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Ceodes umbellifera

J.R.Forst. & G.Forst. (1776)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-CoreEudicot Order: Caryophyllales Family: Nyctaginaceae Genus: Ceodes

pāpala [papala], pāpala kēpau [papala kepau]

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Description

Key Characters:

Growth Form: Large glabrous shrubs or trees up to 10 m tall, loosely branched.

Stems: Branchlets thick, terminal "nodes" enlarged (perhaps best called aggregate nodes), probably really a number of nodes and internodes condensed and bearing pseudo–whorls of leaves and pseudo–terminal cymes, as well as new branchlets, the "terminal" or next proximal internode elongate, 3–15 cm long, and usually straight and weakly quadrangular in cross section, 3–10 mm in diameter.

Roots:

Leaves: Leaves simple. Leaves nearly always in pseudo–whorls of 3–8 or opposite. Blades extremely variable in size and shape, ovate–elliptic to broadly elliptic, obovate, or sometimes broadly spatulate, 5–35 cm long, 5–15 cm wide. Apex subacute or obtuse, rarely somewhat acuminate. Base usually more or less cuneately narrowed or contracted to a short petiole. Blades thick and chartaceous to subcoriaceous. Margins entire to sinuate. Primary lateral veins 8–10(–12) pairs, not very prominent, on most specimens forking and anastomosing toward margins, tertiary and higher order venation obscure. Petioles 0.5–2(–3) cm long. Stipules absent.

Flowers: Flowers in pseudo–terminal, glabrous to minutely puberulent cymes, these 1–5(6) at an aggregate node, occasionally later becoming apparently axillary by vigorous growth of a new branchlet, each one 6–15 cm long at anthesis, enlarging up to twice as long and becoming diffuse in fruit, usually long pedunculate and branching once or twice or more, with aggregate nodes forming an umbellate pattern, sometimes irregularly so, hemispherical to globose in outline, rays 6–10, ultimate umbellules at times somewhat irregular, scale-like bractlets at articulations caducous or absent, often 1 bractlet present part way up the pedicel. Flowers unisexual, rarely appearing bisexual (perfect), strongly fragrant. Calyx of staminate flowers campanulate at anthesis, 4–5-lobed, limb rounded distally, lobes ovate, spreading to somewhat recurved, margins thick, usually revolute. Calyx of pistillate flowers somewhat smaller than staminate flowers. Corolla (petals) absent. Stamens in staminate flowers 6–10, slightly to well exserted; anthers orbicular; pistillate flower staminodes usually not exserted. Ovary superior (pistillate flowers); stigma slightly to strongly exserted, capitate, apically penicillate or covered by hair-like, simple to slightly branched processes, giving a kahili-like appearance. Staminate flowers with a pistillode included or very slightly exserted, capitate to somewhat lobed, processes scarcely evident, apex recurved.

Fruit: Anthocarps prismatic; 3–7 cm long; becoming hard; angles very glutinous; the distal half to third tapering to a sterile portion; forming a rostrum; lobes persistent; spreading and stellate; margins usually tightly revolute or merely thickened; with a triangular portion usually present arising within the margin of the lobe and extending slightly beyond it. Seeds 1 per fruit; closely enclosed by the thin ovary wall.

Ploidy: 2n = 112

Habitat: Wet to moist forests; usually in valleys.

Elevation Range: 90–800 m.

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Pollinators

Bibliography

Name Published In: Char. Gen. Pl.: 142 (1776)

