Description
Key Characters:
Growth Form:
Scentless to weakly scented shrubs, sometimes tree-like.
Stems:
Stems erect or sometimes ascending or prostrate, 5–20(–30) dm long, branched, usually densely grayish mealy pubescent, sometimes young growth greener and only sparsely mealy pubescent.
Roots:
Leaves:
Leaves simple.
Alternate.
Blades broadly deltate to rhombic, (1.5–)2–4.2(–10) cm long, (0.6–)1.3–2.5(–5.5) cm wide.
Base truncate to cuneate.
Surfaces densely mealy pubescent, but Upper surfaces less so and greener than lower surface; blades thick and somewhat fleshy.
Margins usually 3-lobed, sometimes sinuate–dentate, the lobes or teeth obtuse to rounded.
Petioles (0.5–)1.3–2.5(–4.5) cm long.
Stipules absent.
Flowers:
Flowers in small, dense glomerules grouped into large, terminal, leafless or nearly leafless panicles.
Flowers bisexual (perfect), small.
Calyx ca. 0.8 mm long, completely enclosing the fruit at maturity, the lobes ovate, mealy pubescent, margins scarious, especially toward apex.
Corolla (petals) absent.
Stamens 1–5; anthers dithecal, opening by longitudinal slits.
Ovary superior, 2–3(–5)-carpellate, 1-celled; ovule 1, placentation basal; styles 2–5; stigmas dry.
Fruit:
Fruit a simple dry utricle;
Seeds 1 per fruit; dark brown; horizontal or sometimes vertical; ca. 0.8 mm in diameter; the surface papillose; margins indistinct.
Ploidy:
2n = 36
Habitat:
Dry habitats; including coastal; dry forest; and subalpine shrubland.
Elevation Range:
0–2520 m.
Island Status
Ka'ula Endemic
Ni'ihau(Incl. Lehua) Endemic
Kuaihelani (Midway Atoll) Only found in cultivation
Kaua'i
Endemic
O'ahu Endemic
Kapou(Lisianski Island) Endemic
Molokai Endemic
Kamole(Laysan Island) Endemic
Lana'i Endemic
Kaho'olawe Endemic
Lalo (French Frigate Shoals) Endemic
Maui Endemic
Mokumanamana(Necker) Endemic
Hawai'i Endemic
Nihoa Endemic