Description

Key Characters:

Growth Form: Erect perennial herbs.

Stems: Stems apparently short–hispidulous.

Roots:

Leaves: Leaves simple. Opposite. Blades ovate, ca. 11.5–18 cm long, 5.1–9 cm wide. Apex acuminate. Base subcordate to truncate. Surfaces sparsely hirtellous, blades flaccid. Margins crenate. Petioles 4–6.5 cm long, short–hispidulous. Stipules absent.

Flowers: Flowers 2 or 4 per verticillaster, on short, leafless, lateral branches arising in the axils below lowermost leaves, these forming racemose inflorescences 4–6.5 cm long, short–hispidulous throughout, pedicels ca. 2–4 mm long, each flower subtended by spatulate bracts 2–3 mm long. Calyx broadly obconical, ca. 2.5 mm long at anthesis, enlarging in fruit, prominently 10–nerved, hispidulous (a few hairs apparently gland–tipped), usually 5-toothed, the teeth broadly deltate, ca. 0.5–0.7 mm long, apex acute, splitting longitudinally, usually persistent or sometimes only ventral 1/2 persistent. Corolla apparently 6–8 mm long, color unknown, tube narrowly funnelform, usually curved, upper lip entire or 2-lobed, flat or concave, lower lip larger, 3-lobed, lateral lobes ovate, slightly asymmetrical, lower lobe broadly ovate to suborbicular, slightly folded to form a groove, margins somewhat undulate, apex emarginate or entire; nectary disk often present at base of ovary, annular or developed on anterior side only. Stamens 4, both pairs of similar length or upper pair slightly longer, ascending the upper corolla lip, included, aligned with the sinuses; filaments inserted near throat, usually pubescent; anthers dithecal, opening by longitudinal slits; anther sacs divaricate. Ovary superior, 2-carpellate, placentation basal-axile; ovules 1 per ovary lobe; style 1, included, arising between lobes, shortly 2-lobed, the lobes clavate; stigmatic only at apex, 1 lobe often rudimentary; stigmas 2.

Fruit: Nutlets 1.5–2 mm long. Seeds with endosperm absent or scanty and oily.

Ploidy:

Habitat: Known only from the type collection from woods (probably in diverse mesic forest).

Elevation Range: