of saline mud flats near Koko Head. PRESERVED_SPECIMEN Harris, J.A. s.n. Oahu BISH 8/26/1924
146 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kipapa Gulch, Waipahu, near beach Mud flat. PRESERVED_SPECIMEN Hosaka, E.Y. 759 Oahu BISH 9/25/1932
147 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama hamakua marsh, kailua, mauka (mountainside) of hamakua drive Pond dominated by Schoenoplectus mucronatus (70% cover), with Ludwigia palustris, Cuphea carthagenensis, Cyperus halpan, Juncus effusus, Utricularia gibba. PRESERVED_SPECIMEN imada, c.t. engilis, a.; yuen, a.; dr.duk collector number: 92-10 Hawaii BISH 4/4/1992
148 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama wailua golf course, fourth hole water hazard PRESERVED_SPECIMEN flynn, t. collector number: 1249 Kauai BISH 10/3/1985
149 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive Growing in muck and in open water; common. PRESERVED_SPECIMEN Imada, C.T. 92-10 Oahu BISH 4/4/1992
150 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama honolulu PRESERVED_SPECIMEN stokes, j.f.g. Oahu BISH 5/1/1919
151 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama North Kona District, Makalawena, on west edge of Opaeula Pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van Dyke, P. s.n. Hawaii BISH 11/8/2000
152 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama tantalus PRESERVED_SPECIMEN pearsall, g.a. Hawaii US 7/24/1953
153 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapiolani park, honolulu PRESERVED_SPECIMEN munro, g.c. collector number: 1 Oahu BISH 11/22/1948
154 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kipapa gulch, waipahu, near beach Mud flat. PRESERVED_SPECIMEN hosaka, e.y. collector number: 759 Oahu BISH 9/25/1932
155 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Honolulu near Ala Moana Marshes. PRESERVED_SPECIMEN Neal, M.C. 1192 Oahu BISH 12/2/1939
156 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama ewa beach marina Collected from low, wet places in Batis swamp. Colonies are surrounded by Batis mats. No other vegetation in the near area. PRESERVED_SPECIMEN funk, e.j. lee, m.; b.ching collector number: 604 Oahu BISH 5/1/1990
157 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waikiki swamp PRESERVED_SPECIMEN Hitchcock, A.S. 13808 Oahu BISH 6/19/1916
158 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waipio Peninsula; Waipio, Pouhala Marsh between Kapakahi & Waikele Streams On margin of open water & growing in assoc. w/Bacopa, Eleocharis, Cyperus laevigatus, Batis. Occasional. PRESERVED_SPECIMEN Imada, C.T. 96-5 Oahu BISH 12/18/1996
159 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama s.e. molokai In Batis-covered depression back of ocean. PRESERVED_SPECIMEN herbst, d.r. collector number: 1896 Molokai BISH 8/8/1970
160 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Wailua Golf Course, Fourth hole water hazard PRESERVED_SPECIMEN Flynn, T. 1249 Kauai BISH 10/3/1985
161 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waialua PRESERVED_SPECIMEN rock, j.f.c. collector number: 8758 Oahu BISH 5/21/2011
162 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Halalii Lake, S of edge of dry lake bed PreservedSpecimen St. John, H. Tuthill, L.D. Collector Number: 22756 Niihau BISH Specimen 8/12/1947
163 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama unknown PreservedSpecimen Collector Number: s.n. BISH Specimen
164 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama salt lake Water's edge. PRESERVED_SPECIMEN Oahu BISH 12/13/1903
165 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pond's southern end PreservedSpecimen Stokes, J.F.G. Collector Number: s.n. Niihau BISH Specimen
166 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pearl Harbor National Wildlife Refuge, Oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN Crago, L.M. Imada, C.; Silbernagle, M. Collector Number: 2005-065 Oahu BISH 2/23/2005
167 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kahuku, by ditch 300 yds from ocean, leading from sugar mill to ocean PRESERVED_SPECIMEN pearsall, g.a. collector number: 470 Maui BISH 9/12/1959
168 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama ewa beach PRESERVED_SPECIMEN woolford, h.a. Oahu BISH 6/26/1956
169 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama SE Molokai In Batis-covered depression back of ocean. PRESERVED_SPECIMEN Herbst, D.R. 1896 Molokai BISH 8/8/1970
170 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pupuo? Pupao? Pupoo? PRESERVED_SPECIMEN Forbes, C.N. 408.Mo Molokai BISH 8/1/1912
