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Boerhavia repens

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-CoreEudicot Order: Caryophyllales Family: Nyctaginaceae Genus: Boerhavia

alena, anena (ni‘ihau), nena

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Description

Key Characters:

Growth Form: Relatively robust perennial herb.

Stems: Several prostrate stems radiating from the root crown, sparingly to many-branched.

Roots: Roots thickened.

Leaves: Leaves simple. Opposite. Blades usually ovate to elliptic–ovate or suborbicular, 1–5 cm long, 0.5–5 cm wide. Apex obtuse to acute. Base rounded to truncate or subcordate. Surfaces sparsely puberulous to glabrous, glandular punctate. Margins entire to gently sinuate. Petioles 0.3–2 cm long. Stipules absent.

Flowers: Flowers in axillary pedunculate cymes or pseudoumbels, usually from alternate axils, the peduncles appearing to be crowded aside from the axil proper by the growth of an axillary branch, which very soon bears axillary cymes, leaves often reduced gradually distally, creating a paniculate appearance, but with a clear central axis, peduncles branched only near tip, branching basically umbelliform, each branch ending in a glomerule of flowers, or the branch system reduced to a single umbelliform to glomerate cluster of flowers, or the peduncle bearing 1 pair of leaves and a glomerate cluster of flowers. involucre absent. Flowers bisexual (perfect) usually small. Calyx shortly 5-lobed, 2–2.5 mm long, strongly constricted below the middle, lower part glandular, 5–costate, limb pink or white, campanulate. Corolla (petals) absent. Stamens 2–4, subequal with perianth. Ovary superior, sessile; style filiform, unbranched; stigma capitate or somewhat disk–shaped, exserted.

Fruit: Anthocarps clavate to ellipsoid; 3–4 mm long; apex rounded; 5–ribbed; very glandular to subglabrous or glabrous. Seeds 1 per fruit; closely enclosed by the thin ovary wall.

Ploidy: 2n = 52*

Habitat: Formerly much more common; on shores and moderately dry coastal areas and leeward; at least semi–dry lower slopes.

Elevation Range:

Historical Distribution

Images

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Synonyms (35)

Boerhavia angustifolia Span. (1841)
Boerhavia ascendens Schumach. (1827)
Boerhavia bicolor Ehrenb. ex Schweinf. (1867)
Boerhavia cortensis Ehrenb. ex Schweinf. (1867)
Boerhavia diffusa var. eurepens Domin (1926)
Boerhavia diffusa var. gymnocarpa Heimerl (1937)
Boerhavia diffusa var. minor (Delile) Cufod. (1969)
Boerhavia diffusa var. pseudotetrandra Heimerl (1937)
Boerhavia diffusa var. sandwicensis Heimerl (1937)
Boerhavia diffusa var. vulvariifolia (Poir.) Heimerl (1897)
Boerhavia eglandulosa Ehrenb. ex Schweinf. (1867)
Boerhavia hirsuta Willd. (1794)
Boerhavia insularis Bojer (1837)
Boerhavia maroccana Ball (1875)
Boerhavia minutiflora Span. (1841)
Boerhavia mutabilis (Misapplied)
Boerhavia periplocifolia Sieber ex Choisy (1849)
Boerhavia pubescens R.Br. (1810)
Boerhavia repens var. amphilahensis A.Terracc. (1894)
Boerhavia repens var. glabra Choisy (1849)
Boerhavia repens var. maris-indici Fosberg (1979)
Boerhavia repens var. minor Delile (1813)
Boerhavia repens var. pubescens (R.Br.) Fosberg (1978)
Boerhavia repens var. viscosa Choisy (1849)
Boerhavia rupestris Ehrenb. ex Schweinf. (1867)
Boerhavia sessiliflora Steud. (1840)
Boerhavia suberosa C.Sm. (1818)
Boerhavia sumata Zipp. ex Span. (1841)
Boerhavia surinamensis Miq. (1844)
Boerhavia tetrandra (Misapplied)
Boerhavia tomentosa Ehrenb. ex Schweinf. (1867)
Boerhavia undulata Ehrenb. ex Schweinf. (1867)
Boerhavia viscosa Fresen. (1833)
Boerhavia vulvariifolia Poir. (1804)
Dantia nubica Lippert ex Choisy (1849)

Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Ka'ula Indigenous
Mokupāpapa (Kure atoll) Indigenous
Ni'ihau(Incl. Lehua) Indigenous
Kuaihelani (Midway Atoll) Indigenous
Kaua'i Indigenous
Manawai (Pearl & Hermes Atoll) Indigenous
O'ahu Indigenous
Kapou(Lisianski Island) Indigenous
Molokai Indigenous
Kamole(Laysan Island) Indigenous
Lana'i Indigenous
Kaho'olawe Indigenous
Lalo (French Frigate Shoals) Indigenous
Maui Indigenous
Hawai'i Indigenous
Nihoa Indigenous

Dispersal Agents


Pollinators

Bibliography

Name Published In: Sp. Pl.: 3 (1753)