Other References

Wagner et al. 1990:987 (K, O, Mo, L, M, H [as Pisonia umbellifera]); Rossetto et al. 2020:132 (Syn. P. umbellifera = C. umbellifera); Munro 245 BISH (L†, 1913)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Ceodes umbellifera Hanalei District. Wainiha Valley; Halelea Forest Reserve - along road from Powerhouse to dam Secondary forest Preserved_Specimen Tree of 15 ft. in Hibiscus tiliaceus thicket. Leaves mat green. Flowers in bud pink, opening pale whitish-pink. Fragrance semen like. Common along road. Flynn, T.W. 3485 Kauai BISH 1989-07-16
2 Ceodes umbellifera Keanae Valley, Koolau Forest Reserve Small pocket of native plants; 3-4 mi. into valley along jeep road. Preserved_Specimen Sohmer, S.H. 6579 Maui BISH 1970-08-18
3 Ceodes umbellifera Tantalus, near telephone relay station dense bamboo Preserved_Specimen Takeuchi, W.N. 1881 Oahu BISH 1984-12-08
4 Ceodes umbellifera Keanae Valley, Koolau Forest Reserve 3-4 mi. into valley along jeep road Preserved_Specimen Wagner, W.L. 5839 Maui BISH 1988-03-06
5 Ceodes umbellifera Headwaters of Hanalei Rain forest along stream Preserved_Specimen Tree. Ht. 10 m. MacDaniels, L.H. 834 Kauai BISH 1927-02-21
6 Ceodes umbellifera Koolau Forest Reserve, Keanae Valley, 3-4 mi. into valley along jeep road Preserved_Specimen Sohmer, S.H. 6584 Maui BISH 1970-08-18
7 Ceodes umbellifera Halepuaa Section, Nanawale Forest Reserve Metrosideros - Pandanus forest with occasional Diospyros and Aleurites. Preserved_Specimen Tree 20' tall, 22' spread, 10 basal diam. Clarke, G. 698 Hawaii BISH 1982-04-22
8 Ceodes umbellifera Halepuaa Section, Nanawale Forest Reserve Metrosideros - Pandanus forest, with occasional Diospyros and Aleurites. Preserved_Specimen Tree 20 ft tall, 22 ft spread, 10 inch basal diam. Clarke, G. 698 Hawaii BISH 1982-04-22
9 Ceodes umbellifera Laupahoehoe Section of Hilo Forest Reserve, North Hilo District on floor of open canopy Acacia koa-Metrosideros forest with native tree understory. Preserved_Specimen Tree seedling 1 1/2' tall, leaf petiole and mid-rib magenta in color. Cuddihy, L.W. 758 Hawaii BISH 1981-03-03
10 Ceodes umbellifera Waimea Arboretum full sun Preserved_Specimen 5 m tree, multi-trunked; flower white and rose pink, strong odor, morning Lau, J. 2760 Oahu BISH 1986-08-18
11 Ceodes umbellifera west of Wailau Valley on Waiehu land shelf Vegetation: Charpentiera, Syzygium, Morinda, Pandanus, Cordyline, Asplenium. Preserved_Specimen Locally common tree to 7 m. Oppenheimer, H.L. 80508 Molokai BISH 2005-08-17
12 Ceodes umbellifera About 150' above Paaiki Valley Kokia, Gouldia, ohia, Drypetes, lantana, Dodonaea association. North facing slope of valley. Preserved_Specimen Corn, C. Kauai BISH 1977-07-19
13 Ceodes umbellifera Between Keahiakahoe and Kahuauli, gully at cliff base Cyrtandra adpressipilosa Preserved_Specimen Takeuchi, W.N. 1811 Oahu BISH 1984-11-03
14 Ceodes umbellifera Paupauwela. Central Waianae mountains. Mostly disturbed mesic forest dominated by Psidium cattleianum, Toona ciliata, Fraxinus, and Schinus terebinthifolius. Also growing with remnant natives such as Sapindus oahuensis, Pittosporum flocculosum, Microlepia setosa, Planchonella sandwicensis, etc. Preserved_Specimen Large tree with spreading canopy, over 5 meters tall. Leaves whorled on nodes, thick, coriaceous, flowers in small clusters at branch tips. Fruit narrowly ovoid and elongated, very viscous. Thomas, M.K. 616 Oahu BISH 2023-12-30