171 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kailua PRESERVED_SPECIMEN st.john, h. collector number: 10044 Oahu BISH 11/11/1929
172 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama honolulu near ala moana Marshes. PRESERVED_SPECIMEN neal, m.c. collector number: 1192 Oahu BISH 12/2/1939
173 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama koko head One chief species on the edge of a pool of mud by a spring on the N.W. corner of saline mud flats near Koko Head. PRESERVED_SPECIMEN harris, j.a. Oahu BISH 8/26/1924
174 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama backwater between highway and beach near kaunakakai PRESERVED_SPECIMEN herbst, d.r. collector number: 1896 Molokai BISH 8/8/1970
175 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Heeia Bridge, sea level Margin of tidal marsh. PRESERVED_SPECIMEN Egler, F.E. 37-137 Oahu BISH 7/4/1937
176 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Oahu PRESERVED_SPECIMEN Mann, H. 230 Oahu BISH 2/6/1905
177 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama heeia bridge, sea level Margin of tidal marsh. PRESERVED_SPECIMEN egler, f.e. collector number: 37-137 Oahu BISH 7/4/1937
178 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapiolani Park, Honolulu PRESERVED_SPECIMEN Munro, G.C. 1 Oahu BISH 11/22/1948
179 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kaelepulu stream near kailua, o'ahu PRESERVED_SPECIMEN stokes, j.f.g. Oahu BISH 1/2/1920
180 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama james campbell national wildlife refuge, kahuku. in vicinity of bir-viewing gazebo Growing in standing water with Bacopa monnieri, Brachiaria mutica, Paspalum vaginatum, Echinochloa colona. PRESERVED_SPECIMEN imada, c.t. engilis, a.; duk, t. collector number: 92-26 Oahu BISH 4/5/1992
181 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama betw lono harbor and kaunakakai PRESERVED_SPECIMEN corn, c. Molokai BISH 5/18/1982
182 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kailua PRESERVED_SPECIMEN St. John, H. 10044 Oahu BISH 11/11/1929
183 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapiolani Park, Honolulu, sea level Brackish water, wet places. PRESERVED_SPECIMEN Wilder, G.P. 60 Oahu BISH 10/25/1923
184 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Halalii Lake, S of edge of dry lake bed PRESERVED_SPECIMEN St. John, H. 22756 Niihau BISH 8/12/1947
185 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pond's southern end at base of Metrosideros tree PRESERVED_SPECIMEN stokes, j.f.g. Maui BISH 1/8/1912
186 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama oahu PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 230 Oahu BISH
187 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapiolani park, honolulu, sea level Brackish water, wet places. PRESERVED_SPECIMEN wilder, g.p. collector number: 60 Oahu BISH 10/25/1923
188 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pearl harbor national wildlife refuge, oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN crago, l.m. imada, c.; silbernagle, m. collector number: 2005-065 Oahu BISH 2/23/2005
189 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ewa Beach Marina Collected from low, wet places in Batis swamp. Colonies are surrounded by Batis mats. No other vegetation in the near area. PRESERVED_SPECIMEN Funk, E.J. 604 Oahu BISH 5/1/1990
190 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kanaha Bird Reserve, near Kahului Abundant along margins of Kanaha Bird Reserve, with S. validus. PRESERVED_SPECIMEN Henrickson, J.S. 3734 Maui BISH 6/26/1969
191 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Backwater between highway and beach near Kaunakakai PRESERVED_SPECIMEN Herbst, D.R. 1896 Molokai BISH 8/8/1970
192 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kanaha bird reserve, near kahului Abundant along margins of Kanaha Bird Reserve, with S. validus. PRESERVED_SPECIMEN henrickson, j.s. collector number: 3734 Maui BISH 6/26/1969
193 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Between Lono Harbor and Kaunakakai PRESERVED_SPECIMEN Corn, C. Not on sheet Molokai BISH 5/18/1982
194 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama North Kona District, Makalawena, on west edge of Opaeula Pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van Dyke, P. Collector Number: s.n. Hawaii BISH 11/8/2000
195 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kamalo Marshy areas near road. Associated with Batis and Pluchea indica. PRESERVED_SPECIMEN Nagata, K.M. 1762 Molokai BISH 8/9/1978
196 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waialae PRESERVED_SPECIMEN Caum, E.L. s.n. Oahu BISH 5/24/1929
