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Lepidium didymum

L. (1767)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Brassicales Family: Brassicaceae Genus: Lepidium

peppergrass, pepperwort, swinecress, swinecress

Description

Key Characters:

Growth Form: Sparsely pubescent, aromatic annual or biennial herbs.

Stems: Outer stems prostrate, 1–4 dm long.

Roots:

Leaves: Leaves simple. Alternate or rarely opposite. Basal leaves oblong in outline, 3–10 cm long, deeply pinnatifid. stem leaves similar but shorter. Margins lobed, lobes dentate. Petioles of basal leaves 0.5–4(–6) cm. Cauline leaves petiolate to subsessile. Stipules absent.

Flowers: Flowers in racemes usually axillary. Flowers bisexual (perfect), actinomorphic or rarely slightly irregular. Calyx of 4 sepals, deciduous, erect, usually oblong, sometimes the inner 2 with gibbous bases that hold the nectar. Corolla of 4 petals, white, spatulate, scarcely exceeding the sepals, ca. 1 mm long. Stamens 2 tetradynamous, the inner 4 usually in pairs, sometimes connate at base in pairs; anthers dithecal, opening by longitudinal slits. Ovary superior, 2(4?)-carpellate, usually 2-celled by means of a false, but usually complete septum, rarely 1-celled, sessile or rarely stipitate; ovules 1 to numerous, borne on parietal placentas on replum margin at periphery of ovary wall, campylotropous or occasionally anatropous; style 1 or occasionally absent; stigma capitate.

Fruit: Capsules divided into 2 cells by the usually thin and membranous septum; elongate (at least 3 times as long as wide) referred to as a silicle; deeply constricted at apex and base; producing a fruit with 2 globose halves on opposite sides of a narrow partition; indehiscent; reticulate or wrinkled; sometimes tuberculate; didymous; 2–2.5 mm wide; finely wrinkled. Seeds 1 per cell; often becoming mucilaginous when wet; endosperm essentially absent.

Ploidy: 2n = 32

Habitat: Naturalized in dry; disturbed sites; especially common in lawns.

Elevation Range: 0–2100 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Mokupāpapa (Kure atoll) Naturalized
Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
Manawai (Pearl & Hermes Atoll) Naturalized
O'ahu Naturalized
Molokai Naturalized
Kamole(Laysan Island) Naturalized
Lana'i Naturalized
Kaho'olawe Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: Mant. Pl.: 92 (1767)

