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Zanthoxylum kauaense

A.Gray (1854)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Zanthoxylum

a‘e, ae, hea‘e, mānele, zanthoxylum

Description

Key Characters:

Growth Form: Small to medium-sized trees (1–)3–15 m tall.

Stems: Trunk to 40–50 cm in diameter, bark brown to grayish brown, new growth glabrous or finely puberulent, young branches subtended by a pair of bud scales.

Roots:

Leaves: Leaves compound. Alternate. Leaflets ovate to elliptic, sometimes broadly elliptic, broadly elliptic–ovate, or suborbicular, rarely lanceolate, lateral ones usually asymmetrical; leaflets 3–7(–11), thick,4–19(–21) cm long, 2–10(–14) cm wide, lateral petiolules articulated only at base, 0–7(–10) mm long, terminal petiolule once or twice articulated, at base and ± near leaflet, 10–75 mm long. Apex obtuse to rounded, sometimes bluntly acute, rarely acute or weakly acuminate. Base cuneate to rounded, subtruncate, or subcordate, adaxial side ± terminating higher. Surfaces glabrous or sometimes finely puberulent, and then more densely so when young, glandular punctate; Blades coriaceous. Margins entire or sometimes minutely crenulate. Lateral veins 5–10 pairs, spreading, higher order venation forming a conspicuous reticulate pattern. Rachis terete (cylindrical) or winged. Stipules absent.

Flowers: Flowers usually 15–150 or more in axillary, open, cymose inflorescences usually 8–15 cm long, glabrous or finely puberulent, arising at beginning of new growth on stem, and stem continuing vegetatively, peduncles 10–70 mm long, pedicels 1–5 mm long, subtended by a pair of bud scales; bracteoles ca. 1 mm long, usually caducous. Flowers unisexual, actinomorphic. Calyx of 4–5 sepals; sepals deltate, 0.5–1 mm long, ± finely puberulent, ciliolate. Corolla of 4–5 petals, white, sometimes pink or red-tinged, oblong–ovate to oblong–elliptic or oblong–lanceolate, 4–5 mm long. Stamens (3)4–5, alternate with the petals, glabrous, absent in pistillate flowers; anthers medifixed, dithecal. Ovary superior, 1-carpellate; ovule placentation axile, rudimentary in staminate flowers; style oblique; stigma capitate.

Fruit: Follicles 1; falcate; obovoid; 8–12(–15) mm long; apex rounded; exocarp rugulose and pitted. Seeds 1 per fruit; on a slender funicle; black; subglobose; compressed; 7–13 mm in diameter.

Ploidy:

Habitat: Mesic forest; sometimes in dry to wet forest.

Elevation Range: 300–1,980 m.

Historical Distribution

Synonyms (49)

Fagara glandulosa (Hillebr.) Engl. (1896)
Fagara glandulosa var. hiloensis Sherff (1958)
Fagara kauaense (A.Gray) Engl. (1896)
Fagara kauaense var. kohalana Sherff (1958)
Fagara kauaense var. kohuana Skottsb. (1944)
Fagara kauaense var. tenuifolia O.Deg. & Sherff (1958)
Fagara kauaensis (A.Gray) Engl. (1896)
Fagara kauaensis var. kohalana Sherff (1958)
Fagara kauaensis var. tenuifolia O.Deg. & Sherff (1958)
Fagara kauaiensis var. tenuifolia Deg. & Sherff
Fagara mauiensis (H.Mann) Engl. (1896)
Fagara mauiensis f. glabra (Rock) Skottsb.
Fagara mauiensis var. anceps (Rock) Skottsb. (1958)
Fagara mauiensis var. cranwelliae Skottsb. (1944)
Fagara mauiensis var. kaalana Sherff (1958)
Fagara mauiensis var. lanaiensis Sherff (1958)
Fagara mauiensis var. maunahuiensis Sherff (1958)
Fagara mauiensis var. rigida (Rock) Sherff (1958)
Fagara maviensis
Fagara maviensis f. petiolulata (Rock) H.St.John (1945)
Fagara maviensis var. anceps (Rock) H.St.John (1945)
Fagara maviensis var. cranwelliae
Fagara multiflora Engl. (1896)
Fagara multifoliolata O.Deg. & Skottsb. (1936)
Fagara semiarticulata (H.St.John & Hosaka) O.Deg. (1936)
Fagara semiarticulata f. laiense O.Deg. (1936)
Fagara semiarticulata var. sessilis O.Deg. (1936)
Fagara skottsbergii O.Deg. & I.Deg. (1960)
Zanthoxylum glandulosum Hillebr. (1888)
Zanthoxylum hillebrandii P.G. Waterman (1975)
Zanthoxylum hillebrandii var. hiloense (Sherff) H.St.John (1976)
Zanthoxylum kauaense var. kohalanum (Sherff) H.St.John (1973)
Zanthoxylum kauaense var. kohuanum (Skottsb.) H.St.John (1973)
Zanthoxylum kauaense var. tenuifolium (O.Deg. & Sherff) H.St.John (1976)
Zanthoxylum maviense H.Mann (1866)
Zanthoxylum maviense f. glabrum Rock (1913)
Zanthoxylum maviense f. petiolulatum Rock (1913)
Zanthoxylum maviense var. anceps Rock (1913)
Zanthoxylum maviense var. cranwelliae (Skottsb.) H.St.John (1973)
Zanthoxylum maviense var. kaalanum (Sherff) H.St.John (1976)
Zanthoxylum maviense var. lanaiense (Sherff) H.St.John (1976)
Zanthoxylum maviense var. maunahuiense (Sherff) H.St.John (1976)
Zanthoxylum maviense var. rigidum Rock (1913)
Zanthoxylum molokaiense B.C.Stone (1985)
Zanthoxylum semiarticulatum H.St.John & Hosaka (1932)
Zanthoxylum semiarticulatum f. laiense (O.Deg.) Fosberg (1969)
Zanthoxylum semiarticulatum var. sessile Fosberg (1969)
Zanthoxylum semiarticulatum var. sessilis (O.Deg.) Fosberg
Zanthoxylum skottsbergii (O.Deg. & I.Deg.) H.St.John (1973)

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: U.S. Expl. Exped., Phan. 1: 354 (1854)