197 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Koloa District, Mahaulepu. PRESERVED_SPECIMEN T. W. Flynn, W. L. Wagner & D. Mix 7133 Kauai US 10/6/2004
198 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama koloa district, mahaulepu. PRESERVED_SPECIMEN t. w. flynn, w. l. wagner & d. mix 7133 Kauai US 10/6/2004
199 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waialae PRESERVED_SPECIMEN caum, e.l. Oahu BISH 5/24/1929
200 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama central maui, southern coast of isthmus, kealia pond national wildlife refuge. along mokulele hwy access road to refuge main off PRESERVED_SPECIMEN c. imada 98-17 Maui US 10/6/1998
201 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kalaupapa; airport Common on trees and boulders. PRESERVED_SPECIMEN wysong, m.l. collector number: 611 Oahu BISH 2/8/2005
202 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Pearl Harbor National Wildlife Refuge, Oahu Growing on edge of pond and into water. Other species Typha domingensis, Bacopa monnieri, Leptochloa fescua [?]. PRESERVED_SPECIMEN Crago, L.M. 2005-065 Oahu BISH 2/23/2005
203 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ponds southern end PRESERVED_SPECIMEN Stokes, J.F.G. s.n. Niihau BISH 1/1/1912
204 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waikiki, Honolulu, near the Moana Hotel PRESERVED_SPECIMEN Wilder, G.P. not on sheet Oahu BISH 10/27/1930
205 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waikiki swamp PRESERVED_SPECIMEN hitchcock, a.s. collector number: 13808 Oahu BISH 6/19/1916
206 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama in ditch alond kihei rd., at junction with lipoa st In brackish water in ditch along road. Found with Cyperus, Spirodela. PRESERVED_SPECIMEN ishikawa, s. collector number: 363 Maui BISH 12/3/1973
207 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Ewa Beach PRESERVED_SPECIMEN Woolford, H.A. Not on sheet Oahu BISH 6/26/1956
208 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapaa, kawaihau, sea level Rare on marshy flat. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1646 Kauai BISH 10/29/1936
209 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Central Maui, southern coast of isthmus, Kealia Pond National Wildlife Refuge. Along Mokulele Hwy access road to refuge main office Common sedge forming solid patches roadside fronting Batis herbland. Fertile. PRESERVED_SPECIMEN Imada, C.T. 98-17 Maui BISH 10/6/1998
210 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama north kona district, makalawena, on west edge of opaeula pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van dyke, p. collector number: s.n. Hawaii BISH 11/8/2000
211 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kalaupapa; airport seasonal wetland PRESERVED_SPECIMEN Wysong, M.L. 611 Molokai BISH 2/8/2005
212 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama pupuo? pupao? pupoo? PRESERVED_SPECIMEN forbes, c.n. collector number: 408.mo Molokai BISH 8/8/1912
213 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waialua PRESERVED_SPECIMEN Rock, J.F.C. 8758 Oahu BISH 5/21/2011
214 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kailua PRESERVED_SPECIMEN St.John, H. Collector Number: 10044 Oahu BISH 11/11/1929
215 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama halalii lake, s of edge of dry lake bed PRESERVED_SPECIMEN st.john, h. tuthill, l.d. collector number: 22756 Niihau BISH 8/12/1947
216 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kamalo PRESERVED_SPECIMEN nagata, k.m. kimura, b. collector number: 1762 Kauai BISH 8/9/1978
217 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waipio peninsula; waipio, pouhala marsh between kapakahi & waikele streams On margin of open water & growing in assoc. w/Bacopa, Eleocharis, Cyperus laevigatus, Batis. Occasional. PRESERVED_SPECIMEN imada, c.t. nishida, g. collector number: 96-5 Oahu BISH 12/18/1996
218 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama central maui, southern coast of isthmus, kealia pond national wildlife refuge. along mokulele hwy access road to refuge main office Common sedge forming solid patches roadside fronting Batis herbland. Fertile. PRESERVED_SPECIMEN imada, c.t. collector number: 98-17 Maui BISH 10/6/1998
219 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Honolulu PRESERVED_SPECIMEN Stokes, J.F.G. Oahu BISH 5/1/1919
220 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kapaa, Kawaihau, sea level Rare on marshy flat. PRESERVED_SPECIMEN Hosaka, E.Y. 1646 Kauai BISH 10/29/1936