Other References

Wagner et al. 1990:403 (Mi, PH, Ni, K, O, Mo, L, M, Ka, H [as Coronopus didymus]); Staples et al. 2003:9 (La); Al-Shehbaz et al. 2002:6 (Syn. C. didymus)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Lepidium didymum L. honolulu, grounds of bishop museum PRESERVED_SPECIMEN neal, m.c. Oahu BISH 5/6/1952
2 Lepidium didymum L. kalaupapa national historical park Coastal strand vegetation. Plant invading post-deer damage. PRESERVED_SPECIMEN gagné, b.h. trainer, j. collector number: 3153 Molokai BISH 9/17/2000
3 Lepidium didymum L. upper makakilo, adjacent to developed subdivision at end of pueonani street Open area in solid Panicum maximum grassland, with /Waltheria indica, Sida fallax, Eleusine indica, Verbesina encelioides, Sonchus oleraceus. PRESERVED_SPECIMEN imada, c.t. crago, l. collector number: 2004-24 Oahu BISH 3/9/2004
4 Lepidium didymum L. P.T.B.G., Lawai Valley Uncommon in Valley. Native to Eurasia. PRESERVED_SPECIMEN Herbst, D.R. 2274 Kauai BISH 12/30/1971
5 Lepidium didymum L. in pineapple field PRESERVED_SPECIMEN hosaka, e.y. collector number: 470 Lanai BISH 6/23/1931
6 Lepidium didymum L. honolulu PRESERVED_SPECIMEN forbes, c.n. collector number: 1026 Oahu BISH 1/15/1909
7 Lepidium didymum L. east molokai, north coast, kukaiwaa on windswept peninsula Growing in a thick carpet of Fimbristylis pycnocephala, low prostrate. PRESERVED_SPECIMEN hobdy, r.w. collector number: 2185 Molokai BISH 8/27/1984
8 Lepidium didymum L. Southeast Island. Collected from an extensive, nearly pure colony growing on the northern side of the western part of the islan PRESERVED_SPECIMEN Herbst, D.R. 6489 Hawaii BISH 7/7/1980
9 Lepidium didymum L. kaeo cliffs, w molokai Tiny sedge growing in wet mud on pond banks. PRESERVED_SPECIMEN f. r. fosberg 29583 Oahu BISH 2/22/1948
10 Lepidium didymum L. eastern-central part of island,slope of moaula PRESERVED_SPECIMEN cuddihy, l.w. char, w.p. collector number: 433 Kahoolawe BISH 4/25/1980
11 Lepidium didymum L. South Kohala District, east of Queen Kaahumanu Hwy, between Mauna Lani Drive and Puako turnoff. Dry area with scant vegetation; open grassland with scattered Prosopis pallida. PRESERVED_SPECIMEN Funk, E.J. N-27 Hawaii BISH 2/4/1991
12 Lepidium didymum L. Oahu Sugar Co PRESERVED_SPECIMEN Swezey, O.H. s.n. Oahu BISH 5/20/1930
13 Lepidium didymum L. Keauhou Ranch Growing along Pu'u O'o Road near a paddock. PRESERVED_SPECIMEN Davis, J. 725 Hawaii BISH 5/5/1982
14 Lepidium didymum L. Coral plain below Ewa Mill & Sisal PRESERVED_SPECIMEN Forbes, C.N. 2325.O Oahu BISH 5/14/1916
15 Lepidium didymum L. parker ranch, pokanaka, plowed field PRESERVED_SPECIMEN ewart iii, g.r. collector number: 174 Hawaii BISH 4/21/1932
16 Lepidium didymum L. west molokai, kaeo cliffs Common locally at tops of cliffs of volcanic rock PRESERVED_SPECIMEN fosberg, f.r. collector number: 29583 Molokai BISH 2/22/1948
17 Lepidium didymum L. Parker Ranch, Nienie Hawaii PRESERVED_SPECIMEN Rock, J.F.C. 8325 Hawaii BISH 6/1/1910
18 Lepidium didymum L. p.t.b.g., lawai valley Uncommon in Valley. Native to Eurasia. PRESERVED_SPECIMEN herbst, d.r. collector number: 2274 Kauai BISH 12/30/1971
19 Lepidium didymum L. eastern island PRESERVED_SPECIMEN frings, h.w. collector number: 46 Midway Atoll BISH 4/16/1962
20 Lepidium didymum L. oahu PRESERVED_SPECIMEN forbes, c.n. Oahu BISH 3/31/1909
21 Lepidium didymum L. uwekahuna blutt. found in shade Mesic summer-dry forest PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 452 Oahu BISH 3/15/1943
22 Lepidium didymum L. Crest of Koolau Range, head of Kamiloiki valley PRESERVED_SPECIMEN Egler, F.E. 37-110a Oahu BISH 6/12/1937
23 Lepidium didymum L. Tantalus, Honolulu PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 3/31/1908
24 Lepidium didymum L. Lanai PRESERVED_SPECIMEN Munro, G.C. 286 Lanai BISH 12/6/1913
25 Lepidium didymum L. koko crater, dry open slopes PRESERVED_SPECIMEN christophersen, e. collector number: 1301 Oahu BISH 4/6/1930
