Description
Key Characters:
Growth Form:
Erect or strongly ascending, branched shrubs 3–8(–18) dm tall, glabrous throughout except for the leaf margins.
Stems:
Stems often swollen at the nodes, terete (cylindrical), 4–angled or –ridged, lateral branches of primary stems usually elongating several or more nodes below the inflorescence.
Roots:
Leaves:
Leaves simple.
Opposite.
Blades elliptic–ovate or sometimes oblanceolate, 6–12.5 cm long, 2.3–6 (–6.8) cm wide.
Apex weakly mucronate.
Base leaf bases often connate around the stem.
Surfaces green; blades thick, chartaceous at least when dry.
Margins entire, serrulate, the teeth with antrorsely hooked hairs ca. 0.1–0.15 mm long.
3 principal veins, sometimes also with an inconspicuous looping pair of veins near the margins.
Petioles 1.5–3 (–3.8) cm long, pale green, weakly to conspicuously connate around the stem.
Stipules absent.
Flowers:
Flowers in inflorescence pseudoaxillary, with (2–) 18–34 flowers; bracts much smaller than the uppermost leaves, recurved, the lowest pair to 1.4 cm long; peduncles 16–30 mm long, elongating to 50 mm long in fruit, secondary and tertiary intemodes of the lateral inflorescence branches 20–30 mm long; pedicels thinner, 10–20 mm long, elongating uniformly during inflorescence development.
Flowers bisexual (perfect), cleistogamous, pendent.
Calyx not or only slightly opening at anthesis; sepals 4 (5), 6–8 mm long, enlarging to 9–10 mm long and the inner ones to 10 mm wide and the outer ones to 14 mm wide in fruit, greenish white within, outer ones green externally, inner ones white, green toward midrib, elliptic to broadly elliptic, apex obtuse and retuse, or truncate, becoming dark purple and fleshy as fruit matures.
Corolla (petals) absent.
Stamens 10 (–11); filaments ca. 4.5 mm long, subequal; anthers 2.2–2.5 mm long, pale reddish purple at anthesis, changing to a darker reddish purple.
Pollen gray.
Ovary superior, -celled; ovule placentation free-central; styles 4–6 (–7), sometimes variable in number on the same plant, filiform; stigma receptive completely around the distal half or rarely nearly throughout its length, slightly enlarged and papillose.
Fruit:
Capsules 8–12 mm long; ovoid to subglobose; enclosed by or exserted from the persistent dried calyx (in section Alsinidendron calyx fleshy and dark purple); dehiscing at maturity by 4–11 valves.
Seeds relatively few to numerous; 1.4–1.8 mm long; the viable ones black or dark or light brown; the inviable ones usually reddish brown; orbicular–reniform; reniform; or rarely orbicular; usually laterally compressed; the surface rugose; papillose; or sometimes nearly smooth.
Ploidy:
2n = 50–54; 2n = 60
Habitat:
Rare in wet montane forest.
Elevation Range:
(600–) 900–1230 m.