Other References

Wagner et al. 1990:981 (Ku, Mi, PH, Li, La, FF, Ni, K, O, Mo, L, M, Ka, H); Evenhuis & Eldredge 2004:55 (N); Wood & LeGrande 2006:24 (Le); Bryan Jr. 3a BISH (N†, 1923); Caum 13 BISH (Kl†, 1932); Note: Kl added, based on BISH spm. (E.L. Caum 13, 17.viii.1932)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Boerhavia repens Kahuku near beach. Preserved_Specimen Plants luxuriant after rainy season. Bishop, B. Oahu BISH 1963-06-02
2 Boerhavia repens Lahaina, !/2 mi E, along side of Hwy. 34 with Waltheria, Desmanthus Preserved_Specimen prostrate Ishikawa, S. 358 Maui BISH 1973-12-03
3 Boerhavia repens Kaupuu Sand dunes Preserved_Specimen Christensen, C. 142 Niihau BISH 1977-02-12
4 Boerhavia repens E crest sandy Preserved_Specimen Daniel, C.W. 35 Laysan Island BISH 1959-07-27
5 Boerhavia repens E side near beach sandy Preserved_Specimen Tsuda, R.T. 501 Laysan Island BISH 1963-12-05
6 Boerhavia repens North Island central plain to beach, with Tribulus, Lepturus, Lepidium, Sicyos, Solanum, Eragrostis, Scaevola, Messerschmidia Preserved_Specimen Young, A.L. 130 Pearl & Hermes Atoll BISH 1964-08-20
7 Boerhavia repens Seal-Kittery Island Preserved_Specimen decumbent herb with pink flowers Herbst, D.R. 6465 Pearl & Hermes Atoll BISH 1980-07-05
8 Boerhavia repens Manana Island, off Waimanalo Near the beach. Preserved_Specimen Strandtmann, R.W. Oahu BISH 1979-06-12
9 Boerhavia repens Honolulu District, Koko Head Preserved_Specimen Indigenous. Small population on slope facing Hanauma Bay, near Ihiihilauakea Crater. Bruegmann, M.M. 149 Oahu BISH 1984-01-29
10 Boerhavia repens West Maui, Waiehu, parking lot by pavillon Preserved_Specimen prostrate, flowers white in compact clusters Hobdy, R.W. 1982 Maui BISH 1984-02-23
11 Boerhavia repens Kalaupapa National Historic Park. Airport. 100 m W of start of runway Preserved_Specimen slender prostrate herb Wysong, M.L. 590 Molokai BISH 2005-01-27
12 Boerhavia repens Middle Island Preserved_Specimen flowers pinkish, stamens 2-3 Caum, E.L. 47 Pearl & Hermes Atoll BISH 1923-04-27
13 Boerhavia repens Kaohikaipu Islat, Oahu Preserved_Specimen Occasional. Twining in vegetation here and there. Starr, F. 502244 Oahu BISH 2005-02-24
14 Boerhavia repens SE Island Preserved_Specimen creeping, prostrate; flowers tiny, pinkish, 4 stamens; fruit small cylindrical Caum, E.L. 41 Pearl & Hermes Atoll BISH 1923-04-26
15 Boerhavia repens Manana Island. S side of crater Preserved_Specimen Light lavender flrs. Long, C.R. 1683 Oahu BISH 1964-05-02
16 Boerhavia repens Puu Koae Island off the south coast of Kahoolawe. Preserved_Specimen Only one plant found. This is the blunt-leaved type with white flowers. This plant seemed unusually coarsely pilose throughout. Hobdy, R.W. 1038 Kahoolawe BISH 1981-03-21
17 Boerhavia repens Tern Islet Preserved_Specimen Lamoureux, C.H. 1671 French Frigate Shoals BISH 1961-09-02
18 Boerhavia repens Diamond Head Crater Preserved_Specimen Prostrate shrub. Flower pink. Christophersen, E. 1325 Oahu BISH 1930-05-11
19 Boerhavia repens Papawai Pt., Ukumehame Preserved_Specimen crawling vine, 2 m x 1 cm Woolford, H.A. 79 Maui BISH 1955-12-18
20 Boerhavia repens near lagoon, NW part Preserved_Specimen trailing herb Herbst, D.R. 6514 Laysan Island BISH 1980-07-13
21 Boerhavia repens Tern Islet Preserved_Specimen decumbent herb with white flowers Herbst, D.R. 6234 French Frigate Shoals BISH 1978-09-11
22 Boerhavia repens SE Island Preserved_Specimen tiny pinkish flower; fruit small cylindrical; short peduncles Caum, E.L. 40 Pearl & Hermes Atoll BISH 1923-04-26
23 Boerhavia repens National Park, Kau, Halape beach, some distance from W shelter Preserved_Specimen trailing, prostrate Herat, T. 850 Hawaii BISH 1974-06-30
24 Boerhavia repens Kaohikaipu Islet, Makapuu Point Preserved_Specimen Lamoureux, C.H. 4086 Oahu BISH 1967-08-10
25 Boerhavia repens Kure Atoll Preserved_Specimen Lamoureux, C.H. 1882 Kure Atoll BISH 1961-09-12
26 Boerhavia repens along edges of lagoon Preserved_Specimen erect shrub 12-18 high; flower pink Wilder, G.P. 471 Laysan Island BISH 1930-08-10
27 Boerhavia repens Laysan inland sand Preserved_Specimen Bryan, W.A. Laysan Island BISH 1903-04-01