Other References

Wagner et al. 1990:1215 (K, O, Mo, L, M, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Zanthoxylum kauaense A.Gray unknown dryland forest PreservedSpecimen Christensen, C. Collector Number: 199 BISH Specimen 5/7/1977
2 Zanthoxylum kauaense A.Gray unknown Hidden in mamane trees. PreservedSpecimen Fagerlund, G.O. Mitchell, A.L. Collector Number: 1016 BISH Specimen 3/24/1945
3 Zanthoxylum kauaense A.Gray unknown PreservedSpecimen Takeuchi, W.N. Collector Number: 3435 BISH Specimen 2/22/1987
4 Zanthoxylum kauaense A.Gray unknown in remnant patch of intact forest. with Alectryon, Pleomele forbesii, Phyllostegia hirsuta PreservedSpecimen Takeuchi, W.N. Paquin, D. Collector Number: 2814 BISH Specimen 9/6/1986
5 Zanthoxylum kauaense A.Gray unknown forest, with Syzygium, Myrsine, Elaeocarpus PreservedSpecimen Takeuchi, W.N. Collector Number: 2818 BISH Specimen 9/7/1986
6 Zanthoxylum kauaense A.Gray unknown PreservedSpecimen Takeuchi, W.N. Shimabukoro, C. Collector Number: 2819 BISH Specimen 9/7/1986
7 Zanthoxylum kauaense A.Gray West Mau/Molokai PreservedSpecimen Hillebrand, W.B. Lydgate, J.M. Collector Number: s.n. BISH Specimen
8 Zanthoxylum kauaense A.Gray ridge W of Kawaikoa Stream dry lower edge of forest PRESERVED_SPECIMEN St. John, H. 22700 Kauai BISH 8/10/1947
9 Zanthoxylum kauaense A.Gray Maunahui PreservedSpecimen Hillebrand, W.B. Collector Number: s.n. Molokai BISH Specimen
10 Zanthoxylum kauaense A.Gray Kanehoa ridge Metrosideros, Pelea, Psychotria PRESERVED_SPECIMEN Pyle, L. Takeuchi 2029 Oahu BISH 3/3/1985
11 Zanthoxylum kauaense A.Gray Kokee, Honopu Trail PRESERVED_SPECIMEN Herbst, D.R. 2158 Kauai BISH 9/22/1971
12 Zanthoxylum kauaense A.Gray palikea trail PRESERVED_SPECIMEN bennett, t. collector number: prc 223 Oahu BISH 3/11/1981
13 Zanthoxylum kauaense A.Gray west maui, lahainaluna PRESERVED_SPECIMEN hillebrand, w.b. collector number: s.n. Maui BISH
14 Zanthoxylum kauaense A.Gray s slope of haleakala, on ridge between e&w forks of pahihi gulch Dry exotic grassland with native and exotic shrubs and scattered native trees PRESERVED_SPECIMEN warshauer, f.r. mceldowney, p.h. collector number: frw 3091 Maui BISH 8/20/1980
15 Zanthoxylum kauaense A.Gray East Maui, Makawao District, Makawao FR, Kahakapao drainage basin Growing on windward facing slope in mesic forest. Associated vegetation: Acacia, Metrosideros, Dicranopteris, Psychotria, Melicope, Pleomele, Nestegis, Pouteria, Clermontia, Fraxinus, Rubus. PRESERVED_SPECIMEN Oppenheimer, H.L. H70606 Maui BISH 7/12/2006
16 Zanthoxylum kauaense A.Gray waianae mts., trail on e ridge of puu kaua aa lava, waste PRESERVED_SPECIMEN marr, k.l. collector number: 22 Hawaii BISH 4/17/1988
17 Zanthoxylum kauaense A.Gray North Kona District: Puuwaawaa, Waiho. 1/2 mile NNW of Hibiscadelphus grove Hidden in mamane trees. PRESERVED_SPECIMEN Fagerlund, G.O. 1016 Hawaii BISH 3/24/1945
18 Zanthoxylum kauaense A.Gray Awaawapuhi trail, Honopu rainforest PRESERVED_SPECIMEN Woolford, H.A. 430 Kauai BISH 12/29/1956
19 Zanthoxylum kauaense A.Gray Hono O Na Pali NAR, Pohakea region, steep ridge above Hanakapiai Open Metrosideros mesic forest with adjacent transitional wet forest. W/ Kadua affinis, Dicranopteris, Myrsine linearifolia, Scaevola procera, Xylosma hawaiiense, Cyrtandra paludosa, Lysimachia sp, Santalum pyrularium, Dubautia laevigata, Psychotria mariniana, Coprosma PRESERVED_SPECIMEN Wood, K.R. Query, M. Collector Number: 14728 Kauai BISH 7/28/2011
20 Zanthoxylum kauaense A.Gray puhielelu ridge, n PRESERVED_SPECIMEN flynn, t. collector number: 206 Hawaii BISH 5/26/1982
21 Zanthoxylum kauaense A.Gray kawaiiki ridge and nearby drainages to north PRESERVED_SPECIMEN wood, k.r. aguraiuja r. collector number: 15405 Maui BISH 2/15/2013
22 Zanthoxylum kauaense A.Gray w waimea drainage basin PRESERVED_SPECIMEN forbes, c.n. collector number: 1159 k Kauai BISH 7/3/1917
23 Zanthoxylum kauaense A.Gray koele side of summit ridge, along munro trail Metrosideros polymorpha, Dicranopteris mats, Broussaisia arguta, Scaevola chamissoniana, Pipturus rockii, Pittosporum confertiflorum PRESERVED_SPECIMEN carlquist, s. collector number: 2015 Lanai BISH 7/16/1966
24 Zanthoxylum kauaense A.Gray waianae mountains. puu kaua, beside trail from crest right between puu kaua and pohakea pass, third minor ridge from summit of puu kaua Below small stand of Sophora. Close to individual Zanthoxylum dipetalum with strong anixe odor. PRESERVED_SPECIMEN marr, k.l. palmer, d.d. collector number: km55 Oahu BISH 6/24/1988
25 Zanthoxylum kauaense A.Gray honouliuli contour mauka slopes; mesic forest PRESERVED_SPECIMEN takeuchi, w.n. tate, b. collector number: 3579 Oahu BISH 5/31/1987
26 Zanthoxylum kauaense A.Gray papoaea, n slope of haleakala PRESERVED_SPECIMEN forbes, c.n. collector number: 2519,m Maui BISH 6/14/1920
27 Zanthoxylum kauaense A.Gray waimea canyon, toward head from puu kapele PRESERVED_SPECIMEN christensen, c. collector number: 199 Kauai BISH 5/7/1977
28 Zanthoxylum kauaense A.Gray Honouliuli Contour mesic slopes, with Planchonella PRESERVED_SPECIMEN Takeuchi, W.N. s.n. Oahu BISH 2/22/1987