26 Lepidium didymum L. honolulu, bishop museum grounds PRESERVED_SPECIMEN suehiro, a. collector number: s.n. Oahu BISH 2/13/1963
27 Lepidium didymum L. koloa district. mahaulepu. in casuarina grove in dumes behind haula PRESERVED_SPECIMEN flynn, t. collector number: 1969 Kauai BISH 11/25/1986
28 Lepidium didymum L. Between J.C.Campbell Refuge and Beach Tropical Coastal Plants PRESERVED_SPECIMEN Smith, M. 22 Oahu BISH 4/8/1986
29 Lepidium didymum L. Ulupau Head, Mokapu Peninsula (?) Sandy bottom of gully PRESERVED_SPECIMEN Fosberg, F.R. 10905 Oahu BISH 5/26/1935
30 Lepidium didymum L. koko head PRESERVED_SPECIMEN swezey, o.h. Oahu BISH 1/1/1920
31 Lepidium didymum L. eastern island PRESERVED_SPECIMEN frings, h.w. collector number: 50 Midway Atoll BISH 4/16/1962
32 Lepidium didymum L. kokee state park. mohihi rd. near waineke swamp Common along road. PRESERVED_SPECIMEN flynn, t. collector number: 974 Kauai BISH 8/28/1984
33 Lepidium didymum L. popoia islet, oahu on west side near coast In closed Metrosideros forest. Frequent on Metrosideros branches. PRESERVED_SPECIMEN starr, f. starr, k. collector number: 050223-7 Maui BISH 2/23/2005
34 Lepidium didymum L. ewa plains survey-u.s.f.&w. campbell industrial park area PRESERVED_SPECIMEN higashino, p.k. collector number: 8193 Oahu BISH 2/1/1979
35 Lepidium didymum L. Mokuauia Islet, Oahu in central western part of island PRESERVED_SPECIMEN Starr, F. 050222-3 Oahu BISH 2/22/2007
36 Lepidium didymum L. West end of Mahaulepu area Locally common in pockets of soil at top of limestone cliffs overlooking the ocean. PRESERVED_SPECIMEN Fay, J.J. 167 Kauai BISH 2/1/1974
37 Lepidium didymum L. Honolulu, grounds of Bishop Museum PRESERVED_SPECIMEN Neal, M.C. s.n. Oahu BISH 5/6/1952
38 Lepidium didymum L. area between puu moaula and lua makika growing in grassland area PRESERVED_SPECIMEN higashino, p.k. stemmermann, r.l. collector number: 8039 Kahoolawe BISH 11/22/1978
39 Lepidium didymum L. Hawaii PRESERVED_SPECIMEN Mann, H. 331 Hawaii BISH 2/6/1905
40 Lepidium didymum L. Kalaupapa National Historical Park Coastal strand vegetation. Plant invading post-deer damage. PRESERVED_SPECIMEN Gagne, B.H. 3153 Molokai BISH 9/17/2000
41 Lepidium didymum L. lanai city PRESERVED_SPECIMEN degener, o. collector number: 28557 Lanai BISH 8/15/1963
42 Lepidium didymum L. between j.c.campbell refuge and beach Tropical Coastal Plants PRESERVED_SPECIMEN smith, m. collector number: 22 Oahu BISH 4/8/1986
43 Lepidium didymum L. university of hawaii campus PRESERVED_SPECIMEN st.john, h. collector number: 10461 Oahu BISH 4/5/1930
44 Lepidium didymum L. Sand Island. Along disturbed roadsides. PRESERVED_SPECIMEN Herbst, D.R. 6431 Midway Atoll BISH 1980-00-00
45 Lepidium didymum L. Mokapu, offshore islet east of Kalaupapa Peninsula. Chamaesyce celastroides/Chenopodium oahuense coastal dry shrubland with Diospyros forest PRESERVED_SPECIMEN K. R. Wood 8307 Mokapu PTBG 3/8/2000
46 Lepidium didymum L. southeast island. collected from an extensive, nearly pure colony growing on the northern side of the western part of the islan PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6489 Hawaii BISH 7/7/1980
47 Lepidium didymum L. Upper Makakilo, adjacent to developed subdivision at end of Pueonani Street Open area in solid Panicum maximum grassland, with /Waltheria indica, Sida fallax, Eleusine indica, Verbesina encelioides, Sonchus oleraceus. PRESERVED_SPECIMEN Imada, C.T. Crago, L. Collector Number: 2004-24 Oahu BISH 3/9/2004
48 Lepidium didymum L. Airplane field, in ,owed grass, Kapaa Quarry Road, Kailua In mowed grass at edge of airplane field. Soil moist, loamy. Full sun. PRESERVED_SPECIMEN Crago, L.M. 2005-021 Oahu BISH 1/20/2005
49 Lepidium didymum L. napali coast, nualolo kai. west side of valley on talus slope near backwall. PRESERVED_SPECIMEN n. tangalin 2314 LSU 5/3/2010
50 Lepidium didymum L. Along the highway near the lands of Waialua PRESERVED_SPECIMEN Herbst, D.R. 5750 Molokai BISH 3/10/1976
51 Lepidium didymum L. Honolulu PRESERVED_SPECIMEN Forbes, C.N. 1026 Oahu BISH 1/15/1909
52 Lepidium didymum L. Kalaupapa; peninsula, base of lighthouse stairs along rock wall PRESERVED_SPECIMEN Wysong, M.L. 617 Molokai BISH 2/8/2005
53 Lepidium didymum L. Honolulu PRESERVED_SPECIMEN Forbes, C.N. Collector Number: 1026 Oahu BISH 1/15/1909
54 Lepidium didymum L. West Molokai, Kaeo Cliffs Common locally at tops of cliffs of volcanic rock PRESERVED_SPECIMEN Fosberg, F.R. 29583 Molokai BISH 2/22/1948
55 Lepidium didymum L. niihau PRESERVED_SPECIMEN Oahu BISH
56 Lepidium didymum L. ulupau head, mokapu peninsula (?) Sandy bottom of gully PRESERVED_SPECIMEN fosberg, f.r. collector number: 10905 Oahu BISH 5/26/1935
57 Lepidium didymum L. Oahu PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 3/31/1909
58 Lepidium didymum L. molokini islet rare along inner slope. PRESERVED_SPECIMEN hobdy, r.w. collector number: 1228 Maui BISH 2/14/1982
59 Lepidium didymum L. KAUAI: Napali Coast, Nualolo Kai. West side of valley on talus slope near backwall. Talus slope with Polycarpon tetraphyllum, Anagallis arvensis, Xanthium strumarium var. canadense, Stachytarpheta jamaicensis, Pityrogramma austroamericana, Pluchea, Ageratina, Emilia fosbergii, Plantago lanceolata, Momordica charantia, Erigeron karvinskianus, Myoporum sandwicense, Chenopodium murale, Adiantum hispidulum. PRESERVED_SPECIMEN Natalia Tangalin 2314 Kauai PTBG 5/3/2010
60 Lepidium didymum L. KAUAI: Salt Pond Beach Park area, ancient salt pans and littoral vegetation, western end of Hanapepe makai of Burns Field perimeter, shrubland/grassland/herbland with Sida fallax, Jacquemontia ovalifolia, Ipomoea pes-caprae subsp brasiliensis, Sporobolus virginicus, Sesuvium portulacastrum, Scaevola taccada, Vigna marina, Panicum torridum, Panicum fauriei v. latius, Lycium sandwicense PRESERVED_SPECIMEN K. R. Wood 11136 Kauai PTBG 2/14/2005
61 Lepidium didymum L. Kokee State Park. Mohihi Road near Waineke Swamp. PRESERVED_SPECIMEN Tim Flynn 974 Kauai PTBG 12/28/1984
62 Lepidium didymum L. MAUI: W. Maui, Lahaina District, Honokahua. Elev. 200 ft. PRESERVED_SPECIMEN Hank Oppenheimer H700118 Maui PTBG 7/27/2000
63 Lepidium didymum L. Puu Waawaa, Umii Manu cave. PRESERVED_SPECIMEN K. R. Wood 2065 Hawaii PTBG 9/1/1992
64 Lepidium didymum L. Lawai Valley, P.T.B.G. PRESERVED_SPECIMEN Derral Herbst 2274 Kauai PTBG 12/1/2030
65 Lepidium didymum L. West end of Mahaulepu area in pockets of soil on top of limestone cliffs overlooking ocean. PRESERVED_SPECIMEN John J. Fay 167 Kauai PTBG 2/1/1974
66 Lepidium didymum L. KAUAI: Koloa District. Mahaulepu. In Casuarina grove in dune s behind Haula. Elev. ca 100 ft. PRESERVED_SPECIMEN Tim Flynn 1969 Kauai PTBG 11/25/1986
67 Lepidium didymum L. Honolulu, upper Manoa Valley PRESERVED_SPECIMEN Hitchcock, A. S. 13745 Oahu US 6/16/2016
68 Lepidium didymum L. Kure Atoll. Around LORAN buildings and roadsides in that area. PRESERVED_SPECIMEN Derral Herbst 6252 Green PTBG 1/4/1979
69 Lepidium didymum L. In and on the slopes of Makiki PRESERVED_SPECIMEN Heller, A. A. 1974 Oahu US 3/21/1994
70 Lepidium didymum L. Palawa, Kaluanui Flats PRESERVED_SPECIMEN Herbst, D. R.; Spence, G. 5802 Lanai US 4/1/1976
71 Lepidium didymum L. in and on the slopes of makiki PRESERVED_SPECIMEN a. a. heller 1974 Maui US 3/21/1994
72 Lepidium didymum L. honolulu district, koko head. to east and south of ihiihilauakea crater. PRESERVED_SPECIMEN m. bruegmann 145 Oahu US 1/29/1984
73 Lepidium didymum L. james campbell national wildlife refuge, kahuku, in vicinity of bird-viewing gazebo PRESERVED_SPECIMEN c. imada, a. engilis & t. duk 92-17 Oahu US 4/5/1992
74 Lepidium didymum L. Honolulu District, Koko Head. To east and south of Ihiihilauakea Crater. PRESERVED_SPECIMEN Bruegmann, M. M. 145 Oahu US 1/29/1984
75 Lepidium didymum L. Kaeo Cliffs, W Molokai PRESERVED_SPECIMEN Fosberg, F. R. 29583 Molokai US 2/22/1948
76 Lepidium didymum L. Below Ewa & Sisal PRESERVED_SPECIMEN Forbes, C. N. 2325.O. Oahu US 3/14/2016
77 Lepidium didymum L. Napali Coast, Nualolo Kai. West side of valley on talus slope near backwall. PRESERVED_SPECIMEN Tangalin, N. 2314 Kauai US 5/3/2010
78 Lepidium didymum L. James Campbell National Wildlife Refuge, Kahuku, in vicinity of bird-viewing gazebo PRESERVED_SPECIMEN Imada, C. T.; Engilis, A.; Duk, T. 92-17 Oahu US 4/5/1992
79 Lepidium didymum L. Along the highway near the lands of Waialua PRESERVED_SPECIMEN Herbst, D. R.; Spence, G. 5750 Molokai US 3/10/1976
80 Lepidium didymum L. makapuu pt., near waimanalo PRESERVED_SPECIMEN o. degener 17226 Oahu US
81 Lepidium didymum L. Kalaupapa National Historical Park PRESERVED_SPECIMEN B. H. Gagn?? & J. Trainer 3153 Molokai US 9/17/2000
82 Lepidium didymum L. kalaupapa national historical park PRESERVED_SPECIMEN b. h. gagné & j. trainer 3153 Molokai US 9/17/2000
83 Lepidium didymum L. honolulu, upper manoa valley PRESERVED_SPECIMEN a. s. hitchcock 13745 Oahu US 6/16/1916
84 Lepidium didymum L. along the highway near the lands of waialua PRESERVED_SPECIMEN d. r. herbst & g. spence 5750 Molokai US 3/10/1976
85 Lepidium didymum L. below ewa & sisal PRESERVED_SPECIMEN c. n. forbes 2325.o. Oahu US 3/14/1916
86 Lepidium didymum L. palawa, kaluanui flats PRESERVED_SPECIMEN d. r. herbst & g. spence 5802 Lanai US 4/1/1976
87 Lepidium didymum L. Kalaupapa National Historical Park PRESERVED_SPECIMEN Gagn�, B. H.; Trainer, J. 3153 Molokai US 9/17/2000
88 Lepidium didymum L. tantalus, honolulu PRESERVED_SPECIMEN forbes, c.n. Oahu BISH 3/31/1908
89 Lepidium didymum L. hnp, maui halemau trail Grows in a tangles mass with very disagreeable odor. PRESERVED_SPECIMEN olson, g.e. collector number: 62 Maui BISH 7/11/1937
90 Lepidium didymum L. HNP, Maui Halemau Trail Grows in a tangles mass with very disagreeable odor. PRESERVED_SPECIMEN Olson, G.E. 62 Maui BISH 7/11/1937
91 Lepidium didymum L. KAUAI: Menehune Fish Pond, Niumalu, Okada property in Alien vegetation dominated by Rhizophora mangle and Hibiscus tiliaceus with Cocos nucifera, Casuarina equisetifolia, Pluchea indica, Thespesia populnea, Chrysophyllum, Macaranga, Ricinis communis, Panicum maximum PRESERVED_SPECIMEN K. R. Wood 5893 Kauai PTBG 12/20/1996
92 Lepidium didymum L. palawai, kaluanui flats PRESERVED_SPECIMEN herbst, d.r. spence, g. collector number: 5802 Oahu BISH 4/1/1976
93 Lepidium didymum L. E central part of island, between Kaulana Gulch and base of Moaula Growing on eroded slope in Andropogon pertusus grassland. PRESERVED_SPECIMEN Cuddihy, L.W. 423 Kahoolawe BISH 4/25/1980
94 Lepidium didymum L. Southeast Island, part of the Northwest Hawaiian Islands National Wildlife Refuge PRESERVED_SPECIMEN Wegmann, A. s.n. Pearl And Hermes Atoll BISH 10/3/2001
95 Lepidium didymum L. East Maui, Haleakala Crater, Paliku Control Cabin Growing with kikuyu grass in cleared area around cabin. PRESERVED_SPECIMEN Nagata, R. s.n. Maui BISH 7/14/1986
96 Lepidium didymum L. Makapuu Pt., near Waimanalo PRESERVED_SPECIMEN Degener, O. 17226 Oahu US 12/25/2020
97 Lepidium didymum L. hawaii PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 331 Oahu BISH
98 Lepidium didymum L. southeast island, part of the northwest hawaiian islands national wildlife refuge PRESERVED_SPECIMEN wegmann, a. collector number: s.n. Pearl And Hermes Reef BISH 10/3/2001
99 Lepidium didymum L. Alekoko Valley (?) PRESERVED_SPECIMEN s.n. Kauai BISH 6/14/1926
100 Lepidium didymum L. Molokini Islet rare along inner slope. PRESERVED_SPECIMEN Hobdy, R.W. 1228 Maui BISH 2/14/1982
101 Lepidium didymum L. tantalus in honolulu PRESERVED_SPECIMEN forbes, c.n. Oahu BISH 3/31/1909
102 Lepidium didymum L. yard of home H.B. Castle Homestead, Honolulu PRESERVED_SPECIMEN Student at University of Hawaii s.n. Oahu BISH 2/26/1931
103 Lepidium didymum L. Kaeo Cliffs, West Molokai PRESERVED_SPECIMEN Fosberg, F.R. 29583 Molokai BISH 2/22/1948