28 Boerhavia repens French Frigate Shoals Preserved_Specimen not viscid Caum, E.L. 87 French Frigate Shoals BISH 1923-04-22
29 Boerhavia repens Sandy banks above bay at Papakolea Preserved_Specimen Prostrate vine Lane, I.E. 49551 Hawaii BISH 1949-12-26
30 Boerhavia repens Alau Island, eastern coast of East Maui Preserved_Specimen Leaves bright green above, white below; flowers pure white Hobdy, R.W. 1300 Maui BISH 1982-03-07
31 Boerhavia repens Kekepa, Oahu Preserved_Specimen Occasional. Scattered here and there. Starr, F. 504183 Oahu BISH 2005-04-18
32 Boerhavia repens Lehua Rock Sida fallax, Jacquemontia sandwicensis, Sicyos, Waltheria indica, Abutilon grandifolium, Portulaca oleracea, Doryopteris decipiens, Boerhavia repens, Artemisia australis Preserved_Specimen Wood, K.R. 9219 Lehua Rock BISH 2001-12-20
33 Boerhavia repens Ewa Beach Preserved_Specimen Char, W.P. 175 Oahu BISH 1982-02-24
34 Boerhavia repens Kaena Point Preserved_Specimen Forbes, C.N. 1648 Oahu BISH 1911-02-25
35 Boerhavia repens Kure Atoll Preserved_Specimen Lamoureux, C.H. 1876 Kure Atoll BISH 1961-09-12
36 Boerhavia repens Oahu Preserved_Specimen Mann, H. 111 Oahu BISH 1864-01-01
37 Boerhavia repens Tern Islet Preserved_Specimen 15 cm high Lamoureux, C.H. 1675 French Frigate Shoals BISH 1961-09-02
38 Boerhavia repens Manana Island Preserved_Specimen Trailing herb. Lamoureux, C.H. 2568 Oahu BISH 1964-02-29
39 Boerhavia repens Lehua Preserved_Specimen Caum, E.L. 28 Lehua Rock BISH 1931-04-22
40 Boerhavia repens Kaula Rock Preserved_Specimen Caum, E.L. 13 Kaula Rock BISH 1932-08-17
41 Boerhavia repens Diamond Head Preserved_Specimen Forbes, C.N. 1075 Oahu BISH 1909-01-26
42 Boerhavia repens Manana Island Preserved_Specimen Lamoureux, C.H. 2573 Oahu BISH 1964-02-29
43 Boerhavia repens inner NE slope Preserved_Specimen Lamoureux, C.H. 1690 Laysan Island BISH 1961-09-05
44 Boerhavia repens inner NE slope Preserved_Specimen Lamoureux, C.H. 1691 Laysan Island BISH 1961-09-05
45 Boerhavia repens inner NE slope Preserved_Specimen Lamoureux, C.H. 1691 Laysan Island BISH 1961-09-05
46 Boerhavia repens Limestone Point Preserved_Specimen Munro, G.C. 259 Lanai BISH 1918-12-13
47 Boerhavia repens Green Island Preserved_Specimen Lamoureux, C.H. 2785 Kure Atoll BISH 1964-08-14
48 Boerhavia repens Kalawao District, edge of landing field, Kalaupapa airport Preserved_Specimen Wagner, W.L. 4940 Molokai BISH 1983-08-02
49 Boerhavia repens Green Island Preserved_Specimen Herbst, D.R. 6278 Kure Atoll BISH 1979-01-04
50 Boerhavia repens Popoia Island, Kailua Bay Preserved_Specimen Herbst, D.R. 6022 Oahu BISH 1978-02-07
51 Boerhavia repens Keaoi Island, off Halepe Preserved_Specimen Fagerlund, G.O. 356 Hawaii BISH 1943-02-16
52 Boerhavia repens Kaula Rock Preserved_Specimen Caum, E.L. 13 Kaula Rock BISH 1932-08-17
53 Boerhavia repens Lualualei Beach Preserved_Specimen Forbes, C.N. 1922 Oahu BISH 1914-07-01
54 Boerhavia repens Queen's Beach Preserved_Specimen Bennett, T. 135 Oahu BISH 1982-01-28
55 Boerhavia repens East Maui, Nuu Parcel under lower Wiliwili grove west of Na Kula LZ Preserved_Specimen Welton, P. 2323 Maui BISH 2013-01-03
56 Boerhavia repens inner NE slope Preserved_Specimen Lamoureux, C.H. 1689 Laysan Island BISH 1961-09-05
57 Boerhavia repens Ranch Preserved_Specimen Rock, J.F.C. 7034 Molokai BISH 1910-04-01
58 Boerhavia repens Ranch Preserved_Specimen Rock, J.F.C. 7034 Molokai BISH 1910-04-01
59 Boerhavia repens North Island Preserved_Specimen Conant, S. 367 Pearl & Hermes Atoll BISH 1983-08-10
60 Boerhavia repens barking Sands, Mana Preserved_Specimen Degener, O. 30277 Kauai BISH 1926-06-13
61 Boerhavia repens Middle Island Preserved_Specimen Caum, E.L. 48 Pearl & Hermes Atoll BISH 1923-04-27
62 Boerhavia repens Mokulua, North Island, Lanikai Preserved_Specimen Herbst, D.R. 5989 Oahu BISH 1978-02-07
63 Boerhavia repens Kapua, Kona Preserved_Specimen Forbes, C.N. 353 Hawaii BISH 1911-07-14
64 Boerhavia repens inner NE slope Preserved_Specimen Lamoureux, C.H. 1690 Laysan Island BISH 1961-09-05
65 Boerhavia repens Koko Head Preserved_Specimen Rock, J.F.C. Oahu BISH 1912-01-01