29 Zanthoxylum kauaense A.Gray Waianae Mts., Makaha Valley PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 1909-00-00
30 Zanthoxylum kauaense A.Gray kalalau valley, rim forest PRESERVED_SPECIMEN degener, o. degener, i. collector number: 28102 Kauai BISH 3/14/1961
31 Zanthoxylum kauaense A.Gray SW ridge of Kaiholena Gulch, Kamoku PRESERVED_SPECIMEN St.John, H.; R.Cowan BISH 4/4/1947
32 Zanthoxylum kauaense A.Gray north Honouliuli mesic slopes PRESERVED_SPECIMEN Takeuchi, W.N. 3755 Oahu BISH 1/10/1988
33 Zanthoxylum kauaense A.Gray Kalalau Valley, rim forest PRESERVED_SPECIMEN Degener, O. 28102 Kauai BISH 3/14/1961
34 Zanthoxylum kauaense A.Gray West Maui, Lahainaluna PRESERVED_SPECIMEN Hillebrand, W.B. s.n. Maui BISH
35 Zanthoxylum kauaense A.Gray waianae mts., palikea near top of gulley below summit; with Pittosporum, Metrosideros, Dubautia, Lobelia yuccoides PRESERVED_SPECIMEN marr, k.l. palmer, d.d.; nakamura, j. collector number: 65 Oahu BISH 6/26/1988
36 Zanthoxylum kauaense A.Gray Waianae Range, Palehua In decadent, pastured forest PRESERVED_SPECIMEN Degener, O. 10318 Oahu BISH 11/23/1935
37 Zanthoxylum kauaense A.Gray S.slope of Hauula Ridge, 10 ft. below trail, (Kaipapau Forest Reserve) PRESERVED_SPECIMEN Suehiro, A. s.n. Oahu BISH 10/15/1933
38 Zanthoxylum kauaense A.Gray Waimea District. Kawaiiki region, north facing slopes below dividing ridge above twin falls. Metrosideros-Acacia mesic forest with Myrsine lanaiensis, Myoporum sandwicensis, Kadua affinis, Xylosma hawaiiense, Planchonella sandwicensis, Ilex anomala, Perrottetia sandwicensis, Charpentiera elliptica, Chrysodracon aurea, Elaeocar PRESERVED_SPECIMEN Wood, K.R. 17169 Kauai BISH 11/1/2016
39 Zanthoxylum kauaense A.Gray E of Kalalau Lookout forest PRESERVED_SPECIMEN Degener, O. 20600 Kauai BISH 1/31/1950
40 Zanthoxylum kauaense A.Gray schofield, wahiwa, waianae mts PRESERVED_SPECIMEN warshauer, f.r. collector number: 1072a Oahu BISH 2/21/1977
41 Zanthoxylum kauaense A.Gray Growing on edge of bog. Assoc. with Coprosma, Myrsine, Styphelia, Viola maviensis, Astelia, Peperomia and Smilax. PRESERVED_SPECIMEN wilkes expedition Molokai BISH
42 Zanthoxylum kauaense A.Gray Pupukea- Kahuku Trail, Koolau Mts wooded ridge PRESERVED_SPECIMEN St. John, H. 11568 Oahu BISH 3/6/1932
43 Zanthoxylum kauaense A.Gray ridge flute near kamakalii growing in planted stand of Eucalyptus with Psidium cattleianum understory. PRESERVED_SPECIMEN takeuchi, w.n. shimabukoro, c. collector number: 2826 Hawaii BISH 9/1/1986
44 Zanthoxylum kauaense A.Gray east maui, east kaupo gap within dry forest exclusion in haleakala national park South slope cliffs. PRESERVED_SPECIMEN medeiros, a.c. collector number: 214 Maui BISH 7/10/1982
45 Zanthoxylum kauaense A.Gray Puna DISTR. 'Olaa AHU. Tr 32(95/96); E slope of Mauna Loa near Volcano dump; N of Hwy 11 Wet 'ohi'a forest with other native trees and a treefern and native shrub understory. PRESERVED_SPECIMEN Warshauer, F.R. FRW 1256 Hawaii BISH 7/23/1977
46 Zanthoxylum kauaense A.Gray Kipukaki, near Kilauea PRESERVED_SPECIMEN Forbes, C.N. 403 H Hawaii BISH 8/4/1911
47 Zanthoxylum kauaense A.Gray Waianae Palehua PRESERVED_SPECIMEN Skottsberg, C.J.F. 318 Oahu BISH 8/23/1922
48 Zanthoxylum kauaense A.Gray waianae range, w of camp kunia with Pelea, Gouldia, Pittosporum PRESERVED_SPECIMEN wuotila, j.l. Oahu BISH 12/18/1958
49 Zanthoxylum kauaense A.Gray waimea canyon, toward head from puu kapele dry forest PRESERVED_SPECIMEN christensen, c. collector number: 199 Kauai BISH 5/7/1977
50 Zanthoxylum kauaense A.Gray land section of makakupaia, kupaia gulch Associates: Antidesma, Pisonia, Ilex, Athyrium, Metrosideros, Coprosma PRESERVED_SPECIMEN lau, j. collector number: 1451 Molokai BISH 9/24/1985
51 Zanthoxylum kauaense A.Gray middle Palawai Ridge forest PRESERVED_SPECIMEN Degener, O. 10863 Oahu BISH
52 Zanthoxylum kauaense A.Gray Honouliuli, Kaaikukae PRESERVED_SPECIMEN Russ, G.W. s.n. Oahu BISH 10/19/1936
53 Zanthoxylum kauaense A.Gray waianae mts., n branch of kaulaa gulch, s of puu hapapa E facing slope at head of side gulch PRESERVED_SPECIMEN marr, k.l. takeuchi, w.t.; palmer, d.d.; pyle, l. collector number: 3 Oahu BISH 1/10/1988
54 Zanthoxylum kauaense A.Gray waianae slope of kaala above makaleha PRESERVED_SPECIMEN skottsberg, c.j.f. collector number: 376 Kauai BISH 8/30/1922
55 Zanthoxylum kauaense A.Gray Kaala Mt PRESERVED_SPECIMEN Hillebrand, W.B. s.n. Oahu BISH 1861-00-00
56 Zanthoxylum kauaense A.Gray near summit of pupukea-kahuku trail wet cliff PRESERVED_SPECIMEN degener, o. park, k.k.; kwon, m. collector number: 4109 Oahu BISH 12/6/1931
57 Zanthoxylum kauaense A.Gray Waimea District. Makaha, south facing slopes below Milolii road. Acacia-Metrosideros lowland mesic forest and steep slopes of Dicranopteris with Alphitonia ponderosa, Kadua affinis, Planchonella sandwicensis, Caloxylon sandwicensis, Coprosma waimeae, Polyscias kavaiensis, Melicope ovata, Melicope barbigera, Dodonaea viscosa, Nestegis sandwicensis. PRESERVED_SPECIMEN Wood, K.R. 17084 Kauai BISH 7/28/2016