104 Lepidium didymum L. taauhau 2 PRESERVED_SPECIMEN rock, j.f.c. Hawaii BISH 6/1/1910
105 Lepidium didymum L. Honolulu, Bishop Museum Grounds PRESERVED_SPECIMEN Suehiro, A. s.n. Oahu BISH 2/13/1963
106 Lepidium didymum L. Airplane field, in ,owed grass, Kapaa Quarry Road, Kailua In mowed grass at edge of airplane field. Soil moist, loamy. Full sun. PRESERVED_SPECIMEN Crago, L.M. Kennedy, B. Collector Number: 2005-021 Oahu BISH 1/20/2005
107 Lepidium didymum L. sand island area 7: north of taxi strip on northwest shore of sand island, magazine area Densely forested with Scaevola fringe above beach, (including Frigate Point) PRESERVED_SPECIMEN lamoureux, c.h. collector number: 2197 Midway Atoll BISH 12/15/1962
108 Lepidium didymum L. Kalaupapa National Historical Park Coastal strand vegetation. Plant invading post-deer damage. PRESERVED_SPECIMEN Gagn??, B.H. Trainer, J. Collector Number: 3153 Molokai BISH 9/17/2000
109 Lepidium didymum L. Mokuleia Beach, Waialua PRESERVED_SPECIMEN Degener, O. 24342 Oahu BISH 1/1/1957
110 Lepidium didymum L. alekoko valley (?) PRESERVED_SPECIMEN Kauai BISH 6/14/1926
111 Lepidium didymum L. coral plain below ewa mill & sisal PRESERVED_SPECIMEN forbes, c.n. collector number: 2325 Oahu BISH 5/14/1916
112 Lepidium didymum L. kamehameha school campus PRESERVED_SPECIMEN brown, f.b.h. thompson collector number: 1321 Oahu BISH 12/31/1924
113 Lepidium didymum L. U.H. Campus, Mauka Field Laboratory (Dept. of Agronomy and Soils) A weed growing on a dirt pile in full sun. PRESERVED_SPECIMEN Lau, J. 2250 Oahu BISH 4/15/1986
114 Lepidium didymum L. along the highway near the lands of waialua PRESERVED_SPECIMEN herbst, d.r. spence, g. collector number: 5750 Molokai BISH 3/10/1976
115 Lepidium didymum L. University of Hawaii campus PRESERVED_SPECIMEN St. John, H. 10461 Oahu BISH 4/5/1930
116 Lepidium didymum L. Hawaii PRESERVED_SPECIMEN Rock, J.F.C. 2666 Hawaii BISH
117 Lepidium didymum L. Kamehameha school campus PRESERVED_SPECIMEN Brown, F.B.H. 1321 Oahu BISH 12/31/1924
118 Lepidium didymum L. Coral plain below Ewa and Sisal PRESERVED_SPECIMEN Forbes, C.N. 2325.O Oahu BISH 3/14/1916
119 Lepidium didymum L. Eastern Island PRESERVED_SPECIMEN Frings, H.W. 50 Midway Atoll BISH 4/16/1962
120 Lepidium didymum L. u.h. campus, mauka field laboratory (dept. of agronomy and soils) A weed growing on a dirt pile in full sun. PRESERVED_SPECIMEN lau, j. collector number: 2250 Oahu BISH 4/15/1986
121 Lepidium didymum L. parker ranch, nienie hawaii PRESERVED_SPECIMEN rock, j.f.c. collector number: 8325 Hawaii BISH 6/1/1910
122 Lepidium didymum L. moanalua valley PRESERVED_SPECIMEN forbes, c.n. Oahu BISH 2/18/1909
123 Lepidium didymum L. E central part of island, slope of Moaula PRESERVED_SPECIMEN Cuddihy, L.W. 433 Kahoolawe BISH 4/25/1980
124 Lepidium didymum L. mouth of halawa valley, grassy sand flat, much trampled, PRESERVED_SPECIMEN fosberg, f.r. collector number: 13408 Molokai BISH 12/25/1936
125 Lepidium didymum L. pukoo PRESERVED_SPECIMEN Molokai BISH 6/10/1910
126 Lepidium didymum L. James Campbell National Wildlife Refuge, Kahuku. In vicinity of bird-viewing gazebo Weed on dry ground on levee between ponds. Collection made under sponsorship by Ducks Unlimited, Inc. PRESERVED_SPECIMEN Imada, C.T. 92-17 Oahu BISH 4/5/1992
127 Lepidium didymum L. miki PRESERVED_SPECIMEN munro, g.c. collector number: 254 Hawaii BISH 12/21/1913
128 Lepidium didymum L. SAND ISLAND Area 7: North of taxi strip on northwest shore of Sand Island, Magazine Area Densely forested with Scaevola fringe above beach, (including Frigate Point) PRESERVED_SPECIMEN Lamoureux, C.H. 2197 Midway Atoll BISH 12/15/1962
129 Lepidium didymum L. Koko Crater, dry open slopes PRESERVED_SPECIMEN Christophersen, E. 1301 Oahu BISH 4/6/1930
130 Lepidium didymum L. Kokee State Park. Mohihi Rd. near Waineke Swamp Common along road. PRESERVED_SPECIMEN Flynn, T. 974 Kauai BISH 8/28/1984