66 Boerhavia repens Central open plain Preserved_Specimen Clay, H.F. Kure Atoll BISH 1959-10-03
67 Boerhavia repens Ewa Beach Preserved_Specimen Woolford, H.A. Oahu BISH 1956-12-15
68 Boerhavia repens Molokai Preserved_Specimen Stokes, J.F.G. Molokai BISH 1909-01-01
69 Boerhavia repens Na Laau Hawaii Preserved_Specimen Munro, G.C. Oahu BISH 1961-11-18
70 Boerhavia repens Mokuauia Island, near Laie Preserved_Specimen Corn, C. Oahu BISH 1977-11-12
71 Boerhavia repens Waimanalo Beach Preserved_Specimen Suehiro, A. Oahu BISH 1932-05-15
72 Boerhavia repens Ke Kuaaina Preserved_Specimen Munro, G.C. Oahu BISH 1961-11-25
73 Boerhavia repens Kaena Point Preserved_Specimen Christophersen, E. Oahu BISH 1930-12-14
74 Boerhavia repens Palolo Preserved_Specimen Lyon, H.L. Oahu BISH 1908-10-25
75 Boerhavia repens Laysan Preserved_Specimen Butler, G. Laysan Island BISH 1959-04-30
76 Boerhavia repens Kii Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
77 Boerhavia repens Na Laau Hawaii Preserved_Specimen Munro, G.C. Oahu BISH 1961-12-16
78 Boerhavia repens Tern Islet Preserved_Specimen Svihla, A. French Frigate Shoals BISH 1956-02-11
79 Boerhavia repens Koko Head Preserved_Specimen Rock, J.F.C. Oahu BISH 1912-01-01
80 Boerhavia repens Near landing Preserved_Specimen Handy, E.S. Niihau BISH 1931-08-14
81 Boerhavia repens Lisianski Island Preserved_Specimen Richardson, F. Lisianski Island BISH 1954-03-26
82 Boerhavia repens Waimanalo Preserved_Specimen Neal, M.C. Oahu BISH 1930-08-06
83 Boerhavia repens Kaena Preserved_Specimen Pearsall, G.A. Oahu BISH 1958-05-03
84 Boerhavia repens Molokini Islet Preserved_Specimen Richardson, F. Maui BISH 1954-06-01
85 Boerhavia repens Rabbit Island Preserved_Specimen Neal, M.C. Oahu BISH 1930-03-16
86 Boerhavia repens Ewa Beach, lighthouse Preserved_Specimen Woolford, H.A. Oahu BISH 1956-12-26
87 Boerhavia repens Islet off Malaekahana-Mokuauia Preserved_Specimen Neal, M.C. Oahu BISH 1938-06-19
88 Boerhavia repens Near landing Preserved_Specimen Handy, E.S. Niihau BISH 1931-08-14
89 Boerhavia repens Mokapu Beach Preserved_Specimen Swezey, O.H. Oahu BISH 1935-02-10
90 Boerhavia repens Manana Islet, off Waimanalo Preserved_Specimen Strandtmann, R.W. Oahu BISH 1979-06-12
91 Boerhavia repens Laysan Preserved_Specimen Schauinsland, H.H. Laysan Island BISH 1896-01-01
92 Boerhavia repens S Kona, Kapua Bay Preserved_Specimen Higashino, P.K. 6544 Hawaii BISH 1977-07-01
93 Boerhavia repens Lehua Rock Preserved_Specimen St.John, H. 22852 Lehua Rock BISH 1947-08-16
94 Boerhavia repens Kure Atoll Preserved_Specimen Lamoureux, C.H. 1877 Kure Atoll BISH 1961-09-12
95 Boerhavia repens Kure Atoll Preserved_Specimen Lamoureux, C.H. 1881 Kure Atoll BISH 1961-09-12
96 Boerhavia repens Koko Head, Maunalua dry ridge Preserved_Specimen Inafuku, R. Oahu BISH 1931-04-27
97 Boerhavia repens Moomomi sand dunes Preserved_Specimen prostrate vine, leaves thin Fosberg, F.R. 13432 Molokai BISH 1936-12-26
98 Boerhavia repens Kaena Point near shore Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-01-16
99 Boerhavia repens NW end of lagoon Eragrostis and Ipomoea; sand Preserved_Specimen prostrate creeping herb, pinkish white flowers Young, A.L. 147 Laysan Island BISH 1964-09-17
100 Boerhavia repens Barbers Point, Kapolei. Coastal strand, ca. 0.5 km NW of Barbers Point Lighthouse growing with Atriplex suberecta in sandy soil. Preserved_Specimen Perennial herb, sprrawling, ca. 60-70 cm long; corolla pinkish-white, 2-3 plants seen. Ross, M.C. 1956 Oahu BISH 2023-06-30
101 Boerhavia repens Green Island central plain Preserved_Specimen Lamoureux, C.H. 2277 Kure Atoll BISH 1962-12-16
102 Boerhavia repens Ocean Island open Preserved_Specimen vine, prostrate, to 1/3 m high Caum, E.L. 11 Kure Atoll BISH 1923-04-18
103 Boerhavia repens Makua Top of beach. Preserved_Specimen Prostrate creeper. St.John, H. 9947 Oahu BISH 1929-10-20
104 Boerhavia repens East Island coral sand Preserved_Specimen Long, C.R. 2493 French Frigate Shoals BISH 1964-09-27
105 Boerhavia repens Koolau Poko District; Mokapu Peninsula Marine Corps Base Hawaii, Kaneohe Bay, Pyramid Rock Beach Occasional on dry low slopes behind the beach. Preserved_Specimen Prostrate perennial herb with a fleshy tap-root with several stems radiating from it. Flws pink. Herbst, D.R. 9733 Oahu BISH 1995-12-05
