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Heliotropium arboreum

(Blanco) Mabb. (2017)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Boraginales Family: Heliotropiaceae Genus: Heliotropium

east indian velvetleaf, tahinu, tree heliotrope, tree heliotrope

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Description

Key Characters:

Growth Form: Small trees 1–5 m tall, often covered with firm unicellular hairs with a basal cystolith.

Stems:

Roots:

Leaves: Leaves simple. Alternate but appearing whorled, fleshy. Blades oblanceolate to obovate, 10–20 cm long, 3.5–9 cm wide. Apex obtuse. Base attenuate. Surfaces densely silky pubescent. Margins usually entire. Sessile to petiolate. Stipules absent.

Flowers: Flowers in stiff, paniculate, scorpioid cymes that uncoil progressively as the flowers open, cymes usually 6–8 cm long. Flowers bisexual (perfect), small, sessile. Calyx lobes oblong to orbicular, ca. 1.5 mm long, densely silky strigose. Corolla tubular, without scales in the throat, white, ca. 2.5–3.3 mm long, the tube short, 1.5–2 mm long, the lobes elliptic. Stamens slightly exserted; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-celled; ovules usually 4, 2 per carpel, eventually 1 per cell, sometimes fewer by abortion, anatropous, erect, ascending, or nearly horizontal, rarely pendulous; style 1, terminal, gynobasic arising from between the essentially distinct ovary lobes, with an annulus just below the stigma; stigma 2-lobed.

Fruit: Fruit white; drupaceous; fleshy; globose; ca. 4 mm in diameter; glabrous; sometimes with corky endocarp; an adaptation for floating. Seeds 1–4; endosperm absent or; if present; fleshy and scanty.

Ploidy:

Habitat: Naturalized and relatively common in coastal areas.

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Mokupāpapa (Kure atoll) Naturalized
Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
Manawai (Pearl & Hermes Atoll) Naturalized
O'ahu Naturalized
Kapou(Lisianski Island) Naturalized
Molokai Naturalized
Kamole(Laysan Island) Naturalized
Lana'i Naturalized
Lalo (French Frigate Shoals) Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents

Water/Ocean
Birds

Pollinators

Bibliography

Name Published In: Mabberley's Pl.-Book, ed. 4: 1101 (2017)