131 Lepidium didymum L. Moanalua Valley PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 2/18/1909
132 Lepidium didymum L. Eastern Island PRESERVED_SPECIMEN Frings, H.W. 46 Midway Atoll BISH 4/16/1962
133 Lepidium didymum L. mauna kea state park, 18.4 mi along saddle rd. from jct. hawaii belt rd. (hwy. 190) PRESERVED_SPECIMEN wagner, w.l. mill, s.w.; lammers, t.; weller, s.; sakai, a.; davis-jacobsen, j. collector number: 5575 Hawaii BISH 6/25/1985
134 Lepidium didymum L. mokuauia islet, oahu in central western part of island PRESERVED_SPECIMEN starr, f. starr, k. collector number: 050222-3 Oahu BISH 2/22/2007
135 Lepidium didymum L. Koloa District. Mahaulepu. In Casuarina grove in dumes behind Haula PRESERVED_SPECIMEN Flynn, T. 1969 Kauai BISH 11/25/1986
136 Lepidium didymum L. hawaii PRESERVED_SPECIMEN rock, j.f.c. collector number: 2666 Hawaii BISH
137 Lepidium didymum L. mokuleia beach, waialua PRESERVED_SPECIMEN degener, o. degener, i. collector number: 24342 Oahu BISH 1/8/1957
138 Lepidium didymum L. kalaupapa; peninsula, base of lighthouse stairs along rock wall PRESERVED_SPECIMEN wysong, m.l. collector number: 617 Molokai BISH 2/8/2005
139 Lepidium didymum L. Green Island around LORAN Bldgs and roadsides in that area. PRESERVED_SPECIMEN Herbst, D.R. 6252 Kure Atoll BISH 1/4/1979
140 Lepidium didymum L. Area between Puu Moaula and Lua Makika growing in grassland area PRESERVED_SPECIMEN Higashino, P.K. 8039 Kahoolawe BISH 11/22/1978
141 Lepidium didymum L. Keauhou Ranch Growing along Pu'u O'o Road near a paddock. PRESERVED_SPECIMEN Davis, J. Cuddihy, L.W. Collector Number: 725 Hawaii BISH 5/5/1982
142 Lepidium didymum L. In pineapple field PRESERVED_SPECIMEN Hosaka, E.Y. 470 Lanai BISH 6/23/1931
143 Lepidium didymum L. honopu beach PRESERVED_SPECIMEN davis, j. collector number: s.n. Kauai BISH 7/18/1984
144 Lepidium didymum L. ewa beach PRESERVED_SPECIMEN char, w.p. et al. collector number: prc144 Hawaii BISH 1/30/1982
145 Lepidium didymum L. East Molokai, North Coast, Kukaiwaa on windswept peninsula Growing in a thick carpet of Fimbristylis pycnocephala, low prostrate. PRESERVED_SPECIMEN Hobdy, R.W. 2185 Molokai BISH 8/27/1984
146 Lepidium didymum L. Kaeo Cliffs, West Molokai PRESERVED_SPECIMEN Fosberg, F.R. Collector Number: 29583 Molokai BISH 2/22/1948
147 Lepidium didymum L. oahu sugar co PRESERVED_SPECIMEN swezey, o.h. Oahu BISH 5/20/1930
148 Lepidium didymum L. Honopu Beach PRESERVED_SPECIMEN Joyce s.n. Kauai BISH 7/18/1984
149 Lepidium didymum L. crest of koolau range, head of kamiloiki valley PRESERVED_SPECIMEN egler, f.e. collector number: 37-110a Oahu BISH 6/12/1937
150 Lepidium didymum L. auwahi, s slope of haleakala wet, mossy woods PRESERVED_SPECIMEN forbes, c.n. collector number: 2023 Molokai BISH 3/19/1920
151 Lepidium didymum L. keauhou ranch Growing along Pu'u O'o Road near a paddock. PRESERVED_SPECIMEN davis, j. cuddihy, l.w. collector number: 725 Hawaii BISH 5/5/1982
152 Lepidium didymum L. Uwekahuna Blutt. Found in shade PRESERVED_SPECIMEN Fagerlund, G.O. 452 Hawaii BISH 3/15/1943
153 Lepidium didymum L. lanai PRESERVED_SPECIMEN munro, g.c. collector number: 286 Lanai BISH 12/6/1913
154 Lepidium didymum L. Koko Head PRESERVED_SPECIMEN Swezey, O.H. s.n. Oahu BISH 1920-00-00
155 Lepidium didymum L. Palawai, Kaluanui Flats PRESERVED_SPECIMEN Herbst, D.R. 5802 Lanai BISH 4/1/1976
156 Lepidium didymum L. eastern-central part of island, between kaulana gulch and base of moaula Growing on eroded slope in Andropogon pertusus grassland. PRESERVED_SPECIMEN cuddihy, l.w. clarke, g. collector number: 423 Kahoolawe BISH 4/25/1980
157 Lepidium didymum L. Tantalus in Honolulu PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 3/31/1909
158 Lepidium didymum L. Molokai Pukoo PRESERVED_SPECIMEN Faurie, U.J., Abbé s.n. Molokai BISH 6/1/1910
159 Lepidium didymum L. Mouth of Halawa Valley, grassy sand flat, much trampled, PRESERVED_SPECIMEN Fosberg, F.R. 13408 Molokai BISH 12/25/1936