106 Boerhavia repens Kahinaakalani, E of Hawaiian Homes Recreation Center raised reef with thin soil, basaltic boulders and lava outcrops Preserved_Specimen Wagner, W.L. 4911 Molokai BISH 1983-07-29
107 Boerhavia repens Maunalua On dry slope Preserved_Specimen Chong, M. Oahu BISH 1933-03-25
108 Boerhavia repens Mokuauia Island, Laie thin soil on reef rock Preserved_Specimen fls. pale pink Fosberg, F.R. 14178 Oahu BISH 1937-07-04
109 Boerhavia repens Oahu Coral Flats. Preserved_Specimen Flowers pink. Woolford, H.A. Oahu BISH 1957-02-24
110 Boerhavia repens Makua Top of beach. Preserved_Specimen Prostrate creeper. St.John, H. 9947 Oahu BISH 1929-10-20
111 Boerhavia repens Tern Islet white coral sand Preserved_Specimen sends roots down 1 m, top spreads over surface; small elongate oval seed vessel which is viscous Judd, C.S. 9 French Frigate Shoals BISH 1923-06-26
112 Boerhavia repens West Molokai, Ilio Point sand Preserved_Specimen Hobdy, R.W. 2369 Molokai BISH 1985-04-23
113 Boerhavia repens Grass Island with Tribulus, Solanum Preserved_Specimen Herbst, D.R. 6460 Pearl & Hermes Atoll BISH 1980-07-05
114 Boerhavia repens Lisianski Island beach to interior; with Scaevola, Eragrostis, Ipomoea Preserved_Specimen trailing herb with pink to white flowers Young, A.L. 134 Lisianski Island BISH 1964-08-21
115 Boerhavia repens Kaena Point sand dune Preserved_Specimen prostrate, leaves fleshy, fls. pink; fruits glutinous Fosberg, F.R. 13146 Oahu BISH 1936-05-31
116 Boerhavia repens On Keomuku coastline, at Lae Hi On raised coral plain. Preserved_Specimen Hobdy, R.W. 1978 Lanai BISH 1984-02-20
117 Boerhavia repens Queen's Beach rocky, sandy coastal plain. Preserved_Specimen Wagner, W.L. 5337 Oahu BISH 1984-02-19
118 Boerhavia repens Puu Koae Islet Lower W slope in barren rubble Preserved_Specimen Hobdy, R.W. 1959 Kahoolawe BISH 1984-02-11
119 Boerhavia repens Kalani, Moomomi, Kaluakoi on sand dunes. Preserved_Specimen flowers pink, stamens 2-3 St.John, H. 25154 Molokai BISH 1953-12-21
120 Boerhavia repens SE Island, E portion sand Preserved_Specimen sprawling Long, C.R. 2271 Pearl & Hermes Atoll BISH 1964-09-16
121 Boerhavia repens 1.5 mi W Moomomi Beach sand dunes Preserved_Specimen Aidem, J. Molokai BISH 1972-02-17
122 Boerhavia repens Kaena Point Sand dunes near sea-level. Preserved_Specimen Common, brs. spr. to pr., fls. whitish. Webster, G.L. 13848 Oahu BISH 1966-06-22
123 Boerhavia repens Kaluakoi, Kawakiu Road Sand dune, with wind. Preserved_Specimen Prostrate, stems 6-8 ft. long. St.John, H. 12451 Molokai BISH 1932-12-22
124 Boerhavia repens E of Barber's Point Lighthouse and S of Gilbert coral sand beach Preserved_Specimen Harris, J.A. 242188 Oahu BISH 1924-09-02
125 Boerhavia repens open area on W side Scaevola; sand Preserved_Specimen Long, C.R. 2237 Kure Atoll BISH 1964-09-14
126 Boerhavia repens Ahupuaa: Kealia; Kealia trail growing on koa in dry forest. Preserved_Specimen Voluble vine Yamada, R. 20 Oahu BISH 1960-11-20
127 Boerhavia repens Kaena Pt betw end of rd & Natural Area Reserve Coastal strand. Preserved_Specimen perennial; stems prostrate; flower white. Annable, C.R. 3136 Oahu BISH 1996-11-13
128 Boerhavia repens Kapapa, Oahu under partial shade Preserved_Specimen Common to dominant. Forming large mats under partial shade. Starr, F. 5041841 Oahu BISH 2005-04-18
129 Boerhavia repens Milolii Beach, Na Pali Coast growing near the sand with such grasses as Cynodon and Sporobolus with some localized patches of Heteropogon contortus. Preserved_Specimen A prostrate vine Corn, C. 444 Kauai BISH 1979-07-19
130 Boerhavia repens North end sandy soil Preserved_Specimen 20 cm prostrate vine; flowers pink Laysaon, C.W. 23 Laysan Island BISH 1959-07-26
131 Boerhavia repens 300 yds. from N shore sandy, near top of rise; with Tribulus, Scaevola, Eragrostis Preserved_Specimen Shelton, P.C. 402 Lisianski Island BISH 1966-06-16
132 Boerhavia repens Camp Dry, hard, cobble area. Preserved_Specimen Native. Often forming mats with long branches. Flowers are small and white or light pink in color. Al-Bayer, S. Kure Atoll BISH 2018-12-29