15 Ceodes umbellifera Kawela, Kamakou Preserve, gully below Puu Kolekole growing in gully with Dodonaea, Metrosideros, Osmanthus, Lysimachia, and Dissochondrus Preserved_Specimen Tree 8 ft. tall Anderson, S.J. 522 Molokai BISH 1982-05-24
16 Ceodes umbellifera S ridge, Kipapa gulch, Waipio, Koolau Mts moist Preserved_Specimen 30' tree Hosaka, E.Y. 977 Oahu BISH 1933-04-30
17 Ceodes umbellifera Wailupe Stream, E branch under kukui Preserved_Specimen Takeuchi, W.N. 1956 Oahu BISH 1985-01-27
18 Ceodes umbellifera Waiahole, base of Koolau divide exotic riparian Preserved_Specimen large tree Takeuchi, W.N. 2521 Oahu BISH 1985-10-26
19 Ceodes umbellifera Kaluaa, streambed Pipturus, Streblus Preserved_Specimen 4 m tree Takeuchi, W.N. 2383 Oahu BISH 1985-08-10
20 Ceodes umbellifera Kalaupapa National Historical Park Waikolu. Upper waikolu valley floor, above intake tunnel; Waikolu transect 2 w:Me-Df,nt(mf) Preserved_Specimen large tree~10m in height ; rare, only one tree seen Wysong, M.L. 734 Molokai BISH 2005-03-29
21 Ceodes umbellifera ridge S of Kipapa Gulch, Koolau Mts woods Preserved_Specimen 20' x 1 tree; leaves fleshy St.John, H. 10020 Oahu BISH 1929-11-10
22 Ceodes umbellifera Puu Hapapa, Waianae Mts woods Preserved_Specimen Hosaka, E.Y. 181 Oahu BISH 1930-03-16
23 Ceodes umbellifera West Maui, Wailuku, S side of Iao Stream 3-4 mi. into valley along jeep road Preserved_Specimen Wagner, W.L. 5839 Maui BISH 1988-03-06
24 Ceodes umbellifera Haiku Valley, Waiahole FR, Heeia Gleichenia Preserved_Specimen 20' tree St.John, H. 12262 Oahu BISH 1932-12-11
25 Ceodes umbellifera Makaha Valley, Waianae Mts moist forest Preserved_Specimen 15 m x 8 dm tree; flower pink St.John, H. 24856 Oahu BISH 1952-11-30
26 Ceodes umbellifera Hanakapiai Falls Trail, Na Pali Coast, Growing in dense shade which is a frequent occurrence near the stream in the upper regions of the valley. Preserved_Specimen A 25' tree Clarke, G. 336 Kauai BISH 1980-04-11
27 Ceodes umbellifera 15 off dirt road, 1.1m. n. of Hanahanapuni cone Found associated with Toona, Aleurites, Hibiscus. Understory of Nephrolepis, Dicranopteris. Preserved_Specimen Tree 30' - 35', dbh=12, Ishikawa, S. 290 Kauai BISH 1973-08-21
28 Ceodes umbellifera Puu Hapapa, Waianae Mts woods Preserved_Specimen Hosaka, E.Y. 181 Oahu BISH 1930-03-16
29 Ceodes umbellifera Koolau Forest Reserve, Keanae Valley Pocket of Native Plants; 3-4 mi. into valley along jeep road Preserved_Specimen Sohmer, S.H. 6583 Maui BISH 1970-08-18
30 Ceodes umbellifera Koolau Forest Reserve, Keanae Valley, 3-4 mi. into valley along jeep road Preserved_Specimen Sohmer, S.H. 6579 Maui BISH 1970-08-18
31 Ceodes umbellifera Hanalei District. NaPali Coast State Park, Hoolulu Valley Coastal Mesic Forest W/ Charpentiera densiflora, Pandanus, Aleurites, Pittosporum napaliense, Diospyros, Oplismenus hirtellus, Elephantopus mollis, Pritchardia napaliensis, Alyxia, Bobea, Cordyline, Psydrax, Psychotria m Preserved_Specimen Tree 9 ft. tall next to trail. With sticky fruit hanging on long branching diffuse peduncles. Leaves are dark green and smaller than other Pisonia in area. Tangalin, N. 3300 Kauai BISH 2012-09-05
32 Ceodes umbellifera Makawao Stream, E fork, below Puu Lanipo, Kailua, Koolau Mts moist, Aleurites Preserved_Specimen flower red without, white within, heavy sweet fragrance St.John, H. 21551 Oahu BISH 1946-03-20