58 Zanthoxylum kauaense A.Gray Waianae Mts., N branch of Kaulaa Gulch, S of Puu Hapapa E facing slope at head of side gulch PRESERVED_SPECIMEN Marr, K.L. 3 Oahu BISH 1/10/1988
59 Zanthoxylum kauaense A.Gray kalalau trail PRESERVED_SPECIMEN forbes, c.n. collector number: 1091.k Kauai BISH 7/3/1917
60 Zanthoxylum kauaense A.Gray Waianae Mts., Palikea Trail protected hillside PRESERVED_SPECIMEN Crosby, M.R. 1918 Oahu BISH 7/31/1964
61 Zanthoxylum kauaense A.Gray ekahanui-kaua among boulder rocks; in Pelea PRESERVED_SPECIMEN takeuchi, w.n. greer; tate collector number: 3554 Oahu BISH 5/16/1987
62 Zanthoxylum kauaense A.Gray Waimea District, Nuololo, central branch Metrosideros-Acacia mesic forest with Alphitonia ponderosa, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bobea brevipes, Melicope ovata, M. barbigera PRESERVED_SPECIMEN Wood, K.R. 15042 Kauai BISH 6/20/2012
63 Zanthoxylum kauaense A.Gray Olaa Forest Preserve, north of Volcanoe dump PRESERVED_SPECIMEN Marr, M. KM 31 Hawaii BISH 5/31/1988
64 Zanthoxylum kauaense A.Gray Nuololo, north facing slopes above drainage Metrosideros-Acacia montane mesic forest, 40-50% open, up to 20 m tall canopy, W/ Alphitonia, Pouteria, Xylosma hawaiiensis, Claoxylon, Myrsine lanaiensis, Wikstroemia furcata, Dodonaea, Kadua affinis, Melicope ovata, Pleomele, Tetrplasandra kavaiensis, Psychotria greenwelliae PRESERVED_SPECIMEN Wood, K.R. 14541 Kauai BISH 2/28/2011
65 Zanthoxylum kauaense A.Gray ridge S of S. Opaeula Gulch, Paalaa, Koolau Mts wooded PRESERVED_SPECIMEN St. John, H. 11110 Oahu BISH 6/28/1931
66 Zanthoxylum kauaense A.Gray Waianae Mts., Palikea trail, gulley below summit with Cryptomeria, Schinus PRESERVED_SPECIMEN Marr, K.L. 60 Oahu BISH 6/26/1988
67 Zanthoxylum kauaense A.Gray koele, mountains near PRESERVED_SPECIMEN forbes, c.n. collector number: 209.l Oahu BISH 6/1/1913
68 Zanthoxylum kauaense A.Gray East Maui, East Kaupo Gap within dry forest exclusion in Haleakala National Park South slope cliffs. PRESERVED_SPECIMEN Medeiros, A.C. 214 Maui BISH 7/10/1982
69 Zanthoxylum kauaense A.Gray west maui, maunahuoma PRESERVED_SPECIMEN forbes, c.n. caum, e.l. Maui BISH 5/1/1910
70 Zanthoxylum kauaense A.Gray waimea PRESERVED_SPECIMEN knudsen collector number: 146 Kauai BISH
71 Zanthoxylum kauaense A.Gray near pohakea, at the divide in rock crevices; with Schinus PRESERVED_SPECIMEN takeuchi, w.n. paquin, d. Oahu BISH 8/8/1983
72 Zanthoxylum kauaense A.Gray between puu manawahua and mauna kapu, nanakuli side PRESERVED_SPECIMEN degener, o. greenwell, a.; hatheway, w.h.; caindec collector number: 20847 Hawaii BISH 6/28/1950
73 Zanthoxylum kauaense A.Gray Waianae Mts., along ridge extending downward from Puu Hapapa. Entomological collection area code: P-23 Associated with Dracaena and Tetraplasandra. PRESERVED_SPECIMEN Gagne, W.C. s.n. Oahu BISH 4/26/1970
74 Zanthoxylum kauaense A.Gray East Maui, Haleakala, E. Kaupo gap steep ridge with dryland forest PRESERVED_SPECIMEN Crivellone, C. s.n. Maui BISH 6/10/1983
75 Zanthoxylum kauaense A.Gray Makawao, Haleakala PRESERVED_SPECIMEN Rock, J.F.C. 10207 Maui BISH 3/1/1912
76 Zanthoxylum kauaense A.Gray Awehi, E Lanai PRESERVED_SPECIMEN Montgomery, S.L. s.n. Lanai BISH 6/6/1971
77 Zanthoxylum kauaense A.Gray Waianae Mts., W side of main ridge separating Makaha and Waianae Kai Valleys, N of Puu Kawiwi PRESERVED_SPECIMEN Herbst, D.R. 5044b Oahu BISH 9/21/1974
78 Zanthoxylum kauaense A.Gray north palawai gulch just north of palikea, southern waianae range Under a canopy of taller trees, sloping terrain, mesic-summer, dry habitat; three trees, voucher pistillate PRESERVED_SPECIMEN obata, j.k. palmer, d. collector number: 427 Oahu BISH 3/30/1980
79 Zanthoxylum kauaense A.Gray Kokee forest PRESERVED_SPECIMEN Degener, O. 28098 Kauai BISH 11/19/1960
80 Zanthoxylum kauaense A.Gray Hilo PRESERVED_SPECIMEN Lydgate, J.M. Collector Number: s.n. Hawaii BISH
81 Zanthoxylum kauaense A.Gray upper awaawapuhi In wet forest dominated by uluhe, ohia, manono, pilo PRESERVED_SPECIMEN takeuchi, w.n. pyle, l. collector number: 3689 Kauai BISH 10/25/1987
82 Zanthoxylum kauaense A.Gray Waianae Mts., Mokuleia, Makaleha Trail Metrosideros and Pelea forest, lateral ridge PRESERVED_SPECIMEN St. John, H. 25284 Oahu BISH 2/28/1954
83 Zanthoxylum kauaense A.Gray waianae range, south of palikea Moderately dry slope near the summit ridge PRESERVED_SPECIMEN degener, o. potter; topping collector number: 10159 Oahu BISH 1/5/1936
84 Zanthoxylum kauaense A.Gray Kanehoa ridge, 300' below summit PRESERVED_SPECIMEN Pyle, L. 2029 Oahu BISH 3/3/1985
85 Zanthoxylum kauaense A.Gray Kapua Gulch, S branch forest PRESERVED_SPECIMEN Degener, O. 28530 Kauai BISH 8/25/1963
86 Zanthoxylum kauaense A.Gray puaulu, kilaueu PRESERVED_SPECIMEN rock, j.f.c. collector number: 10202 Hawaii BISH 7/1/1911
87 Zanthoxylum kauaense A.Gray not far from summit of kunia trail, which ends on summit ridge of waianae mts. between puu kanehoa and puu hapapa On steep slopes in scrubby Metrosideros polymorpha forest, with Styphelia tameiameiae, Antidesma, Diospyros, Alyxia oliviformis, Pelea spp. PRESERVED_SPECIMEN carlquist, s. collector number: 1903 Oahu BISH 7/2/1966
