Themeda villosa

(Poir.) A.Camus (1922)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Poaceae Genus: Themeda

lyon's grass

img

Description

Key Characters:

Growth Form: Robust perennials.

Stems: Culms tufted, stout, 20–40 dm tall, solid, culm base compressed flabellate.

Roots: Fibrous root system.

Leaves: Leaves simple. Alternate. Blades up to 150 cm long, up to 20 mm wide. Apex acuminate. Base narrowed to the sheath. Surfaces glabrous, midrib white, prominent, margins and nerves scabrous on upper surface. Margins entire. Veins parallel. Sheaths keeled, ± compressed, margins overlapping, glabrous; ligule membranous, ciliate. Stipules absent.

Flowers: Flowers in inflorescences large compound, drooping, paniculate, spathes 3–3.5 cm long, subtending a flattened rachis, apex pubescent, racemes of spikelet pairs at about the middle of the spathe. involucral and pedicellate spikelets reddish brown, similar in shape, narrowly fusiform, acuminate, 15–20 mm long, pedicels rufous pubescent; sessile spikelet with a callus ca. 2 mm long, densely pubescent with rufous hairs, first glume ovate, 10–nerved, the nerves reticulate toward apex, densely rufous pubescent between the nerves, apex acute, first lemma 1–nerved, apex acute; second lemma 1–nerved, hyaline, prolonged into a twisted pubescent awn up to 42 mm long, second palea lanceolate, nerveless, hyaline. Stamens 3 (when present). Ovary superior.

Fruit: Caryopsis reddish brown; fusiform; ca. 5 mm long.

Ploidy: 2n = 32; 40

Habitat: Dry pastures; along roadsides; and other disturbed sites.

Elevation Range: 0–700 m.

Historical Distribution

Images

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: H.Lecomte, Fl. Indo-Chine 7: 364 (1922)

Other References

Wagner et al. 1990:1599 (K, O, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
2 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
3 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
4 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
5 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
6 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
7 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
8 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
9 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
10 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
11 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
12 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
13 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
14 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
15 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
16 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
17 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
18 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
19 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
20 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
21 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
22 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
23 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
24 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
25 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
26 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
27 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
28 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
29 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
30 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
31 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
32 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
33 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
34 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
35 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
36 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
37 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
38 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
39 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
40 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
41 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
42 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
43 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
44 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
45 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
46 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
47 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
48 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
49 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
50 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
51 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
52 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
53 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
54 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
55 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
56 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
57 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
58 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
59 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
60 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
61 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
62 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
63 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
64 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
65 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
66 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
67 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
68 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
69 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
70 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
71 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
72 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
73 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
74 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
75 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
76 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
77 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
78 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
79 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
80 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
81 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
82 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
83 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
84 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
85 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
86 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
87 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
88 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
89 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
90 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
91 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
92 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
93 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
94 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
95 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
96 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
97 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
98 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
99 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
100 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
101 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
102 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
103 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
104 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
105 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
106 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
107 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
108 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
109 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
110 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
111 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
112 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
113 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
114 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
115 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
116 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
117 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
118 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
119 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
120 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
121 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
122 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
123 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
124 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
125 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
126 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
127 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
128 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
129 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
130 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
131 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
132 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
133 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
134 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
135 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
136 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
137 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
138 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
139 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
140 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
141 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
142 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
143 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
144 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
145 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
146 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
147 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
148 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
149 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
150 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
151 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
152 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
153 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
154 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
155 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
156 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
157 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
158 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
159 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
160 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
161 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
162 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
163 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
164 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
165 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
166 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
167 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
168 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
169 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
170 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
171 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
172 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
173 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
174 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
175 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
176 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
177 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
178 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
179 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
180 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
181 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
182 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
183 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
184 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
185 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
186 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
187 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
188 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
189 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
190 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
191 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
192 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
193 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
194 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
195 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
196 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
197 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
198 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
199 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
200 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
201 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
202 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
203 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
204 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
205 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
206 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
207 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
208 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
209 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
210 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
211 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
212 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
213 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
214 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
215 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
216 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
217 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
218 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
219 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
220 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
221 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
222 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
223 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
224 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
225 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936