Other References

Wagner et al. 1990:398 (Ku, Mi, PH, Li, La, FF, Ni, K, O, Mo, L, M, H [as Tournefortia argentea]); Staples & Herbst 2005:197 (KEY), 199 (DESCR); Craven 2005:379 (Syn. T. argentea = H. foertherianum); Mabberley 2017:1101 (Syn. H. foertherianum = H. arboreum)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Heliotropium arboreum Kona HUMAN_OBSERVATION Friday, J.B. Hawaii 12/29/2003
2 Heliotropium arboreum Waikiki PRESERVED_SPECIMEN Rock, J.F.C. 2515 Oahu BISH 4/1/1909
3 Heliotropium arboreum Kahuku; Turtle Bay Coastal Shorline. Sandy soil. PRESERVED_SPECIMEN Sanchez, S. SS 03 Oahu BISH 3/20/1986
4 Heliotropium arboreum Mokuleia PRESERVED_SPECIMEN Degener, O. 12266 Oahu BISH 6/1/1937
5 Heliotropium arboreum w. danford's, kekaha, kauai PRESERVED_SPECIMEN blackman, l.g. bryan, i. Kauai BISH
6 Heliotropium arboreum Laysan One tree growing at head of NW beach. PRESERVED_SPECIMEN Tsuda, R.T. 506 Laysan BISH 12/8/1963
7 Heliotropium arboreum Eastern Island PRESERVED_SPECIMEN Frings, H.W. 35 Midway Atoll BISH 4/16/1962
8 Heliotropium arboreum Eastern I. Also found on Sand I PRESERVED_SPECIMEN Neff, J.A. 3 Midway Atoll BISH 11/1/1954
9 Heliotropium arboreum kapapa islet, kaneohe bay Raised coral reef with shallow, sandy soil. PRESERVED_SPECIMEN herbst, d.r. collector number: 6131 Oahu BISH 5/24/1978
10 Heliotropium arboreum Near end of Kalanianiole Hwy. Keaukaha PRESERVED_SPECIMEN Ishikawa, S. 222 Hawaii BISH 1/12/1973
11 Heliotropium arboreum kau district. along halape beach same from the shelter PRESERVED_SPECIMEN herat, t. herat, r.; higashino, p.k. collector number: 846 Hawaii BISH 6/30/1974
12 Heliotropium arboreum Kau district. Along Halape beach same from the shelter PRESERVED_SPECIMEN Herat, T. 846 Hawaii BISH 6/30/1974
13 Heliotropium arboreum burm island, keehi lagoon PRESERVED_SPECIMEN herbst, d.r. walker, r. collector number: 6071 Oahu BISH 4/30/1978
14 Heliotropium arboreum kapapa, oahu Found on margins of islet. PRESERVED_SPECIMEN starr, f. starr, k.; smith, d. collector number: 050418-49 Oahu BISH 4/18/2005
15 Heliotropium arboreum n lanai, beach kaiolohia bay, growing in solitary groove along the beach PRESERVED_SPECIMEN spence, g. collector number: 303 Oahu BISH 10/18/1973
16 Heliotropium arboreum near end of kalanianiole hwy. keaukaha PRESERVED_SPECIMEN ishikawa, s. collector number: 222 Hawaii BISH 1/12/1973
17 Heliotropium arboreum near poaiwa Near sandy shore PRESERVED_SPECIMEN degener, o. degener, i. collector number: 28608 Lanai BISH 7/24/1963
18 Heliotropium arboreum mokuleia PRESERVED_SPECIMEN degener, o. collector number: 12266 Oahu BISH 6/1/1937
19 Heliotropium arboreum strand, hanauma bay Growing in mixed stand with Thespesia populnea. PRESERVED_SPECIMEN crosby, m.r. anderson, w.r. collector number: 1551 Oahu BISH 6/25/1964
20 Heliotropium arboreum Near Kealia Planted along beaches, introduced. PRESERVED_SPECIMEN Little Jr., E.L. 31183 Kauai BISH 9/2/1976
21 Heliotropium arboreum east maui, hana district, kauiki, on the e side of hana bay Red sand beach PRESERVED_SPECIMEN flynn, t. bub, p. collector number: 2503 Maui BISH 10/21/1987
22 Heliotropium arboreum southeast island Collected on southern side of the eastern part of the island PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6485 Pearl And Hermes Reef BISH 7/7/1980
23 Heliotropium arboreum Kailua Beach, Kailua cultivated on beach PRESERVED_SPECIMEN Fosberg, F.R. 13291 Oahu BISH 9/26/1936
24 Heliotropium arboreum west coast, central PRESERVED_SPECIMEN Conant, S. 193 Lisianski BISH 8/8/1983
25 Heliotropium arboreum Southeast Island Collected on southern side of the eastern part of the island PRESERVED_SPECIMEN Herbst, D.R. 6485 Pearl And Hermes Atoll BISH 7/7/1980
26 Heliotropium arboreum wailuku district, found off waiehu beach road PRESERVED_SPECIMEN ishikawa, s. collector number: 160 Maui BISH 10/16/1972