88 Zanthoxylum kauaense A.Gray Puuwaawaa PRESERVED_SPECIMEN Rock, J.F.C. 3716 Hawaii BISH 6/15/1909
89 Zanthoxylum kauaense A.Gray unknown PRESERVED_SPECIMEN takeuchi, w.n. collector number: 3435 Oahu BISH
90 Zanthoxylum kauaense A.Gray awaawapuhi trail, honopu Moist Metrosideros woods PRESERVED_SPECIMEN st.john, h. collector number: 26012 Kauai BISH 12/29/1956
91 Zanthoxylum kauaense A.Gray E slope of Puu Kaua woods PRESERVED_SPECIMEN Degener, O. 28083 Oahu BISH 8/29/1962
92 Zanthoxylum kauaense A.Gray Waimea Canyon PRESERVED_SPECIMEN Christensen, C. 194 Kauai BISH 4/27/1977
93 Zanthoxylum kauaense A.Gray Makaleha PRESERVED_SPECIMEN Russ, G.W. s.n. Oahu BISH 10/1/1928
94 Zanthoxylum kauaense A.Gray ekahanui gulch, ascending ridge to puu kaua, ridge top, at junction of last adjoining sub-ridge to the ascending ridge associated with Psychotria, Pelea, Koa, etc. PRESERVED_SPECIMEN obata, j.k. pyle, l. collector number: s.n. Oahu BISH 12/23/1984
95 Zanthoxylum kauaense A.Gray Makaha Valley, N of Puu Kawiwi, Kamaileunu Ridge, Waianae Mts crest of minor spur; with Schinus, Acacia, Diospyros PRESERVED_SPECIMEN Nagata, K.M. 1101 Oahu BISH 12/28/1972
96 Zanthoxylum kauaense A.Gray Hawaii PRESERVED_SPECIMEN Remy, M.J. 614 Hawaii BISH 1/24/1905
97 Zanthoxylum kauaense A.Gray Waianae Mtns. Kaluaa Gulch. Same site as KM 2, 3, 69-71, 73 Other species: Metrosideros, Streblus, Morinda trimera, Antidesma, Acacia koa, Syzygium sandwicensis, Nothocestrum, Alectryon macrococcum, Charpentiera obovata, Pisonia spp, Pipturus albidus, Urera glabra, Pelea spp, Platydesma cornuta. PRESERVED_SPECIMEN Marr, K.L. KM 68 Oahu BISH 7/17/1988
98 Zanthoxylum kauaense A.Gray pupukea fr, koolauloa Metrosideros forest PRESERVED_SPECIMEN grant, m.l. collector number: 7157 Oahu BISH 8/1/1934
99 Zanthoxylum kauaense A.Gray auhi, ulupalakua rocky hillside PRESERVED_SPECIMEN munro, g.c. collector number: 406 Oahu BISH 2/12/1915
100 Zanthoxylum kauaense A.Gray laie-malaekahana ridge, koolau mts woods PRESERVED_SPECIMEN st.john, h. collector number: 11533 Oahu BISH 2/14/1932
101 Zanthoxylum kauaense A.Gray e slope of puu kaua woods PRESERVED_SPECIMEN degener, o. obata, j. collector number: 28083 Oahu BISH 8/29/1962
102 Zanthoxylum kauaense A.Gray Puaulu, Kilaueu PRESERVED_SPECIMEN Rock, J.F.C. 10202 Hawaii BISH 7/1/1911
103 Zanthoxylum kauaense A.Gray Kipuka near kilauea PRESERVED_SPECIMEN Forbes, C.N. 403 H Hawaii BISH 12/4/1911
104 Zanthoxylum kauaense A.Gray Waianae Mtns. Palikea. Bottom of gulch leading to summit in Cryptomeria stand PRESERVED_SPECIMEN Marr, K.L. KM 59 Oahu BISH 6/26/1988
105 Zanthoxylum kauaense A.Gray Along highest ridge overlooking Kahuku along Pupukea-Kahuku Trail In rainforest PRESERVED_SPECIMEN Degener, O. 4109 Oahu BISH 12/6/1931
106 Zanthoxylum kauaense A.Gray south opaeula gulch, paala, koolau mts., wooded south ridge PRESERVED_SPECIMEN st.john, h. collector number: 11089 Oahu BISH 5/3/1931
107 Zanthoxylum kauaense A.Gray Koolau Range, summit above Kahuku PRESERVED_SPECIMEN Rock, J.F.C. 25765 Oahu BISH 4/3/1957
108 Zanthoxylum kauaense A.Gray lanai PRESERVED_SPECIMEN munro, g.c. collector number: 439 Lanai BISH 4/29/1915
109 Zanthoxylum kauaense A.Gray makaleha PRESERVED_SPECIMEN russ, g.w. Oahu BISH 10/1/1928
110 Zanthoxylum kauaense A.Gray Kanehoa, East side PRESERVED_SPECIMEN Russ, G.W. s.n. Oahu BISH 6/19/1930
111 Zanthoxylum kauaense A.Gray Waianae Range, Honouliuli, Puu Kanehoa dry wooded ridge PRESERVED_SPECIMEN Webster, G.L. 1630 Oahu BISH 6/13/1948
112 Zanthoxylum kauaense A.Gray waianae kai. on saddle ridge near poles with koa, ohia, Grevillea, Dodonaea, maile, pilo, Scaevola PRESERVED_SPECIMEN perlman, s.p. collector number: 5047 Oahu BISH 1/8/1987
113 Zanthoxylum kauaense A.Gray makaha valley, n of puu kawiwi, kamaileunu ridge, waianae mts crest of minor spur; with Schinus, Acacia, Diospyros PRESERVED_SPECIMEN nagata, k.m. spence, g. collector number: 1101 Oahu BISH 12/28/1972
114 Zanthoxylum kauaense A.Gray Waianae Mtns. North branch of Kaluaa Gulch, at head of side gulch, south of Puu Hapapa PRESERVED_SPECIMEN Marr, K.L. KM 2 Oahu BISH 1/10/1988
115 Zanthoxylum kauaense A.Gray kokee PRESERVED_SPECIMEN degener, o. degener, i. collector number: 28098 Oahu BISH 11/19/1960
116 Zanthoxylum kauaense A.Gray Kepuka Puaulu. PRESERVED_SPECIMEN Russ, G.W. s.n. Hawaii BISH 8/1/1927
117 Zanthoxylum kauaense A.Gray mts. near koele PRESERVED_SPECIMEN forbes, c.n. collector number: 37.l Lanai BISH 6/1/1913
118 Zanthoxylum kauaense A.Gray ridge s of s. opaeula gulch, paalaa, koolau mts Dwarf shrub & grass community in fine ash pockets. PRESERVED_SPECIMEN st.john, h. collector number: 11523 Hawaii BISH 1/17/1932
119 Zanthoxylum kauaense A.Gray Lahainaluna PRESERVED_SPECIMEN Hillebrand, W.B. s.n. Maui BISH
120 Zanthoxylum kauaense A.Gray lateral ridge, r side of olowalu valley PRESERVED_SPECIMEN forbes, c.n. collector number: 2255.m Maui BISH 5/7/1920