160 Lepidium didymum L. kaimuki, honolulu, hawaii PRESERVED_SPECIMEN topping, d.l. collector number: 2995 Oahu BISH 1/21/1925
161 Lepidium didymum L. east maui, haleakala crater, paliku control cabin Growing with kikuyu grass in cleared area around cabin. PRESERVED_SPECIMEN nagata, r. collector number: s.n. Maui BISH 7/14/1986
162 Lepidium didymum L. Kaimuki, Honolulu, Hawaii PRESERVED_SPECIMEN Topping, D.L. 2995 Oahu BISH 1/21/1925
163 Lepidium didymum L. Auwahi, S slope of Haleakala PRESERVED_SPECIMEN Forbes, C.N. 2023.M Maui BISH 3/19/1920
164 Lepidium didymum L. kaeo cliffs, west molokai PRESERVED_SPECIMEN fosberg, f.r. collector number: 29583 Molokai BISH 2/22/1948
165 Lepidium didymum L. james campbell national wildlife refuge, kahuku. in vicinity of bird-viewing gazebo Weed on dry ground on levee between ponds. Collection made under sponsorship by Ducks Unlimited, Inc. PRESERVED_SPECIMEN imada, c.t. engilis, a.; dr.duk, mccafferty, t. collector number: 92-17 Oahu BISH 4/5/1992
166 Lepidium didymum L. yard of home h.b. castle homestead, honolulu PRESERVED_SPECIMEN student at university of hawaii Oahu BISH 2/26/1931
167 Lepidium didymum L. airplane field, in ,owed grass, kapaa quarry road, kailua Mesic forest PRESERVED_SPECIMEN crago, l.m. kennedy, b. collector number: 2005-021 Oahu BISH 1/20/2005
168 Lepidium didymum L. West Maui, Lahaina District, Honokohau, Pohakupule PRESERVED_SPECIMEN B. H. Gagn???? & J. Trainer 3153 Maui HAPI 9/17/2000
169 Lepidium didymum L. koko head, maunalua PRESERVED_SPECIMEN krauss, n.h. Maui BISH 3/25/1933
170 Lepidium didymum L. Miki PRESERVED_SPECIMEN Munro, G.C. 254 Lanai BISH 12/21/1913
171 Lepidium didymum L. coral plain below ewa and sisal PRESERVED_SPECIMEN forbes, c.n. collector number: 2325 Oahu BISH 3/14/1916
172 Lepidium didymum L. Lanai City PRESERVED_SPECIMEN Degener, O. 28557 Lanai BISH 8/15/1963
173 Lepidium didymum L. Ewa Plains Survey-U.S.F.&W. Campbell Industrial Park area PRESERVED_SPECIMEN Higashino, P.K. 8193 Oahu BISH 2/1/1979
174 Lepidium didymum L. Popoia Islet, Oahu on west side near coast growing in sands inside limestone hole. PRESERVED_SPECIMEN Starr, F. 050223-7 Oahu BISH 2/23/2005
175 Lepidium didymum L. Upper Makakilo, adjacent to developed subdivision at end of Pueonani Street Open area in solid Panicum maximum grassland, with /Waltheria indica, Sida fallax, Eleusine indica, Verbesina encelioides, Sonchus oleraceus. PRESERVED_SPECIMEN Imada, C.T. 2004-24 Oahu BISH 3/9/2004
176 Lepidium didymum L. Parker ranch, Pokanaka, plowed field PRESERVED_SPECIMEN Ewart III, G.R. 174 Hawaii BISH 4/21/1932
177 Lepidium didymum L. laniloa (?) PRESERVED_SPECIMEN cooke, c.m. Oahu BISH 2/1/1919
178 Lepidium didymum L. pacific remote islands national wildlife refuge, in flat rocky area, ca 200 m from beach Collected at former guano mining site, in area with high concentration of albatross nestings. Most species in area are alien: Sporobolus, Solanum americanum, Conyza bonariensis. PRESERVED_SPECIMEN martin, a. collector number: 2 Laysan BISH 3/12/2002
179 Lepidium didymum L. Pacific Remote Islands National Wildlife Refuge, in flat rocky area, ca 200 m from beach Collected at former guano mining site, in area with high concentration of albatross nestings. Most species in area are alien: Sporobolus, Solanum americanum, Conyza bonariensis. PRESERVED_SPECIMEN Martin, A. 2 Laysan BISH 3/12/2002
180 Lepidium didymum L. along road leading to kaluanui valley moist, wooded PRESERVED_SPECIMEN corn, c. collector number: s.n. Oahu BISH 3/21/1978
181 Lepidium didymum L. Ewa Beach PRESERVED_SPECIMEN Char, W.P. PRC 144 Oahu BISH 1/30/1982
182 Lepidium didymum L. Laniloa (?) PRESERVED_SPECIMEN Cooke, C.M. s.n. Oahu BISH 2/1/1919
183 Lepidium didymum L. Taauhau 2 PRESERVED_SPECIMEN Rock, J.F.C. s.n. Hawaii BISH 6/1/1910
184 Lepidium didymum L. Honopu Beach PRESERVED_SPECIMEN Davis, J. Collector Number: s.n. Kauai BISH 7/18/1984
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