27 Heliotropium arboreum Hawaii PRESERVED_SPECIMEN Rock, J.F.C. s.n. Hawaii BISH 7/15/1911
28 Heliotropium arboreum wailuku district, beach nw kahului Sandy flat PRESERVED_SPECIMEN fosberg, f.r. collector number: 9832 Maui BISH 8/17/1933
29 Heliotropium arboreum honolulu, iwilei, vacant lot; Growing in high tidepool. PRESERVED_SPECIMEN fosberg, f.r. collector number: 27148 Molokai BISH 10/31/1946
30 Heliotropium arboreum moanakai rd, kapaa PRESERVED_SPECIMEN ishikawa, s. collector number: 235 Oahu BISH 4/13/1973
31 Heliotropium arboreum northeast part of the island at interface of bunchgrass association and beach. PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6531 Laysan BISH 7/14/1980
32 Heliotropium arboreum Mokuauia Islet, Oahu PRESERVED_SPECIMEN Starr, F. 050222-25 Oahu BISH 2/22/2007
33 Heliotropium arboreum waialua, on lot owned bymrs.laura wilder Introduced, the species introduced at least 45 years ago, from Palmyra? (Guam trade) PRESERVED_SPECIMEN judd jr., a.f. collector number: 39 Oahu BISH 12/6/1925
34 Heliotropium arboreum Kapapa Islet, Kaneohe Bay Raised coral reef with shallow, sandy soil. PRESERVED_SPECIMEN Herbst, D.R. 6131 Oahu BISH 5/24/1978
35 Heliotropium arboreum Strand, Hanauma Bay Growing in mixed stand with Thespesia populnea. PRESERVED_SPECIMEN Crosby, M.R. 1551 Oahu BISH 6/25/1964
36 Heliotropium arboreum Sand island U.jS. Naval Station at east end of E-W air strip PRESERVED_SPECIMEN Beauchamp, R.M. 1274 Midway Atoll BISH 8/4/1970
37 Heliotropium arboreum green island In greenhouse. PRESERVED_SPECIMEN herbst, d.r. corn, c.; lamoureux, c.h. collector number: 6277 Oahu BISH 4/1/1997
38 Heliotropium arboreum East central edge of island PRESERVED_SPECIMEN Herbst, D.R. 6497 Lisianski BISH 7/7/1980
39 Heliotropium arboreum W Maui, Lahaina District, Honokahua PRESERVED_SPECIMEN St.John, H. Catto, R.J. Collector Number: 17715 Maui BISH 12/20/1936
40 Heliotropium arboreum N Lanai, Beach Kaiolohia Bay, Growing in solitary groove along the beach PRESERVED_SPECIMEN Spence, G. 303 Lanai BISH 10/18/1973
41 Heliotropium arboreum Midway - Sand Island PRESERVED_SPECIMEN Herbst, D.R. 6386 Midway Atoll BISH 6/29/1980
42 Heliotropium arboreum east island On the interior of the island, in sand, many perching birds. PRESERVED_SPECIMEN long, c.r. collector number: 2489 Midway Atoll BISH 9/27/1964
43 Heliotropium arboreum puuhonua o honaunau national historical park PRESERVED_SPECIMEN higashino, p.k. stemmermann, r.l.; allen, m.s. collector number: 10290 Hawaii BISH 4/8/1984
44 Heliotropium arboreum eastern i. also found on sand i PRESERVED_SPECIMEN neff, j.a. demont, p.a. collector number: 3 Midway Atoll BISH 11/1/1954
45 Heliotropium arboreum Kona HUMAN_OBSERVATION Friday, J.B. Hawaii 12/23/2017
46 Heliotropium arboreum hawaii PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 479 Hawaii BISH
47 Heliotropium arboreum Waikiki PRESERVED_SPECIMEN Rock, J.F.C. Collector Number: 2515 Oahu BISH 4/1/1909
48 Heliotropium arboreum growing in back of beach, kaloko Gully, dense Clidemia and Dicranopteris; no canopy PRESERVED_SPECIMEN herbst, d.r. collector number: 760 Oahu BISH 11/29/1967
49 Heliotropium arboreum kailua beach, kailua cultivated on beach PRESERVED_SPECIMEN fosberg, f.r. collector number: 13291 Oahu BISH 9/26/1936
50 Heliotropium arboreum kahuku, oahu Beach free naturalized at Kahuku, Oahu. Plant cultivated in Degener Mokuleia garden seeding itself along beach. First tree O.D., saw was at Waikiki Aquarium in 1922 PRESERVED_SPECIMEN degener, o. degener, i. Oahu BISH 11/1/1978
51 Heliotropium arboreum malaekahan state recreation area Assoc w. Leucaena, Terminalia & Panicum. PRESERVED_SPECIMEN annable, c.r. atha, d. collector number: 3132 Oahu BISH 11/11/1996
52 Heliotropium arboreum Green Island PRESERVED_SPECIMEN Herbst, D.R. 6277 Kure Atoll BISH 1/4/1979