121 Zanthoxylum kauaense A.Gray bird park kipuka puaulu moderate forest PRESERVED_SPECIMEN morley, h. collector number: s.n. Oahu BISH 8/11/1934
122 Zanthoxylum kauaense A.Gray Lihue Distric; Top of Mt. Haupu; Metrosideros Lowland Mixed Mesic Forest with North facing Native cliffs; Associated Plants: Bidens valida, Artemisia, Sida fallax, Hedyotis acuminata & H. fluviatilis, Bonamia, Isodendrion, Zanthoxylum, Carex wahuensis, Eragrotis variabilis, Hibiscus kokio, Lobelia niihauensis, Machaerina, Pipturus, Lepidium bidentatum, Mariscus, Melicope anisata. PRESERVED_SPECIMEN Wood, K.R. 2037 Kauai BISH 7/27/1992
123 Zanthoxylum kauaense A.Gray waianae mts., near halona valley PRESERVED_SPECIMEN welch, d'a. Oahu BISH 6/4/1936
124 Zanthoxylum kauaense A.Gray nuololo, north facing slopes above drainage Metrosideros-Acacia montane mesic forest, 40-50% open, up to 20 m tall canopy, W/ Alphitonia, Pouteria, Xylosma hawaiiensis, Claoxylon, Myrsine lanaiensis, Wikstroemia furcata, Dodonaea, Kadua affinis, Melicope ovata, Pleomele, Tetrplasandra kavaiensis, Psychotria greenwelliae PRESERVED_SPECIMEN wood, k.r. query, m. collector number: 14541 Kauai BISH 2/28/2011
125 Zanthoxylum kauaense A.Gray Rim of Waimea Canyon, below road approx. 11.8 miles from Waimea. Same location as 5 Z. dipetalum individuals PRESERVED_SPECIMEN Marr, K.L. K.M. 11 Kauai BISH 3/26/1988
126 Zanthoxylum kauaense A.Gray kipuka near kilauea PRESERVED_SPECIMEN forbes, c.n. collector number: 403 h Maui BISH 12/4/1911
127 Zanthoxylum kauaense A.Gray east maui, wailou spring area,olinda on slopes, near gulch bottom PRESERVED_SPECIMEN hobdy, r.w. collector number: 795 Maui BISH 4/20/1980
128 Zanthoxylum kauaense A.Gray central ridge of Makaleha Valley forest near summit PRESERVED_SPECIMEN Degener, O. 20759 Oahu BISH 8/19/1950
129 Zanthoxylum kauaense A.Gray Haupu, talus slope, foot of cliff talus slope PRESERVED_SPECIMEN MacDaniels, L.H. 882 Kauai BISH 2/26/1927
130 Zanthoxylum kauaense A.Gray Lanai PRESERVED_SPECIMEN Munro, G.C. 439 Lanai BISH 4/29/1915
131 Zanthoxylum kauaense A.Gray w waimea drainage basin PRESERVED_SPECIMEN forbes, c.n. collector number: 826 k Oahu BISH 7/3/1917
132 Zanthoxylum kauaense A.Gray olaa forest reseve, behind volcano dump Wet forest habitat. PRESERVED_SPECIMEN marr, k.l. collector number: km 81 Hawaii BISH 7/30/1988
133 Zanthoxylum kauaense A.Gray waianae mts., dupont trail to puu kaala PRESERVED_SPECIMEN krajina, v.j. collector number: 620225340 Oahu BISH 2/25/1962
134 Zanthoxylum kauaense A.Gray Makaha Valley, on steep slope of ridge leading down into Makaha Valley from near well know Pinnacle Rocks, Waianae,Kai area, east of Kamaileunu ridge PRESERVED_SPECIMEN Perlman, S. 5058 Oahu BISH 1/8/1987
135 Zanthoxylum kauaense A.Gray land section of Makakupaia, Kupaia Gulch Associates: Antidesma, Pisonia, Ilex, Athyrium, Metrosideros, Coprosma PRESERVED_SPECIMEN Lau, J. 1451 Molokai BISH 9/24/1985
136 Zanthoxylum kauaense A.Gray East Maui, Wailou spring area,Olinda on slopes, near gulch bottom PRESERVED_SPECIMEN Hobdy, R.W. 795 Maui BISH 4/20/1980
137 Zanthoxylum kauaense A.Gray puuwaawaa PRESERVED_SPECIMEN forbes, c.n. collector number: 61.h Hawaii BISH 6/8/1911
138 Zanthoxylum kauaense A.Gray haupu, talus slope, foot of cliff talus slope PRESERVED_SPECIMEN macdaniels, l.h. collector number: 882 Kauai BISH 2/26/1927
139 Zanthoxylum kauaense A.Gray West Maui, Lahaina District, slopes of Helu, Puehuehunui, first Gulch NW of Luakoi Ridge On S facing slope. Vegetation: Metrosideros, Cheirodendron, Coprosma, Broussaisia, Clermontia, Santalum, Alyxia, Freycinetia, Microlepia, Perrottetia, Diplazium, Ilex, Sadleria. PRESERVED_SPECIMEN Oppenheimer, H.L. H50709 Maui BISH 5/7/2007
140 Zanthoxylum kauaense A.Gray Kokee. Kawaikoa Stream Trail Above trail on S. side of stream, 70 m before 1/2 mile marker, Growing between two Zanthoxylum depetalum (KM 45, 46) PRESERVED_SPECIMEN Marr, K.L. KM 47 Kauai BISH 7/15/1988
141 Zanthoxylum kauaense A.Gray palikea trail, waianae mts PRESERVED_SPECIMEN gagné, w.c. collector number: 504 Oahu BISH 4/18/1970
142 Zanthoxylum kauaense A.Gray keauhou mauka: koa niu PRESERVED_SPECIMEN herbst, d.r. collector number: 2139 Hawaii BISH 9/16/1971
143 Zanthoxylum kauaense A.Gray Koa Niu, Keauhou Mauka Remnant of a mesic forest, now with scattered trees and kikuya grass. PRESERVED_SPECIMEN Herbst, D.R. 2138 Hawaii BISH 9/16/1971
144 Zanthoxylum kauaense A.Gray Malaekahana Trail, Laie (Near No. 10083) Forested ridge half way to summit PRESERVED_SPECIMEN Degener, O. 10086 Oahu BISH 7/29/1935
145 Zanthoxylum kauaense A.Gray Koolau Mts., Pupukea-Kahuku trail ridge PRESERVED_SPECIMEN Stone, B.C. 2816 Oahu BISH 5/4/1959
146 Zanthoxylum kauaense A.Gray paalaa, koolau mts marsh PRESERVED_SPECIMEN krauss, n.h. Oahu BISH 12/10/1933
147 Zanthoxylum kauaense A.Gray waianae mtns. kaluaa gulch. same site as km 2, 3, 69-71, 73 Other species: Metrosideros, Streblus, Morinda trimera, Antidesma, Acacia koa, Syzygium sandwicensis, Nothocestrum, Alectryon macrococcum, Charpentiera obovata, Pisonia spp, Pipturus albidus, Urera glabra, Pelea spp, Platydesma cornuta. PRESERVED_SPECIMEN marr, k.l. takeuchi, w.t. collector number: km 68 Oahu BISH 7/17/1988
