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Dioscorea pentaphylla

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Dioscoreales Family: Dioscoreaceae Genus: Dioscorea

fiveleaf yam, pi‘a [pia] Hawai‘i

Description

Key Characters:

Growth Form: Vines.

Stems: Stems twining to the left, = angled, up to 10 m long, pilose to hirsute, glabrate with age, usually with numerous scattered prickles, tubers nearly always solitary, variable in shape, growing near the surface or over 1 m underground.

Roots: Fibrous root system.

Leaves: Leaves compound (palmate) or rarely simple. Alternate. Leaflets 3–5, elliptic or oblong, 5–10 cm long, 2–4 cm wide, the central one oblanceolate or obovate, the lateral ones somewhat asymmetrical. Apex obtuse or acute to acuminate. Surfaces with yellowish brown pubescent at least on lower surface, axils with bulbils and occasionally bearing globose to horseshoe shaped tubers ca. 1 cm in diameter. Leaflet margins entire. Petioles 5-11 cm long, densely reddish brown pubescent. Stipules absent.

Flowers: Staminate flowers in axillary, spike-like racemes up to 30 cm long, near the ends of the branches, reddish pubescent, pistillate flowers in long pendent spikes 5–25 cm long. Flowers unisexual, the plants dioecious. Calyx of staminate flowers with Tepals broadly lanceolate, ca. 1 mm long. Calyx of pistillate flowers with tepals ca. 1.5 mm long. Staminate flower with 3 stamen and 3 staminodes (longer than stamens); pistillate flowers with 6 rudimentary staminodes. Ovary inferior, villous, 3-carpellate.

Fruit: Capsules oblong; 20–25 mm long; glabrous or glabrate; shallowly emarginate at both ends; the wings up to 6 mm wide. Seeds winged on basal side only.

Ploidy: 2n = 40; 70; 80; 144

Habitat: Naturalized primarily in windward mesic valleys.

Elevation Range: 115–325 m.

Historical Distribution

Uses and Culture

USES

  • Tubers eaten in times of famine (Krauss 1993:11,12).

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Specimens

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Bibliography

Name Published In: Sp. Pl.: 1032 (1753)

Other References

Wagner et al. 1990:1437 (K, O, Mo, M, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
2 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
3 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
4 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
5 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
6 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
7 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
8 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
9 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
10 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
11 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
12 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
13 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
14 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
15 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
16 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
17 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
18 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
19 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
20 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
21 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
22 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
23 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
24 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
25 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
26 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
27 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
28 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
29 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
30 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
31 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
32 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
33 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
34 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
35 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
36 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
37 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
38 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
39 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
40 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
41 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
42 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
43 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
44 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
45 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
46 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
47 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
48 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
49 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
50 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
51 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
52 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
53 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
54 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
55 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
56 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
57 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
58 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
59 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
60 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
61 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
62 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
63 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
64 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
65 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
66 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
67 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
68 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
69 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
70 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
71 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
72 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
73 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
74 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
75 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
76 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
77 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
78 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
79 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
80 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
81 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
82 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
83 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
84 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
85 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
86 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
87 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
88 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
89 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
90 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
91 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
92 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
93 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
94 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
95 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
96 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
97 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
98 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
99 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
100 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
101 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
102 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
103 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
104 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
105 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
106 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
107 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
108 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
109 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
110 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
111 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
112 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
113 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
114 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
115 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
116 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
117 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
118 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
119 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
120 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
121 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
122 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
123 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
124 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
125 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
126 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
127 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
128 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
129 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
130 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
131 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
132 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
133 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
134 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
135 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
136 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
137 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
138 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
139 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
140 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
141 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
142 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
143 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
144 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
145 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
146 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
147 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
148 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
149 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
150 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
151 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
152 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
153 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
154 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
155 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
156 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
157 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
158 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
159 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
160 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
161 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
162 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
163 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
164 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
165 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
166 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
167 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
168 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
169 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
170 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
171 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
172 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
173 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
174 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
175 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
176 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
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