53 Heliotropium arboreum east central edge of island medium-wet forest, on trunks of small trees PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6497 Oahu BISH 7/7/1980
54 Heliotropium arboreum Moomomi Beach PRESERVED_SPECIMEN Spaulding III, P. s.n. Molokai BISH 1/17/1983
55 Heliotropium arboreum kahuku; turtle bay Coastal Shorline. Sandy soil. PRESERVED_SPECIMEN sanchez, s. collector number: ss 03 Oahu BISH 3/20/1986
56 Heliotropium arboreum Keauhou PRESERVED_SPECIMEN Fagerlund, G.O. Mitchell, A.L. Collector Number: 385 Hawaii BISH 2/19/1943
57 Heliotropium arboreum Northwestern Hawaiian Islands Growing on a sand beach at the edge of the vegetation line; on a west central beach. About 9 living trees on west side close to camp site and sign and Casuarina tree. PRESERVED_SPECIMEN Conant, S. 172 Laysan BISH 7/25/1983
58 Heliotropium arboreum Kaohikaipu, Oahu on west end of island PRESERVED_SPECIMEN Starr, F. 050224-6 Oahu BISH 2/24/2005
59 Heliotropium arboreum Near Poaiwa Near sandy shore PRESERVED_SPECIMEN Degener, O. 28608 Lanai BISH 7/24/1963
60 Heliotropium arboreum Hawaii PRESERVED_SPECIMEN Mann, H. 479 Hawaii BISH 2/6/1905
61 Heliotropium arboreum Kekaha Kai State Park HUMAN_OBSERVATION Friday, J.B. Hawaii 12/27/2016
62 Heliotropium arboreum Kekaha Kai State Park HUMAN_OBSERVATION Friday, J.B. Hawaii 12/27/2016
63 Heliotropium arboreum Northeast part of the island at interface of bunchgrass association and beach. PRESERVED_SPECIMEN Herbst, D.R. 6531 Laysan BISH 7/14/1980
64 Heliotropium arboreum Kapapa Island PRESERVED_SPECIMEN Munro, G.C. 13 Oahu BISH 6/8/1946
65 Heliotropium arboreum queen's beach Rocky, sandy coastal plain. PRESERVED_SPECIMEN wagner, w.l. collector number: 5342 Oahu BISH 2/19/1984
66 Heliotropium arboreum eastern island PRESERVED_SPECIMEN frings, h.w. collector number: 35 Midway Atoll BISH 4/16/1962
67 Heliotropium arboreum Tern Islet Very common along the south side of the island, especially the southeast quarter, Most appear to be about the same age. PRESERVED_SPECIMEN Herbst, D.R. 6241 French Frigate Shoals BISH 9/11/1978
68 Heliotropium arboreum Laysan One plant at beach crest ca. 200m. North of campsite. PRESERVED_SPECIMEN Lamoureux, C.H. 1700 Laysan BISH 9/5/1961
69 Heliotropium arboreum Koolau Poko District, Mokapu Peninsula, Marine Corps Base Hawaii. Kaneohe Bay. Wildlife Management Area PRESERVED_SPECIMEN Herbst, D.R. 9766 Oahu BISH 5/23/1996
70 Heliotropium arboreum Near end of Kalanianiole Hwy. Keaukaha PRESERVED_SPECIMEN Ishikawa, S. Collector Number: 222 Hawaii BISH 1/12/1973
71 Heliotropium arboreum beach 1/2 mile from ewa golf course PRESERVED_SPECIMEN woolford, h.a. collector number: s.n. Oahu BISH 10/2/1956
72 Heliotropium arboreum french frigate shoals PRESERVED_SPECIMEN rainwater, h.i. French Frigate Shoals BISH 10/29/1953
73 Heliotropium arboreum East Maui, Hana District, Kauiki, on the E side of Hana Bay Red sand beach PRESERVED_SPECIMEN Flynn, T. 2503 Maui BISH 10/21/1987
74 Heliotropium arboreum kalae paoo, mahaulepu Sandy beach, sealevel PRESERVED_SPECIMEN stone, b.c. collector number: 729 Kauai BISH 6/14/1955
75 Heliotropium arboreum northwestern hawaiian islands Growing on a sand beach at the edge of the vegetation line; on a west central beach. About 9 living trees on west side close to camp site and sign and Casuarina tree. PRESERVED_SPECIMEN conant, s. collector number: 172 Laysan BISH 7/25/1983
76 Heliotropium arboreum eastern island PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6429 Midway Atoll BISH 6/30/1980
77 Heliotropium arboreum Moanakai Rd, Kapaa near sealevel PRESERVED_SPECIMEN Ishikawa, S. 235 Kauai BISH 4/13/1973
78 Heliotropium arboreum Small sand islet at inner north-west corner of triangle formed by seaplane runways, Keehi Lagoon PRESERVED_SPECIMEN Herbst, D.R. 1939 Oahu BISH 6/13/1971
