Description
Key Characters:
Growth Form:
Erect shrubs 1.5–3 m tall.
Stems:
Branches pubescent with erect and ± appressed hairs.
Roots:
Leaves:
Leaves simple.
Alternate.
Blades elliptic, elliptic–ovate, or ovate, 7–15 cm long, 3–5.5 cm wide.
Apex long-acuminate or acute.
Base cuneate to rounded.
Upper surfaces sparsely silky pubescent, the hairs somewhat spreading or appressed, 0.2–0.4 mm long; lower surfaces moderately silky pubescent, the hairs appressed or somewhat spreading, 0.3–0.6 mm long, their tips primarily directed toward apex and margins, grayish to greenish on lower surface, thin, with disciform cystoliths.
Margins coarsely dentate above the middle or entire.
Palmately 3-veined from base.
Petioles 0.8–3 cm long, pubescent with appressed and spreading hairs.
Stipules small, intrapetiolar.
Flowers:
Flowers in axillary clusters, staminate flowers on pedicels 0–1 mm long; pistillate flowers sessile.
Flowers unisexual (and the plants dioecious), actinomorphic.
Calyx of staminate flowers silky pubescent, the hairs ascending, lobes 4, boat-shaped, the lobes 2–3.5 mm long, coherent in bud, separating at anthesis, pilose; pistillate Calyx conspicuously angled, silky pubescent, the hairs ascending to erect, also with some short, erect, uncinate hairs, apex 4-toothed, the teeth acuminate to long–acuminate.
Corolla (petals) absent.
Stamens 4 in staminate flowers, staminal filaments folded elastically in bud, reflexed when pollen is shed, strap–shaped, 2–5 mm long; anthers reniform, dithecal, opening by longitudinal slits.
Pollen white.
Ovary superior (pistillate flowers), pseudomonomerous, 1-celled; ovule 1, basal; style 1; stigma 2–8 mm long, lacking receptive hairs on 1 surface. Staminate flower ovary vestigial and sterile.
Fruit:
Achenes 1.5–2 mm long; angled; surrounded by the conspicuously angled and ridged; fleshy calyx; apical portion conical; not separated from basal portion by a constriction.
Seeds 1 per achene; ovoid.
Ploidy:
Habitat:
Diverse mesic forest.
Elevation Range:
450–825 m.