148 Zanthoxylum kauaense A.Gray near kolekole PRESERVED_SPECIMEN takeuchi, w.n. paquin, d. collector number: 2814 Lanai BISH 9/6/1986
149 Zanthoxylum kauaense A.Gray Waianae Mts., trail to Puu Kanehoa just below top of ridge on N facing slope PRESERVED_SPECIMEN Marr, K.L. 38 Oahu BISH 6/7/1988
150 Zanthoxylum kauaense A.Gray kumueli gulch PRESERVED_SPECIMEN oppenheimer, h.l. perlman, s.; tangalin, n. collector number: h20722 Kauai BISH 2/22/2007
151 Zanthoxylum kauaense A.Gray Lihue E. side PRESERVED_SPECIMEN Russ, G.W. s.n. Kauai BISH 10/1/1929
152 Zanthoxylum kauaense A.Gray ridge S of S. Opaeula Gulch, Paalaa, Koolau Mts wooded PRESERVED_SPECIMEN St. John, H. 11523 Oahu BISH 1/17/1932
153 Zanthoxylum kauaense A.Gray Waianae Mts., Waianae Kai, saddle between Mt. Kaala and Puu Kawiwi, ~100 m from power lines with Cibotium, Bidens, Alyxia, Psidium, koa, ohia, Cyanea, Joinvillea PRESERVED_SPECIMEN Marr, K.L. 66 Oahu BISH 7/4/1988
154 Zanthoxylum kauaense A.Gray Waimea District. Kohua Ridge and south facing slopes above Koaie. Metrosideros-Acacia montane mesic forest w/ Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Dracaena aurea, Polyscias kavaiensis, Planchonella sandwicensis, Bob PRESERVED_SPECIMEN Wood, K.R. 17177 Kauai BISH 11/3/2016
155 Zanthoxylum kauaense A.Gray kahanahaiki gulch side; N sheltered slope PRESERVED_SPECIMEN judd, c.s. collector number: 73 Oahu BISH 3/29/1928
156 Zanthoxylum kauaense A.Gray PRESERVED_SPECIMEN o. degener 4109 MPM 12/6/1931
157 Zanthoxylum kauaense A.Gray waianae mts., kaluaa gulch E facing slope of small gulley in N part of gulch PRESERVED_SPECIMEN marr, k.l. takeuchi, w.t. collector number: 69 Oahu BISH 7/17/1988
158 Zanthoxylum kauaense A.Gray makaleha Growing in fallow taro loi, 3-5 inches of water PRESERVED_SPECIMEN russ, g.w. Hawaii BISH 10/1/1929
159 Zanthoxylum kauaense A.Gray waimea district. makaha valley, 1.3 miles down makaha ridge road from hwy 550, below kokio keokeo picnic area On coral rubble. PRESERVED_SPECIMEN flynn, t. uchida, g. collector number: 2774 BISH 2/24/1988
160 Zanthoxylum kauaense A.Gray anahulu trail, koolau range rainy PRESERVED_SPECIMEN degener, o. et al. collector number: 10084 Oahu BISH 6/16/1935
161 Zanthoxylum kauaense A.Gray puu hapapa, ridge buttress PRESERVED_SPECIMEN takeuchi, w.n. paquin, d. collector number: 2804 Oahu BISH 9/1/1986
162 Zanthoxylum kauaense A.Gray n head of kahanahaiki valley dense forest PRESERVED_SPECIMEN degener, o. hatheway, w.h. collector number: 20639 Oahu BISH 5/7/1950
163 Zanthoxylum kauaense A.Gray Kokee, jct. Kalalau Rd. and Kalua-Puhi trail PRESERVED_SPECIMEN Herbst, D.R. 1194 Kauai BISH 8/10/1968
164 Zanthoxylum kauaense A.Gray near Kolekole on sliding rubble PRESERVED_SPECIMEN Takeuchi, W.N. 2815 Oahu BISH 9/6/1986
165 Zanthoxylum kauaense A.Gray slopes of mt. kaala PRESERVED_SPECIMEN takeuchi, w.n. et al. Oahu BISH 12/1/1982
166 Zanthoxylum kauaense A.Gray kula, s of sanitorium PRESERVED_SPECIMEN forbes, c.n. collector number: 2176.m Maui BISH 4/8/1920
167 Zanthoxylum kauaense A.Gray Waimea Canyon, toward head from Puu Kapele dryland forest PRESERVED_SPECIMEN Christensen, C. 199 Kauai BISH 5/7/1977
168 Zanthoxylum kauaense A.Gray waianae mts., w side of main ridge separating makaha and waianae kai valleys, n of puu kawiwi PRESERVED_SPECIMEN herbst, d.r. collector number: 5044b Oahu BISH 9/21/1974
169 Zanthoxylum kauaense A.Gray buttress ridge NW of Kaluaa gulch, near summit divide; interval between Puu Hapapa and Kanehoa a local concentration on steep slopes PRESERVED_SPECIMEN Takeuchi, W.N. 2048 Oahu BISH 3/3/1985
170 Zanthoxylum kauaense A.Gray makaha valley, on steep slope of ridge leading down into makaha valley from near well know pinnacle rocks, waianae,kai area, east of kamaileunu ridge PRESERVED_SPECIMEN perlman, s.p. collector number: 5058 Oahu BISH 1/8/1987
171 Zanthoxylum kauaense A.Gray mountains near koele PRESERVED_SPECIMEN forbes, c.n. collector number: 113.l Lanai BISH 6/1/1913
172 Zanthoxylum kauaense A.Gray koolau range, malaekahana-waimea, summit trail Cultivated. PRESERVED_SPECIMEN st.john, h. collector number: 18666 Kauai BISH 1/9/1938
173 Zanthoxylum kauaense A.Gray Waianae Mts., near Halona Valley PRESERVED_SPECIMEN Welch, D.A. s.n. Oahu BISH 6/4/1936
174 Zanthoxylum kauaense A.Gray Kapapala PRESERVED_SPECIMEN Russ, G.W. s.n. Hawaii BISH 3/1/1928
175 Zanthoxylum kauaense A.Gray Palikea Trail PRESERVED_SPECIMEN Bennett, T. PRC 223 Oahu BISH 3/11/1981
176 Zanthoxylum kauaense A.Gray waianae mts., pukaloa gulch, above schofield, off ridge trail to puu kalena, near kumakalii wet forest, with Metrosideros, Alyxia, Canthium, Nestegis, Chamaesyce, Psidium, Antidesma, Pleomele, Claoxylon, Urera, Nothocestrum longiflorum, Embelia, Pisonia, Bobea, Psychotria, Scaevola, Melicope, Astelia, Labordia, Perrottetia PRESERVED_SPECIMEN perlman, s.p. hill, r. collector number: 5644 Oahu BISH 4/16/1987
177 Zanthoxylum kauaense A.Gray middle palawai ridge PRESERVED_SPECIMEN degener, o. takamoto collector number: 10863 Oahu BISH
178 Zanthoxylum kauaense A.Gray Waimea, Kohua Ridge, Kawaia Trail PRESERVED_SPECIMEN Cranwell, L.M. 3855 Kauai BISH 8/19/1938
179 Zanthoxylum kauaense A.Gray Waianae Range, DuPont trail above Wailua steep slope, wind swept wet forest PRESERVED_SPECIMEN Stone, B.C. 3184 Oahu BISH 12/20/1959
180 Zanthoxylum kauaense A.Gray kukui trail, waimea canyon PRESERVED_SPECIMEN degener, o. degener, i.; hansen, h. collector number: 23961 Maui BISH 9/20/1955
181 Zanthoxylum kauaense A.Gray waianae mts., palikea, e of summit N facing slope; with Cryptomeria japonica PRESERVED_SPECIMEN marr, k.l. linney, g. collector number: 5 Oahu BISH 2/13/1988
182 Zanthoxylum kauaense A.Gray Naihakua, near PRESERVED_SPECIMEN Munro, G.C. 423 Lanai BISH 4/28/1925
183 Zanthoxylum kauaense A.Gray Anahulu Trail, Koolau Range rainy PRESERVED_SPECIMEN Degener, O. 10084 Oahu BISH 6/16/1935
184 Zanthoxylum kauaense A.Gray lihue distric; top of mt. haupu; Metrosideros Lowland Mixed Mesic Forest with North facing Native cliffs; Associated Plants: Bidens valida, Artemisia, Sida fallax, Hedyotis acuminata & H. fluviatilis, Bonamia, Isodendrion, Zanthoxylum, Carex wahuensis, Eragrotis variabilis, Hibiscus kokio, Lobelia niihauensis, Machaerina, Pipturus, Lepidium bidentatum, Mariscus, Melicope anisata. PRESERVED_SPECIMEN wood, k.r. perlman, s.; lorence, d.h.; marr, k. collector number: 2037 Kauai BISH 7/27/1992
185 Zanthoxylum kauaense A.Gray huliwai PRESERVED_SPECIMEN takeuchi, w.n. paquin, d. collector number: 2846 Oahu BISH 9/20/1986
186 Zanthoxylum kauaense A.Gray makaha valley, 1.5 mi. down ridge road with Metrosideros, Acacia, Zanthoxylum dipetalum PRESERVED_SPECIMEN flynn, t. collector number: 1218 Kauai BISH 9/10/1985
187 Zanthoxylum kauaense A.Gray kanehoa ridge 4m tree on slopes just below crestline PRESERVED_SPECIMEN takeuchi, w.n. holt, r.a.; perlman, s.; obata, j.; imada, c. collector number: 1852 Oahu BISH 11/18/1984
188 Zanthoxylum kauaense A.Gray mt. kaala, waianae range PRESERVED_SPECIMEN gillett, g.w. hardy, e.; nicharat, s.; st.john, h. collector number: 1720 Oahu BISH 3/12/1965
189 Zanthoxylum kauaense A.Gray Puu Kaua, Waianae Mts woods near summit PRESERVED_SPECIMEN Gillespie, J.W. 1064 Oahu BISH 7/6/1927
190 Zanthoxylum kauaense A.Gray Ridge S. of S.Opaeula Gulch, Paalaa, Koolau Mts Open moist woods PRESERVED_SPECIMEN St. John, H. 13339 Oahu BISH 9/24/1933
191 Zanthoxylum kauaense A.Gray Kula, S of sanitorium PRESERVED_SPECIMEN Forbes, C.N. 2176.M Maui BISH 4/8/1920
192 Zanthoxylum kauaense A.Gray W Maui, headwater region of Honokahua Valley. Riparian zone of ohia montane mesic forest within Unit 4 of Puu Kukui Watershed Mgmnt Area On E bank 15 ft. above intermittent stream. Assoc. with Dicranopteris linearis, Metrosideros polymorpha, Pipturus albidus, Tibouchina herbacea, Cheirodendron trigynum, Bobea elatior, Psychotria mariniana, Clermontia arborescens, Ageratina adenophorus, Nothocestrum longifolium, Cyanea scabra, C. macrostegia, Coniogramme pilosa, Amauropelta globulifera, Peperomia PRESERVED_SPECIMEN Meidell, J.S. 100 Maui BISH 10/18/1995
193 Zanthoxylum kauaense A.Gray lanai Open ocky or grassy ridge PRESERVED_SPECIMEN munro, g.c. collector number: 236 Kauai BISH 6/19/1915
194 Zanthoxylum kauaense A.Gray West Maui, Hanaula, Manawainui steep slope near gulch bottom PRESERVED_SPECIMEN Hobdy, R.W. 1720 Maui BISH 12/15/1982
195 Zanthoxylum kauaense A.Gray piko trail, makua valley, kawaihapai forest at top PRESERVED_SPECIMEN degener, o. park, k.k.; nitta, y. collector number: 8597 Oahu BISH 7/10/1932
196 Zanthoxylum kauaense A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 2103 Kauai BISH 3/3/1909
197 Zanthoxylum kauaense A.Gray waianae mts., s gulch between puu hapapa and puu kanehoa open, N facing slope, 40 m above bottom PRESERVED_SPECIMEN marr, k.l. takeuchi, w.t. collector number: 28 Oahu BISH 5/14/1988
198 Zanthoxylum kauaense A.Gray Hanalei Distric; Kalalau Rim; Kalahu side below first Kalalau lookout; Steep Diverse Montane Mesic forest with; Poa mannii, Lysimachia glutinosa, Zanthoxylum kauaense, Nototrichium sp. nov., Labordia helleri, Dryopteris unidentata, Metrosideros, Coprosma, Peucedanum, Vaccinium dentatum, Lipochaeta, Wilkesia. PRESERVED_SPECIMEN Wood, K.R. 1147 Kauai BISH 8/15/1991
199 Zanthoxylum kauaense A.Gray hauola PRESERVED_SPECIMEN munro, g.c. collector number: 441 Lanai BISH 6/9/1915
200 Zanthoxylum kauaense A.Gray N head of Kahanahaiki Valley dense forest PRESERVED_SPECIMEN Degener, O. 20639 Oahu BISH 5/7/1950
201 Zanthoxylum kauaense A.Gray waimea canyon, puu ka pele PRESERVED_SPECIMEN hobdy, r.w. collector number: 41 Kauai BISH 12/1/1968
202 Zanthoxylum kauaense A.Gray ridge crest between kaua and kanehoa Chamaesyce clearing PRESERVED_SPECIMEN takeuchi, w.n. paquin, d. collector number: 2847 Oahu BISH 9/20/1986
203 Zanthoxylum kauaense A.Gray Kaluapuhi Trail, Kokee State Park. PRESERVED_SPECIMEN Stern, W.L. 3154 Kauai BISH 4/6/1974
204 Zanthoxylum kauaense A.Gray Kumuwela PRESERVED_SPECIMEN Rice, R.C.A.