Description

Key Characters:

Growth Form: Herbaceous to suffrutescent perennial or annual herbs or subshrubs, pubescence various, the hairs primarily stellate and simple.

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Roots:

Leaves: Leaves simple. Alternate. Blades lanceolate to broadly ovate, rarely slightly 3-lobed. Apex acute to acuminate or obtuse. Base cordate-truncate to broadly cuneate. Margins serrate to crenate or dentate. Petioles shorter than blades. Stipules linear to triangular, usually curved, +/- persistent.

Flowers: Flowers axillary, solitary or glomerate in the leaf axils or in terminal spikes or racemes, subsessile or pedicellate; involucral bracts 3, linear to lanceolate or unguiculate, distinct or adnate basally to calyx. Flowers bisexual (perfect). Calyx composed of connate sepals, 5-lobed, often angled in bud, the lobes valvate in bud. Corolla actinomorphic to moderately zygomorphic, yellow to yellowish orange, rotate to campanulate or sometimes not opening, the corolla of 5 petals, obovate, distinct from each other but adnate at base to staminal column, convolute in bud. Stamens 5 to numerous, monadelphous, forming a staminal column, staminal column shorter than corolla and same in color, terminated by filaments; anthers monothecal. Pollen globose, echinate. Ovary superior, carpels 5-18, borne in a single whorl or rarely seemingly superposed whorls, placentation axile; ovules 1 per carpel, ascending; style exceeding the staminal column, branches as many as carpels; stigmas terminal, capitate.

Fruit: Schizocarp; mericarps transverse-elliptic to quadrangular in lateral view; unarmed or with 1-3 apical mucros or awns; completely or partly dehiscent loculicidally or indehiscent; endoglossum present or absent. Seeds reddish brown to black; subreniform; compressed; glabrous; with or without endosperm.

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