33 Ceodes umbellifera Kahuna Stream above intake, moist cliffs Preserved_Specimen 5 m x 1 dm tree St.John, H. 17671 Maui BISH 1936-12-19
34 Ceodes umbellifera Kahana, Waikane-Schofield trail, Koolau Mts forest Preserved_Specimen Hartung, M. Oahu BISH 1935-04-28
35 Ceodes umbellifera Wainiha Moist lower forest Preserved_Specimen 10 ft. shrub St.John, H. 13923 Kauai BISH 1934-01-01
36 Ceodes umbellifera Mt. Kalena, Waianae Aleurites forest Preserved_Specimen 35' tree Grant, M.L. 7371 Oahu BISH 1934-09-30
37 Ceodes umbellifera Pig God Trail, Punaluu dark forest Preserved_Specimen Degener, O. 9553 Oahu BISH 1931-05-30
38 Ceodes umbellifera West Maui: Wailena Gulch Growing in moist woods. Preserved_Specimen Tree, 75-100 feet tall, 14-18 in. in diameter. Flowers reddish orange; fruits exceedingly sticky. Crosby, M.R. 1751 Maui BISH 1964-07-15
39 Ceodes umbellifera Waimea Arboretum full sun Preserved_Specimen 5 m tree, multi-trunked; flower white and rose pink, strong odor, morning Lau, J. 2673 Oahu BISH 1986-07-28
40 Ceodes umbellifera Located on the Mauna Loa Strip Road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 ° north of road. (plot 19) Acacia - Sapindus savanna community Preserved_Specimen Tree Newell, C.L. 153 Hawaii BISH 1966-07-26
41 Ceodes umbellifera Lualualei, below Kanehoa, valley 9 Open valley bottom Preserved_Specimen 5 m tree; flowers; female Christophersen, E. 3711 Oahu BISH 1932-06-29
42 Ceodes umbellifera Kuliouou Valley Aleurites, Psidium cattleianum Preserved_Specimen St.John, H. 26906 Oahu BISH 1978-01-29
43 Ceodes umbellifera Napali coast; Hanakapiai, Maunapuluo On cliff. Preserved_Specimen St.John, H. 10891 Kauai BISH 1930-12-30
44 Ceodes umbellifera Makaha Valley, Waianae Mts moist forest Preserved_Specimen 15 m x 8 dm tree; flower pink St.John, H. 24856 Oahu BISH 1952-11-30
45 Ceodes umbellifera Kahuna Stream above intake, moist cliffs Preserved_Specimen 5 m x 1 dm tree St.John, H. 17671 Maui BISH 1936-12-19
46 Ceodes umbellifera Koloa District, Lawai Valley. Cultivated the National Tropical Botanical Garden Nat'l Tropical Botanical Garden Preserved_Specimen Densely branched tree of 10 ft.; leaves medium green above with pale green midvein, paler below; corolla whitish-green, flowers fragrant (l.30p) From seed wild collected Mt. Haupu. Flynn, T.W. 5538 Kauai BISH 1994-03-28
47 Ceodes umbellifera ahupuaa of Halepuaa in the Puna District of Hawaii Growing in Guava scrub in heavy shade. Preserved_Specimen Tree is approximately 30 ft. tall. Kubo, R. 28 Hawaii BISH 1979-05-04
48 Ceodes umbellifera Na Pali Kona Coast Trail between Haena Point and Hanakoa Valley Growing in protected valley Preserved_Specimen Tree, 12 in. dia. Stern, W.L. 1259 Kauai BISH 1961-08-01
49 Ceodes umbellifera Lualualei, head, below Puu Kanehoa open forest Preserved_Specimen 5-6 m tree; flower flesh colored; male Christophersen, E. 3662 Oahu BISH 1932-05-25
50 Ceodes umbellifera Kipapa Gulch, Waipio, Koolau Mts along stream bed Preserved_Specimen 30' tree Hosaka, E.Y. 1051 Oahu BISH 1933-05-28
51 Ceodes umbellifera Kipapa Gulch, Koolau Mts moist gully Preserved_Specimen 35' tree Hosaka, E.Y. 1055 Oahu BISH 1933-06-04
52 Ceodes umbellifera ridge N of Kuapa Pond edge of forest Preserved_Specimen Gillespie, J.W. 1095 Oahu BISH 1927-08-08
53 Ceodes umbellifera Dive to Kipuka Puaulu located about 1.5 miles up the Mauna Loa Strip Road from intersection with Kona-Hilo Highway. Plant was about halfway around Kipuka Puaulu Nature Trail on side trail leading to the Big Koa tree. (plot 62) Acacia - Sapindus - Microlepia community Preserved_Specimen Tree Newell, C.L. 1034 Hawaii BISH 1967-06-28
54 Ceodes umbellifera Kaluaa Gulch, above Schofield, between Kanehoa and Hapapa deep shade Preserved_Specimen flower dull rose red Hutchison, P.C. 2834 Oahu BISH 1967-01-15
55 Ceodes umbellifera Waikane, Schofield trail woods Preserved_Specimen 15' tree St.John, H. 10167 Oahu BISH 1930-01-19
56 Ceodes umbellifera Kauai Preserved_Specimen Faurie, A. 713 Kauai BISH 1910-01-01
57 Ceodes umbellifera Napali coast; Hanakapiai, Maunapuluo On cliff. Preserved_Specimen St.John, H. 10891 Kauai BISH 1930-12-30
58 Ceodes umbellifera East branch Kilauea river Preserved_Specimen Tree. Ht. 15 m. MacDaniels, L.H. 643 Kauai BISH 1927-02-11
59 Ceodes umbellifera Waianae Mts., Puu Kanehoa Ridge Preserved_Specimen tree; fruit, flower pinkish brown Stone, B.C. 3621 Oahu BISH 1961-08-27
60 Ceodes umbellifera Koolau Mountains; Waipio; in moderately moist gully of Kipapa Gulch Preserved_Specimen Tree 35' tall. Hosaka, E.Y. 1055 Oahu BISH 1933-06-04
61 Ceodes umbellifera Lualualei Valley, Palikea Preserved_Specimen to 20' tree, hollow trunk Takeuchi, W.N. 8511 Oahu BISH 1992-09-01
62 Ceodes umbellifera Lualualei divide, Puu Hapapa, dry streambed Preserved_Specimen large tree; sterile Takeuchi, W.N. 2451 Oahu BISH 1985-09-14
63 Ceodes umbellifera Waihee, at beginning of trail Preserved_Specimen 5-10 m tree DLNR staff 25 Maui BISH
64 Ceodes umbellifera Inside forest reserve boundary, Kaluanui Valley, near the falls (Sacred Falls) Underneath kukui and mountain apple trees. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
65 Ceodes umbellifera Koolau Mountains; Waipio; in gully of Kipapa Gulch Preserved_Specimen Tree 30' tall. Hosaka, E.Y. 1056 Oahu BISH 1933-06-04
66 Ceodes umbellifera Palolo Valley Preserved_Specimen Swezey, O.H. 12760 Oahu BISH
67 Ceodes umbellifera Tantalus Preserved_Specimen Brown, F.B.H. 139 Oahu BISH 1920-06-05
68 Ceodes umbellifera Waianae Mts., gulch below Palikea Preserved_Specimen Leaves thick, coriaceous, shiny on both sides; flower pinkish white Cowan, R.S. 631 Oahu BISH 1947-06-15
69 Ceodes umbellifera Mahana Preserved_Specimen female Munro, G.C. 245 Lanai BISH 1913-11-22
70 Ceodes umbellifera Pauoa Valley Preserved_Specimen Rock, J.F.C. 1091 Oahu BISH 1909-01-07
71 Ceodes umbellifera Lualualei Valley, Palikea Preserved_Specimen to 20' tree, hollow trunk Takeuchi, W.N. 8511 Oahu BISH 1992-09-01
72 Ceodes umbellifera Ka Loko Reservoir, Kilauea Preserved_Specimen Forbes, C.N. 526 Kauai BISH 1916-10-17
73 Ceodes umbellifera Wahiawa Mts Preserved_Specimen Forbes, C.N. 180 Kauai BISH 1909-08-01
74 Ceodes umbellifera East Maui, Hana District, ~15 mi from Hana, off hwy Preserved_Specimen leaves obovate, cuneate at base; short fleshy petiole; upper leaves whorled, lower opposite; perianth uniseriate, limb of perigone 5 lobed; inflorescences axillary, compactly paniculate Ishikawa, S. 199 Maui BISH 1972-12-19
75 Ceodes umbellifera Manoa Falls trail, near falls Preserved_Specimen tree Kerr, M. 23 Oahu BISH 1945-12-15
76 Ceodes umbellifera Wailau Valley Preserved_Specimen Male. Tree 20 feet. St.John, H. 13245 Molokai BISH 1933-07-03
77 Ceodes umbellifera N Tanatalus Preserved_Specimen 4 m tree MacDaniels, L.H. 71 Oahu BISH 1926-10-09