133 Boerhavia repens North Island, N side sand Preserved_Specimen Long, C.R. 2310 Pearl & Hermes Atoll BISH 1964-09-17
134 Boerhavia repens Nihoa Island sand beach Preserved_Specimen trailing Bryan Jr., E.H. 3 Nihoa BISH 1923-06-14
135 Boerhavia repens Near Honuapo aa lava flow Preserved_Specimen Degener, O. 30279 Hawaii BISH 1926-07-25
136 Boerhavia repens near Puu Keke, Mahaulepu dunes along beach; with Scaevola taccada and Vitex Preserved_Specimen prostrate spreading as much as 1.5 m in all directions from a central tap root Fay, J.J. 163 Kauai BISH 1974-01-14
137 Boerhavia repens Mokapu Peninsula, Pyramid Rock, Heeia Flats, Heleloa lava hill Preserved_Specimen Fosberg, F.R. 10566 Oahu BISH 1933-03-25
138 Boerhavia repens highway between Laie & Kahuku Slope: Flat. Soil: Loam. Preserved_Specimen Inconspicuous weeds. Leaves are 0.5-1.5 in. long opposite. Flowers are borne on slender stalks. Lee, P. 2 Oahu BISH 1975-02-08
139 Boerhavia repens Nualolo-Kai, Waimea District shore flats Preserved_Specimen trailing, flowers white, stamens 2 St.John, H. 26691 Kauai BISH 1965-04-24
140 Boerhavia repens Diamond Head Crater Southern crest of crater. Preserved_Specimen Corn, C. Oahu BISH 1977-12-13
141 Boerhavia repens Whale-Skate Island coarse sand and coral rubble; with Lepturus, Tribulus Preserved_Specimen stems from a large underground tuberous rootstock Carlquist, S. 2274 French Frigate Shoals BISH 1966-09-13
142 Boerhavia repens West Molokai, Moomomi flat eroded volcanic soil back of dunes Preserved_Specimen prostrate; leaves pallid beneath; flowers pink, stamens 2-3 Fosberg, F.R. 29607 Molokai BISH 1948-02-23
143 Boerhavia repens Tern Islet, SE part sand Preserved_Specimen flowers were lavender Long, C.R. 2508 French Frigate Shoals BISH 1964-09-28
144 Boerhavia repens inner cresent Panicum torridum, Chenopodium murale, Ageratum conyzoides, Portulaca oleracea, Jacquemontia ovalifolia, Sicyos maximowiczii Preserved_Specimen sub shrub; stems decumbent; leaves medium green; corolla pink Wood, K.R. 11344 Lehua Rock BISH 2005-05-14
145 Boerhavia repens Nihoa Island among rocks above tide zone Preserved_Specimen sprawling; flowers pinkish white Caum, E.L. 79 Nihoa BISH 1923-06-17
146 Boerhavia repens Ocean Island open in Scaevola Preserved_Specimen vine, prostrate, to 1/3 m high Caum, E.L. 6 Kure Atoll BISH 1923-04-17
147 Boerhavia repens Kaena Point on sand flats. associates include Sida fallax, Jacquemontia sandwicensis, Heliotropium anomalum, and Lipochaeta integrifolia Preserved_Specimen common creeper Takeuchi, W.N. 32 Oahu BISH 1984-01-28
148 Boerhavia repens Sealife Park Beach, N. of Makapuu Point On sandy dunes Preserved_Specimen Prostrate, much branched, fleshy herb. Stems often reddish. Leaves light green, almost white below. Flowers white. van Royen, P. 10188 Oahu BISH 1967-09-04
149 Boerhavia repens Kalaeahole clay and rock below low ocean cliffs Preserved_Specimen Degener, O. 24248 Lanai BISH 1957-08-11
150 Boerhavia repens 300 yds from N shore sandy, near top of rise; with Tribulus, Scaevola, Eragrostis Preserved_Specimen prostrate Shelton, P.C. 402 Lisianski Island BISH 1966-06-16
151 Boerhavia repens N Kona Talus, cliff Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
152 Boerhavia repens SE Island, S half of the E part Tribulus Preserved_Specimen dense mat Herbst, D.R. 6492 Pearl & Hermes Atoll BISH 1980-07-07
153 Boerhavia repens Kapapa Islet, Kaneohe Bay Common on coral sandy soil on raised reef Preserved_Specimen Leaves pale beneath. Fosberg, F.R. 14023 Oahu BISH 1937-06-06
154 Boerhavia repens Kalani, Kaluakoi inland coral sand dunes Preserved_Specimen trailing, flowers pink, stamens 3-4 St.John, H. 23485 Molokai BISH 1948-12-24
155 Boerhavia repens Makua, top of beach Beach. Preserved_Specimen Prostrate creeper. St.John, H. 9947 Oahu BISH 1929-10-20
156 Boerhavia repens Laysan inland sand Preserved_Specimen Bryan, W.A. Laysan Island BISH 1903-04-01
157 Boerhavia repens Kapapa Islet, Kaneohe Bay Sandy coral soil on raised reef Preserved_Specimen Prostrate; leaves pallid beneath; flowers pinkish. Fosberg, F.R. 14024 Oahu BISH 1937-06-06