221 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama In ditch alond Kihei Rd., at junction with Lipoa St In brackish water in ditch along road. Found with Cyperus, Spirodela. PRESERVED_SPECIMEN Ishikawa, S. 363 Maui BISH 12/3/1973
222 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kahuku, by ditch 300 yds from ocean, leading from sugar mill to ocean On ditch bank, brackish water. PRESERVED_SPECIMEN Pearsall, G.A. 470 Oahu BISH 9/12/1959
223 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Salt Lake Water's edge. PRESERVED_SPECIMEN s.n. Oahu BISH 12/13/1903
224 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama James Campbell National Wildlife Refuge, Kahuku. In vicinity of bir-viewing gazebo Growing in standing water with Bacopa monnieri, Brachiaria mutica, Paspalum vaginatum, Echinochloa colona. PRESERVED_SPECIMEN Imada, C.T. 92-26 Oahu BISH 4/5/1992
225 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waikiki, honolulu, near the moana hotel PRESERVED_SPECIMEN wilder, g.p. Oahu BISH 10/27/1930
226 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Koko Head One chief species on the edge of a pool of mud by a spring on the N.W. corner of saline mud flats near Koko Head. PRESERVED_SPECIMEN Harris, J.A. s.n. Oahu BISH 8/26/1924
227 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Kipapa Gulch, Waipahu, near beach Mud flat. PRESERVED_SPECIMEN Hosaka, E.Y. 759 Oahu BISH 9/25/1932
228 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama hamakua marsh, kailua, mauka (mountainside) of hamakua drive Pond dominated by Schoenoplectus mucronatus (70% cover), with Ludwigia palustris, Cuphea carthagenensis, Cyperus halpan, Juncus effusus, Utricularia gibba. PRESERVED_SPECIMEN imada, c.t. engilis, a.; yuen, a.; dr.duk collector number: 92-10 Hawaii BISH 4/4/1992
229 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama wailua golf course, fourth hole water hazard PRESERVED_SPECIMEN flynn, t. collector number: 1249 Kauai BISH 10/3/1985
230 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive Growing in muck and in open water; common. PRESERVED_SPECIMEN Imada, C.T. 92-10 Oahu BISH 4/4/1992
231 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama honolulu PRESERVED_SPECIMEN stokes, j.f.g. Oahu BISH 5/1/1919
232 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama North Kona District, Makalawena, on west edge of Opaeula Pond Fairly common sedge along margin of pond, between stands of bulrush. Associated with Schoenoplectus californicus, Sesuvium portulacastrum, Cyperus laevigatus. PRESERVED_SPECIMEN van Dyke, P. s.n. Hawaii BISH 11/8/2000
233 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama tantalus PRESERVED_SPECIMEN pearsall, g.a. Hawaii US 7/24/1953
234 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kapiolani park, honolulu PRESERVED_SPECIMEN munro, g.c. collector number: 1 Oahu BISH 11/22/1948
235 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama kipapa gulch, waipahu, near beach Mud flat. PRESERVED_SPECIMEN hosaka, e.y. collector number: 759 Oahu BISH 9/25/1932
236 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Honolulu near Ala Moana Marshes. PRESERVED_SPECIMEN Neal, M.C. 1192 Oahu BISH 12/2/1939
237 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama ewa beach marina Collected from low, wet places in Batis swamp. Colonies are surrounded by Batis mats. No other vegetation in the near area. PRESERVED_SPECIMEN funk, e.j. lee, m.; b.ching collector number: 604 Oahu BISH 5/1/1990
238 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waikiki swamp PRESERVED_SPECIMEN Hitchcock, A.S. 13808 Oahu BISH 6/19/1916
239 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Waipio Peninsula; Waipio, Pouhala Marsh between Kapakahi & Waikele Streams On margin of open water & growing in assoc. w/Bacopa, Eleocharis, Cyperus laevigatus, Batis. Occasional. PRESERVED_SPECIMEN Imada, C.T. 96-5 Oahu BISH 12/18/1996
240 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama s.e. molokai In Batis-covered depression back of ocean. PRESERVED_SPECIMEN herbst, d.r. collector number: 1896 Molokai BISH 8/8/1970
241 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Wailua Golf Course, Fourth hole water hazard PRESERVED_SPECIMEN Flynn, T. 1249 Kauai BISH 10/3/1985
242 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama waialua PRESERVED_SPECIMEN rock, j.f.c. collector number: 8758 Oahu BISH 5/21/2011
243 Bolboschoenus maritimus subsp. paludosus (A.Nelson) T.Koyama Halalii Lake, S of edge of dry lake bed PreservedSpecimen St. John, H. Tuthill, L.D. Collector Number: 22756 Niihau BISH Specimen 8/12/1947
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