79 Heliotropium arboreum midway - sand island PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6386 Midway Atoll BISH 6/29/1980
80 Heliotropium arboreum Honolulu, Iwilei, vacant lot; PRESERVED_SPECIMEN Fosberg, F.R. Collector Number: 27148 Oahu BISH 10/31/1946
81 Heliotropium arboreum kau district. along halape beach same distance from the shelter PRESERVED_SPECIMEN herat, t. herat, r.; higashino, p.k. collector number: 846 Hawaii BISH 6/30/1974
82 Heliotropium arboreum Kapapa Islet, Kaneohe Bay Sandy Coral Soil, on raised reef. PRESERVED_SPECIMEN Fosberg, F.R. 14036 Oahu BISH 6/6/1937
83 Heliotropium arboreum Laysan One plant at beach crest, ca. 200 m. North of campsite. PRESERVED_SPECIMEN Lamoureux, C.H. 1700 Laysan BISH 9/5/1961
84 Heliotropium arboreum Honolulu, Iwilei, vacant lot; PRESERVED_SPECIMEN Fosberg, F.R. 27148 Oahu BISH 10/31/1946
85 Heliotropium arboreum laysan PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1700 Oahu BISH 9/5/1961
86 Heliotropium arboreum Strand, Hanauma Bay Growing in mixed stand with Thespesia populnea. PRESERVED_SPECIMEN Crosby, M.R. Anderson, W.R. Collector Number: 1551 Oahu BISH 6/25/1964
87 Heliotropium arboreum along beach near waialua PRESERVED_SPECIMEN little jr., e.l. collector number: 31225 Molokai BISH 9/16/1976
88 Heliotropium arboreum Lihue District. Nawiliwili Small Boat Harbor. In grass median between parking lot and slips. PRESERVED_SPECIMEN Flynn, T. 8886 Kauai BISH 4/26/2018
89 Heliotropium arboreum East Maui, Hana District, Kauiki, on the E side of Hana Bay Red sand beach PRESERVED_SPECIMEN Flynn, T. Bub, P. Collector Number: 2503 Maui BISH 10/21/1987
90 Heliotropium arboreum small sand islet at inner north-west corner of triangle formed by seaplane runways, keehi lagoon PRESERVED_SPECIMEN herbst, d.r. pontes, j. collector number: 1939 Oahu BISH 6/13/1971
91 Heliotropium arboreum Kapapa, Oahu Found on margins of islet. PRESERVED_SPECIMEN Starr, F. 050418-49 Oahu BISH 4/18/2005
92 Heliotropium arboreum Queen's Beach Rocky, sandy coastal plain. PRESERVED_SPECIMEN Wagner, W.L. 5342 Oahu BISH 2/19/1984
93 Heliotropium arboreum Keauhou PRESERVED_SPECIMEN Fagerlund, G.O. 385 Hawaii BISH 2/19/1943
94 Heliotropium arboreum Popoia Islet, Oahu PRESERVED_SPECIMEN Starr, F. 050223-11 Oahu BISH 2/23/2005
95 Heliotropium arboreum moomomi beach PRESERVED_SPECIMEN spaulding iii, p. collector number: s.n. Molokai BISH 1/17/1983
96 Heliotropium arboreum kaaawa PRESERVED_SPECIMEN mccantoria collector number: s.n. Oahu BISH 10/6/1958
97 Heliotropium arboreum mokuauia islet, oahu PRESERVED_SPECIMEN starr, f. starr, k. collector number: 050222-25 Oahu BISH 2/22/2007
98 Heliotropium arboreum kaohikaipu, oahu on west end of island In fallow pineapple field; very common. PRESERVED_SPECIMEN starr, f. starr, k. collector number: 050224-6 Oahu BISH 2/24/2005
99 Heliotropium arboreum waikiki PRESERVED_SPECIMEN rock, j.f.c. collector number: 2515 Oahu BISH 4/1/1909
100 Heliotropium arboreum Along beach near Waialua PRESERVED_SPECIMEN Little Jr., E.L. 31225 Molokai BISH 9/16/1976
101 Heliotropium arboreum Wailuku District, found off Waiehu Beach Road PRESERVED_SPECIMEN Ishikawa, S. 160 Maui BISH 10/16/1972
102 Heliotropium arboreum keauhou PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 385 Hawaii BISH 2/19/1943
103 Heliotropium arboreum kapapa island PRESERVED_SPECIMEN munro, g.c. collector number: 13 Oahu BISH 6/8/1946
104 Heliotropium arboreum Kaaawa PRESERVED_SPECIMEN McCantoria s.n. Oahu BISH 10/6/1958
105 Heliotropium arboreum kahuku Coastal Plants PRESERVED_SPECIMEN guinto, j. collector number: jg 8 Oahu BISH 2/4/1986
106 Heliotropium arboreum French Frigate Shoals PRESERVED_SPECIMEN Rainwater, H.I. s.n. French Frigate Shoals BISH 10/29/1953
107 Heliotropium arboreum Wailuku District, Beach NW Kahului Sandy flat PRESERVED_SPECIMEN Fosberg, F.R. 9832 Maui BISH 8/17/1933