158 Boerhavia repens Alau, islet SW of Hana coastal vegetation; Scaevola sericea, Sesuvium portulacastrum, Vigna marina, Portulaca lutea, Schiedea globosa Preserved_Specimen herb, prostrate; stems yellow, slight pink tinge; corolla white, yellow tinge Wood, K.R. 11186 Maui BISH 2005-04-05
159 Boerhavia repens Kiekie, 0.5 mi NE Sand near shore Preserved_Specimen Trailing; flowers pink St.John, H. 22791 Niihau BISH 1947-08-14
160 Boerhavia repens Kalawao, Kalaupapa Airport edge of landing field Preserved_Specimen Wagner, W.L. 4940 Molokai BISH 1983-08-02
161 Boerhavia repens Tern Islet near shore with Ipomoea pes-caprae Preserved_Specimen Carlquist, S. 2271 French Frigate Shoals BISH 1966-09-12
162 Boerhavia repens S side of runway sand Preserved_Specimen Long, C.R. 2228 Kure Atoll BISH 1964-09-14
163 Boerhavia repens Mokulua, north islet near the beach, on volcanic soil. Preserved_Specimen Locally common. Egler, F.E. 37255 Oahu BISH 1937-08-22
164 Boerhavia repens Manana island, off Waimanalo, Oahu Near the beach. W.(?) side. Preserved_Specimen Strandtmann, R.W. Oahu BISH 1979-06-12
165 Boerhavia repens Mokolii A few large patches. Preserved_Specimen Starr, F. 5041930 Oahu BISH 2005-04-19
166 Boerhavia repens NW end of lagoon sand; with Eragrostis and Ipomoea Preserved_Specimen prostrate herb 0.1-0.2 m high with pinkish-white flowers Young, A.L. 147 Laysan Island BISH 1964-09-17
167 Boerhavia repens Wailuku rocky shore Preserved_Specimen Faurie, A. 557 Maui BISH 1909-08-01
168 Boerhavia repens Kapapa Islet, Kaneohe Bay Raised coral reef with shallow, sandy soil. Preserved_Specimen Herbst, D.R. 6129 Oahu BISH 1978-05-24
169 Boerhavia repens Kekepa Islet Islet is a raised limestone reef. Plants growing in soil pockets or in thin soil layer over limestone base. Preserved_Specimen Herbst, D.R. 5860 Oahu BISH 1976-06-06
170 Boerhavia repens Laie sand flat under trees Preserved_Specimen fls. pinkish, plant prostrate. Fosberg, F.R. 14186 Oahu BISH 1937-07-04
171 Boerhavia repens East of Waialua Beach Park Collected on sand. Other common strand species also present. Preserved_Specimen Stemmermann, R.L. 3700 Oahu BISH 1979-02-11
172 Boerhavia repens Trig Island, E end sand; with Lepturus, Tribulus and Portulaca Preserved_Specimen flowers lavender Long, C.R. 2471 French Frigate Shoals BISH 1964-09-27
173 Boerhavia repens National Park, Kau, near Halape shelter by beach, open, sunny Preserved_Specimen trailing Herat, T. 864 Hawaii BISH 1974-06-30
174 Boerhavia repens Kau, Mahana Paddock open pasture, close to ocean Preserved_Specimen sprawling; flowers white Kawasaki, A. 28 Hawaii BISH 1964-01-28
175 Boerhavia repens Mahana, Halulu, edge of beach sand along edge of beach, dry Preserved_Specimen prostrate, pale pink flowers Herbst, D.R. 5778 Lanai BISH 1976-03-30
176 Boerhavia repens Lisianski Island bunchgrass Preserved_Specimen Herbst, D.R. 6494 Lisianski Island BISH 1980-07-11
177 Boerhavia repens Waihee, Maui Coastal Land Trust property, near western stream boundary Beach dune, with Sida fallax, Jacquemontia ovalifolia var. sandwicensis, Melanthera sp. Preserved_Specimen Decumbent vine; white flowers. James, S.A. 326 Maui BISH 2004-03-04
178 Boerhavia repens Poipu Beach top of coral sand Preserved_Specimen flowers pinkish, in a single head with 2-4 stamens St.John, H. 23098 Kauai BISH 1947-12-27
179 Boerhavia repens W shore sandy Preserved_Specimen prostrate vine, large mats, 12 cm high; flower lavender Daniel, C.W. 3 Laysan Island BISH 1959-07-24
180 Boerhavia repens Barking Sands, N end, Waimea open sand dunes Preserved_Specimen trailing; flowers white or pinkish; stamens 3 St.John, H. 22965 Kauai BISH 1947-12-23
181 Boerhavia repens Kaluakoi, Kawakiu Road Sand dune, with wind. Preserved_Specimen Prostrate, stems 6-8 ft. long. St.John, H. 12451 Molokai BISH 1932-12-22
182 Boerhavia repens Mokulua, north islet near the beach, on volcanic soil. Preserved_Specimen Locally common. Egler, F.E. 37255 Oahu BISH 1937-08-22
183 Boerhavia repens Waihu Beach sand, with Sesuvium portulacastrum Preserved_Specimen trailing vine, 1/2 m across cluster; flower white Woolford, H.A. 194 Maui BISH 1956-01-01
