Themeda villosa

(Poir.) A.Camus (1922)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Poaceae Genus: Themeda

lyon's grass

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Description

Key Characters:

Growth Form: Robust perennials.

Stems: Culms tufted, stout, 20–40 dm tall, solid, culm base compressed flabellate.

Roots: Fibrous root system.

Leaves: Leaves simple. Alternate. Blades up to 150 cm long, up to 20 mm wide. Apex acuminate. Base narrowed to the sheath. Surfaces glabrous, midrib white, prominent, margins and nerves scabrous on upper surface. Margins entire. Veins parallel. Sheaths keeled, ± compressed, margins overlapping, glabrous; ligule membranous, ciliate. Stipules absent.

Flowers: Flowers in inflorescences large compound, drooping, paniculate, spathes 3–3.5 cm long, subtending a flattened rachis, apex pubescent, racemes of spikelet pairs at about the middle of the spathe. involucral and pedicellate spikelets reddish brown, similar in shape, narrowly fusiform, acuminate, 15–20 mm long, pedicels rufous pubescent; sessile spikelet with a callus ca. 2 mm long, densely pubescent with rufous hairs, first glume ovate, 10–nerved, the nerves reticulate toward apex, densely rufous pubescent between the nerves, apex acute, first lemma 1–nerved, apex acute; second lemma 1–nerved, hyaline, prolonged into a twisted pubescent awn up to 42 mm long, second palea lanceolate, nerveless, hyaline. Stamens 3 (when present). Ovary superior.

Fruit: Caryopsis reddish brown; fusiform; ca. 5 mm long.

Ploidy: 2n = 32; 40

Habitat: Dry pastures; along roadsides; and other disturbed sites.

Elevation Range: 0–700 m.

Historical Distribution

Images

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: H.Lecomte, Fl. Indo-Chine 7: 364 (1922)

Other References

Wagner et al. 1990:1599 (K, O, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
2 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
3 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
4 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
5 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
6 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
7 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
8 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
9 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
10 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
11 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
12 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
13 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
14 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
15 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
16 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
17 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
18 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
19 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
20 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
21 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
22 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
23 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
24 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
25 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
26 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
27 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
28 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
29 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
30 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
31 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
32 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
33 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
34 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
35 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
36 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
37 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
38 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
39 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
40 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
41 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
42 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
43 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
44 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
45 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
46 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
47 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
48 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
49 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
50 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
51 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
52 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
53 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
54 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
55 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
56 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
57 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
58 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
59 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
60 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
61 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
62 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
63 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
64 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
65 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
66 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
67 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
68 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
69 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
70 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
71 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
72 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
73 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
74 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
75 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
76 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
77 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
78 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
79 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
80 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
81 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
82 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
83 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
84 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
85 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
86 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
87 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
88 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
89 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
90 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
91 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
92 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
93 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
94 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
95 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
96 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
97 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
98 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
99 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
100 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
101 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
102 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
103 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
104 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
105 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
106 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
107 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
108 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
109 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
110 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
111 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
112 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
113 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
114 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
115 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
116 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
117 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
118 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
119 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
120 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
121 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
122 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
123 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
124 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
125 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
126 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
127 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
128 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
129 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
130 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
131 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
132 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
133 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
134 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
135 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
136 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
137 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
138 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
139 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
140 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
141 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
142 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
143 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
144 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
145 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
146 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
147 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
148 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
149 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
150 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
151 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
152 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
153 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
154 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
155 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
156 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
157 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
158 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
159 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
160 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
161 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
162 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
163 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
164 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
165 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
166 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
167 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
168 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
169 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
170 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
171 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
172 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
173 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
174 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
175 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
176 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
177 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
178 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
179 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
180 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
181 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
182 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
183 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
184 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
185 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
186 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
187 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
188 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
189 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
190 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
191 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
192 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
193 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
194 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
195 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
196 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
197 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
198 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
199 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
200 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
201 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
202 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
203 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
204 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
205 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
206 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
207 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
208 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
209 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
210 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
211 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
212 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
213 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
214 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
215 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
216 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
217 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
218 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
219 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
220 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
221 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
222 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
223 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
224 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
225 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936