108 Heliotropium arboreum Small sand islet at inner north-west corner of triangle formed by seaplane runways, Keehi Lagoon PRESERVED_SPECIMEN Herbst, D.R. Pontes, J. Collector Number: 1939 Oahu BISH 6/13/1971
109 Heliotropium arboreum near kealia #NAME? PRESERVED_SPECIMEN little jr., e.l. collector number: 31183 Oahu BISH 9/2/1976
110 Heliotropium arboreum tern island PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1653 French Frigate Shoals BISH 9/2/1961
111 Heliotropium arboreum Eastern Island PRESERVED_SPECIMEN Herbst, D.R. 6429 Midway Atoll BISH 6/30/1980
112 Heliotropium arboreum Kamalino, 1 mile N of Top of beach PRESERVED_SPECIMEN St. John, H. 23610 Niihau BISH 4/30/1950
113 Heliotropium arboreum honolulu, kawaikui beach park ridge PRESERVED_SPECIMEN lau, j. collector number: 2142 Maui BISH 3/17/1986
114 Heliotropium arboreum kure island yew trees j@ NW end of island PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1895 Kure Atoll BISH 9/12/1961
115 Heliotropium arboreum Kure Island yew trees j@ NW end of island PRESERVED_SPECIMEN Lamoureux, C.H. 1895 Kure Atoll BISH 9/12/1961
116 Heliotropium arboreum Kahuku; Turtle Bay Coastal Shorline. Sandy soil. PRESERVED_SPECIMEN Sanchez, S. Collector Number: SS 03 Oahu BISH 3/20/1986
117 Heliotropium arboreum Beach just south of mouth of Wailua River, eastern Kauai. With Scaevola, Casuarina in sand near shore. PRESERVED_SPECIMEN Stern, W.L. 1353 Kauai BISH 1961-00-00
118 Heliotropium arboreum Mokuleia PRESERVED_SPECIMEN Degener, O. Collector Number: 12266 Oahu BISH 6/1/1937
119 Heliotropium arboreum Growing in back of beach, Kaloko Growing in arid, sandy soil. PRESERVED_SPECIMEN Herbst, D.R. Collector Number: 760 Oahu BISH 11/29/1967
120 Heliotropium arboreum Malaekahan State Recreation Area Assoc w. Leucaena, Terminalia & Panicum. PRESERVED_SPECIMEN Annable, C.R. 3132 Oahu BISH 11/11/1996
121 Heliotropium arboreum W. Danford's, Kekaha, Kauai PRESERVED_SPECIMEN Blackman, L.G. s.n. Kauai BISH
122 Heliotropium arboreum west coast, central PRESERVED_SPECIMEN conant, s. collector number: 193 Lisianski BISH 8/8/1983
123 Heliotropium arboreum Kau district. Along Halape beach same from the shelter PRESERVED_SPECIMEN Herat, T. Herat, R.; Higashino, P.K. Collector Number: 846 Hawaii BISH 6/30/1974
124 Heliotropium arboreum Kau district. Along Halape beach same distance from the shelter PRESERVED_SPECIMEN Herat, T. 846 Hawaii BISH 6/30/1974
125 Heliotropium arboreum Beach 1/2 mile from Ewa Golf Course PRESERVED_SPECIMEN Woolford, H.A. s.n. Oahu BISH 10/2/1956
126 Heliotropium arboreum Kahului PRESERVED_SPECIMEN Aborn, R. s.n. CM 5/1/1962
127 Heliotropium arboreum Beach 1/2 mile from Ewa Golf Course PRESERVED_SPECIMEN Woolford, H.A. Collector Number: s.n. Oahu BISH 10/2/1956
128 Heliotropium arboreum East Island On the interior of the island, in sand, many perching birds. PRESERVED_SPECIMEN Long, C.R. 2489 Midway Atoll BISH 9/27/1964
129 Heliotropium arboreum waimanalo bay state recreation area grassy picnic area behind beach PRESERVED_SPECIMEN pollack, g.h.; tetteris, a. 33 CM 11/18/1989
130 Heliotropium arboreum pequena comunidade de pukoo cultivated PRESERVED_SPECIMEN thorne, k. 9135 CM 5/2/1991
131 Heliotropium arboreum kau district, s coast, whittington beach state park, ne of na'alehu Leptospermum. PRESERVED_SPECIMEN weber, w.a.; bujakiewicz, a. 16581 IBUNAM 4/13/1983
132 Heliotropium arboreum kahuku, degener mokulua garden PRESERVED_SPECIMEN degener, o. 35753d DUKE
133 Heliotropium arboreum kahuku, oahu Beach garden PRESERVED_SPECIMEN otto degener; isa degener 35753a IND 12/8/1982
134 Heliotropium arboreum PRESERVED_SPECIMEN Stern WL; Herbst DR Stern, WL; Herbst, DR 2992 L
135 Heliotropium arboreum Lawai Valley. Cultivated plants of th e Pacific Tropical Botanical Garden. Near Pump 6 on west sid e of Lawai Stream near banana grove. PRESERVED_SPECIMEN Tim Flynn 845 Kauai PTBG 5/10/1984