184 Boerhavia repens Barking Sands sand dunes Preserved_Specimen stems prostrate, 1-2 m long, flowers white Fosberg, F.R. 12739 Kauai BISH 1935-12-30
185 Boerhavia repens U.H. Campus, University High School In a grass lawn in full sun. Preserved_Specimen A prostrate subshrub forming a mat 1.5 m across. The flowers are purple. Lau, J. 1383 Oahu BISH 1985-09-02
186 Boerhavia repens French Frigate Shoals sand Preserved_Specimen flower red Caum, E.L. 86 French Frigate Shoals BISH 1923-06-22
187 Boerhavia repens Laysan sandy, near Casuarina equisetifolia Preserved_Specimen prostrate Tsuda, R.T. 490 Laysan Island BISH 1963-12-04
188 Boerhavia repens Nualolo-Kai, Waimea District shore flats Preserved_Specimen trailing, flowers white, stamens 2 St.John, H. 26691 Kauai BISH 1965-04-24
189 Boerhavia repens Koloa District. Mahaulepu, south of quarry and horse stables. Scaevola taccada - Fimbristylis cymosa coastal shrubland on raised limestone w/ Sida fallax, Euphorbia degeneri, Capparis sandwichiana, Jacquemontia ovalifolia subsp. sandwicensis, Lycium sandwichense, Boerhavia repens, Osteomeles anthyllidifolia, Sesuvium portulacastrum Preserved_Specimen Prostrate herb, creeping w/ long stems yellow-green, in flower, corolla white, common Wood, K.R. 18680 Kauai BISH 2021-05-06
190 Boerhavia repens Alau In sheer open areas. Preserved_Specimen Starr, F. 5040512 Maui BISH 2005-04-05
191 Boerhavia repens Kalani, Moomomi, Kaluakoi on sand dunes. Preserved_Specimen flowers pink, stamens 2-3 St.John, H. 25154 Molokai BISH 1953-12-21
192 Boerhavia repens Lihue District. Coastal strand around Nukolii Beach Park, Weast Kauai Naupaka Kahakai (Scaevola) Shrubland Sporobolus virginicus, Scaevola taccada, Tournefortia argentea, Schinus teribinthifolius, Cynodon dactylon, Ipomoea pes-caprae, I. imperati, Euphorbia serpens, Vigna luteola, Portulaca lutea, Sphagnet. Growing in low dunes intertwined with Sporobolus virginicus, Euphorbia serpens, and Canavalia sericea, just seaward of Scaevola shrubland and above the high water mark Preserved_Specimen Mat-forming perennial, a meter or so across. Flowers light pink. Leaves thick, darker adaxially. Williams, A.M. 744 Kauai BISH 2022-09-23
193 Boerhavia repens North Island, central, slightly depressed on W end Solanum, Eragrostis Preserved_Specimen Long, C.R. 2307 Pearl & Hermes Atoll BISH 1964-09-17
194 Boerhavia repens S end, S crest sandy soil Preserved_Specimen 15 cm prostrate vine; flowers white Laysaon, C.W. 19 Laysan Island BISH 1959-07-25
195 Boerhavia repens Manana Island sandy soil E end of island, W side numerous creeping stems. Preserved_Specimen Lavender flrs. Long, C.R. 1688 Oahu BISH 1964-05-02
196 Boerhavia repens NW Kahalui sand flat Preserved_Specimen prostrate Fosberg, F.R. 9827 Maui BISH 1933-08-17
197 Boerhavia repens Mokulua South, Oahu sheer areas near the coast and along ridges Found in open sheer areas, near the coast, and along ridges mixed in with other plants. Preserved_Specimen Occasional to common. Starr, F. 5022330 Oahu BISH 2005-02-23
198 Boerhavia repens North Island Solanum, Boerhavia Preserved_Specimen Herbst, D.R. 6472 Pearl & Hermes Atoll BISH 1980-07-06
199 Boerhavia repens Southeast Island, W side With Boerhavia. Preserved_Specimen Prostrate herb. Conant, S. 146 Pearl & Hermes Atoll BISH 1983-07-21
200 Boerhavia repens Kaluakoi, Kawakiu Rd Sand dune, with wind. Preserved_Specimen Prostrate, stems 6-8 ft long. St.John, H. 12451 Molokai BISH 1932-12-22
201 Boerhavia repens Whale-Skate Island sand Preserved_Specimen Long, C.R. 2477 French Frigate Shoals BISH 1964-09-27
202 Boerhavia repens Paopao Point beach Preserved_Specimen Degener, O. 24249 Lanai BISH 1957-08-20
203 Boerhavia repens S end, near lagoon sandy Preserved_Specimen prostrate vine; flower lavender Daniel, C.W. 15 Laysan Island BISH 1959-07-25
204 Boerhavia repens Kahuku - Between coast and wildlife re Slope: 1%. Soil: Sandy. Vegetation type:Tropical coastal plants. Preserved_Specimen Low-lying perennial herb. Distribution: Species found on islands of Pacific. Diffusa is endemic to Hawaiian Is. Smith, M. 33 Oahu BISH 1986-04-08
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