136 Heliotropium arboreum whittingham beach, honuapo rocky lava shoreline. scaveola, grass. 0-50 feet elevation PRESERVED_SPECIMEN k. thorne; h. black, b. frost 6926 NCSC 7/31/1989
137 Heliotropium arboreum malaekahana state recreation area PRESERVED_SPECIMEN c. r. annable 3132 NY 11/11/1996
138 Heliotropium arboreum kahului PRESERVED_SPECIMEN aborn, r. s.n. NY 5/1/1962
139 Heliotropium arboreum ka`u district, along halape beach, hawaii volcanoes national park PRESERVED_SPECIMEN t. heart, r. herat & p. higashino 846 Hawaii US 6/30/1974
140 Heliotropium arboreum Nawiliwili Small Boat Harbor. In grass median between parking lot and slips. Cultivated. PRESERVED_SPECIMEN Tim Flynn 8886 Kauai PTBG 4/26/2018
141 Heliotropium arboreum Aliomanu. In vegetation line at th e back of the beach. PRESERVED_SPECIMEN Tim Flynn 3728 Kauai PTBG 1/3/1990
142 Heliotropium arboreum Kahuai, at Huialoha Church. Near sea level. PRESERVED_SPECIMEN Hank Oppenheimer H60112 Maui PTBG 6/10/2001
143 Heliotropium arboreum Strand at Ke'e. PRESERVED_SPECIMEN W. L. Stern 2992 Kauai PTBG 4/1/2003
144 Heliotropium arboreum Burns Field, Hanapepe. In weedy area just beyond high tide mark. PRESERVED_SPECIMEN Tim Flynn 1606 Kauai PTBG 3/18/1986
145 Heliotropium arboreum Coastal PRESERVED_SPECIMEN J.C. Watson JCW029 Laysan PTBG 9/27/2014
146 Heliotropium arboreum Larson's Beach between Moloaa and Kilauea. PRESERVED_SPECIMEN Tim Flynn 1322 Kauai PTBG 10/25/1985
147 Heliotropium arboreum Kauiki, on the eastern side of Hana Bay . Dominated by Casuarina planted in late 1930's but with numerous other alien species (Terminalia, Syzygium, Ardisia). elev. 0-386 ft. PRESERVED_SPECIMEN Tim Flynn 2503 Maui PTBG 10/21/1987
148 Heliotropium arboreum laysan, leeward island. PRESERVED_SPECIMEN c. h. lamoureux 1700 UPS 9/5/1961
149 Heliotropium arboreum off waiehu beach road PRESERVED_SPECIMEN s. s. ishikawa 160 Maui US 10/16/1972
150 Heliotropium arboreum north islet PRESERVED_SPECIMEN a. young 129 Kauai US 8/20/1964
151 Heliotropium arboreum beach just s of mouth of wailua river, e kauai PRESERVED_SPECIMEN w. l. stern & w. carlquist 1353 Kauai US
152 Heliotropium arboreum kawaihau district; aliomanu, at the back of the beach. PRESERVED_SPECIMEN t. w. flynn & b. schaeffer 3728 Kauai US 1/3/1990
153 Heliotropium arboreum north islet, along s and w side of island just above the high tide line PRESERVED_SPECIMEN c. long 2300 Pearl And Hermes Atoll US 9/17/1964
154 Heliotropium arboreum PRESERVED_SPECIMEN w. l. stern & s. carlquist 1353 Kauai US
155 Heliotropium arboreum east part of kauai, near kealia. PRESERVED_SPECIMEN e. l. little jr. 31183 Kauai US 9/2/1976
156 Heliotropium arboreum degener's mokuleia beach garden PRESERVED_SPECIMEN o. degener 36593 Oahu US
157 Heliotropium arboreum ko'olau poko distr., mokapu peninsula, marine corps base hawaii. kane'ohe bay, wildlife management area PRESERVED_SPECIMEN d. r. herbst 9766 Oahu US 5/23/1996
158 Heliotropium arboreum PRESERVED_SPECIMEN d. r. herbst & w. n. takeuchi 6386 Oahu US 6/29/1980
159 Heliotropium arboreum trig islet, central portion of islet PRESERVED_SPECIMEN c. long 2463 French Frigate Shoals US 9/27/1964
160 Heliotropium arboreum kapapa islet, kaneohe bay PRESERVED_SPECIMEN d. r. herbst 6131 Kauai US 5/24/1978
161 Heliotropium arboreum queens beach PRESERVED_SPECIMEN t. w. flynn 151b Maui US
162 Heliotropium arboreum nw part of island PRESERVED_SPECIMEN d. r. herbst & w. n. takeuchi 6531 Laysan US 7/14/1980
163 Heliotropium arboreum mokuauia PRESERVED_SPECIMEN c. long 1740 Kauai US 5/16/1964
164 Heliotropium arboreum tern island PRESERVED_SPECIMEN d. r. herbst 6241 Oahu US 9/11/1978
165 Heliotropium arboreum along beach near waialua, e molokai PRESERVED_SPECIMEN e. l. little jr. 31225 Molokai US 9/16/1976
166 Heliotropium arboreum honolulu, iwilei PRESERVED_SPECIMEN f. r. fosberg 27148 Oahu US 10/31/1946
167 Heliotropium arboreum off moanakai rd., kapaa, near sea level PRESERVED_SPECIMEN s. s. ishikawa 235 Pearl And Hermes Atoll US 4/13/1973
168 Heliotropium arboreum queens beach PRESERVED_SPECIMEN t. w. flynn 146b Oahu US
169 Heliotropium arboreum beach kaiolohia bay, north lanai PRESERVED_SPECIMEN g. spence 303 Lanai US 10/18/1973
170 Heliotropium arboreum along beach near end of kalanianiole hwy., keaukaha PRESERVED_SPECIMEN s. s. ishikawa 222 Hawaii US 1/12/1973
171 Heliotropium arboreum donkey beach PRESERVED_SPECIMEN Kauai US 1/8/2005
172 Heliotropium arboreum Burm Island, Keehi Lagoon PRESERVED_SPECIMEN Herbst, D.R. 6071 Oahu BISH 4/30/1978
173 Heliotropium arboreum Inland, in mineral rich, damp soil. PRESERVED_SPECIMEN Al-Bayer, S. Not on sheet Kure Atoll BISH 1/21/2019
174 Heliotropium arboreum tern islet PRESERVED_SPECIMEN herbst, d.r. collector number: 6241 Kauai BISH 9/11/1978
175 Heliotropium arboreum Kahuku, Oahu Beach free naturalized at Kahuku, Oahu. Plant cultivated in Degener Mokuleia garden seeding itself along beach. First tree O.D., saw was at Waikiki Aquarium in 1922 PRESERVED_SPECIMEN Degener, O. s.n. Oahu BISH 11/1/1978
176 Heliotropium arboreum Along beach near Waialua PRESERVED_SPECIMEN Little Jr., E.L. Collector Number: 31225 Molokai BISH 9/16/1976
177 Heliotropium arboreum kamalino, 1 mile n of top of beach PRESERVED_SPECIMEN st.john, h. collector number: 23610 Niihau BISH 4/30/1950
178 Heliotropium arboreum Kau district. Along Halape beach same distance from the shelter PRESERVED_SPECIMEN Herat, T. Herat, R.; Higashino, P.K. Collector Number: 846 Hawaii BISH 6/30/1974
179 Heliotropium arboreum sand island u.js. naval station at east end of e-w air strip In gulch PRESERVED_SPECIMEN beauchamp, r.m. collector number: 1274 Oahu BISH 8/4/1970
180 Heliotropium arboreum kapapa islet, kaneohe bay Sandy Coral Soil, on raised reef. PRESERVED_SPECIMEN fosberg, f.r. collector number: 14036 Oahu BISH 6/6/1937
181 Heliotropium arboreum w maui, lahaina district, honokahua PRESERVED_SPECIMEN st.john, h. catto, r.j. collector number: 17715 Maui BISH 12/20/1936
182 Heliotropium arboreum Tern Island PRESERVED_SPECIMEN Lamoureux, C.H. 1653 French Frigate Shoals BISH 9/2/1961
183 Heliotropium arboreum Honolulu, Kawaikui beach Park PRESERVED_SPECIMEN Lau, J. 2142 Oahu BISH 3/17/1986
184 Heliotropium arboreum W Maui, Lahaina District, Honokahua PRESERVED_SPECIMEN St. John, H. 17715 Maui BISH 12/20/1936
185 Heliotropium arboreum popoia islet, oahu PRESERVED_SPECIMEN starr, f. starr, k. collector number: 050223-11 Oahu BISH 2/23/2005
186 Heliotropium arboreum laysan PRESERVED_SPECIMEN tsuda, r.t. collector number: 506 Hawaii BISH 12/8/1963
187 Heliotropium arboreum koolau poko district, mokapu peninsula, marine corps base hawaii. kaneohe bay. wildlife management area PRESERVED_SPECIMEN herbst, d.r. collector number: 9766 Oahu BISH 5/23/1996
188 Heliotropium arboreum hawaii PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Hawaii BISH 7/15/1911
189 Heliotropium arboreum KaLae Paoo, Mahaulepu Sandy beach, sealevel PRESERVED_SPECIMEN Stone, B.C. 729 Kauai BISH 6/14/1955
190 Heliotropium arboreum Kalaupapa; airport, oceanview pavilion PRESERVED_SPECIMEN Wysong, M.L. 434 Molokai BISH 11/11/2004
191 Heliotropium arboreum e central edge of island PRESERVED_SPECIMEN d. r. herbst & w. n. takeuchi 6497 Lisianski US 7/11/1980
192 Heliotropium arboreum kalaupapa; airport, oceanview pavillion PRESERVED_SPECIMEN wysong, m.l. collector number: 434 Molokai BISH 11/11/2004
193 Heliotropium arboreum Puuhonua o Honaunau National Historical Park PRESERVED_SPECIMEN Higashino, P.K. 10290 Hawaii BISH 4/8/1984
194 Heliotropium arboreum Kahuku Coastal Plants PRESERVED_SPECIMEN Guinto, J. JG 8 Oahu BISH 2/4/1986
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