78 Ceodes umbellifera Lualualei divide, Puu Hapapa Preserved_Specimen in flower; perigone cream colored Takeuchi, W.N. 2450 Oahu BISH 1985-09-14
79 Ceodes umbellifera Kului Gulch west of Hawaiiloa Ridge Preserved_Specimen Fruits sticky, leaves in whorls. Kiehn, M. 94071812 Oahu BISH 1994-01-01
80 Ceodes umbellifera Kahauiki Gulch, Kalihi, behind Fort Shafter, Koolau Mts rain forest Preserved_Specimen Yoshioka, T. Oahu BISH 1933-12-25
81 Ceodes umbellifera Waikane, Koolau Mts Preserved_Specimen Morley, H. Oahu BISH 1934-05-13
82 Ceodes umbellifera Haena-Kalalau Trail, about 3 1/2 miles from the Haena end Preserved_Specimen Herbst, D.R. 2861 Kauai BISH 1973-01-19
83 Ceodes umbellifera Tantalus Preserved_Specimen Brown, F.B.H. 139 Oahu BISH 1920-06-05
84 Ceodes umbellifera Pauoa Valley Preserved_Specimen Rock, J.F.C. 583 Oahu BISH 1908-11-01
85 Ceodes umbellifera Olaa forest Preserved_Specimen tree 20 ft. high, branches angular, fls. rich pink Stephens, E.W. Hawaii BISH 1957-12-01
86 Ceodes umbellifera Oahu Preserved_Specimen Brown, F.B.H. 117 Oahu BISH
87 Ceodes umbellifera Lualualei Valley, Palikea Preserved_Specimen to 20' tree, hollow trunk Takeuchi, W.N. 8511 Oahu BISH 1992-09-01
88 Ceodes umbellifera Kaala, Waianaeuka Preserved_Specimen tree; white flower Hume, E.P. 35 Oahu BISH 1930-11-30
89 Ceodes umbellifera near the head of Nualolo Valley, Na Pali Coast Preserved_Specimen Corn, C. 348 Kauai BISH 1979-07-18
90 Ceodes umbellifera [Hawaiian Islands; U.S.A. Preserved_Specimen BISH
91 Ceodes umbellifera Punaluu Valley, trail W. of Castle Trail Preserved_Specimen Tree up to 4 m with very brittle white wood. Lv dark green. Fl pinkish. van Royen, P. 10227 Oahu BISH 1968-05-26
92 Ceodes umbellifera Koolau, Palolo valley Preserved_Specimen Skottsberg, C.J.F. 905 Oahu BISH 1922-10-23
93 Ceodes umbellifera East Maui, Nahiku Preserved_Specimen Forbes, C.N. 254 Maui BISH 1910-07-01
94 Ceodes umbellifera Lanai Preserved_Specimen Munro, G.C. 380 Lanai BISH
95 Ceodes umbellifera Manoa Falls trail, near falls Preserved_Specimen Kerr, M. 3681 Oahu BISH 1945-12-15
96 Ceodes umbellifera Ka Loko Reservoir, Kilauea Preserved_Specimen Forbes, C.N. 526 Kauai BISH 1916-10-17
97 Ceodes umbellifera Tantalus ridge Preserved_Specimen Lyon, H.L. Oahu BISH 1909-09-05
98 Ceodes umbellifera On cliffs, Maunapuluo, Hanakapiai, Napali Coast Preserved_Specimen St.John, H. 10891 Kauai BISH 1930-12-30
99 Ceodes umbellifera Hii Mts Preserved_Specimen Forbes, C.N. 645 Kauai BISH 1916-10-20
100 Ceodes umbellifera Pauoa Valley Preserved_Specimen Rock, J.F.C. 1086 Oahu BISH 1909-01-07
101 Ceodes umbellifera Kolekole Pass, Waianae Mts Preserved_Specimen Forbes, C.N. 2033 Oahu BISH 1912-02-01
102 Ceodes umbellifera Mts back of Honolulu Preserved_Specimen Rock, J.F.C. 17157 Oahu BISH 1912-09-01
103 Ceodes umbellifera Olowalu Valley, lateral ridge, L side Preserved_Specimen Forbes, C.N. 2291 Maui BISH 1920-05-10
104 Ceodes umbellifera East Maui, Makawao Preserved_Specimen Faurie, A. 717 Maui BISH 1909-08-01
105 Ceodes umbellifera Mahana Preserved_Specimen Munro, G.C. 205 Lanai BISH 1913-11-22
106 Ceodes umbellifera Manoa Cliffs Trail Preserved_Specimen Sohmer, S.H. 6659 Oahu BISH 1970-10-03
107 Ceodes umbellifera Ka Loko Reservoir, Kilauea Preserved_Specimen Forbes, C.N. 584 Kauai BISH 1916-10-12
108 Ceodes umbellifera Manoa Cliffs Trail Preserved_Specimen Sohmer, S.H. 6113 Oahu BISH 1969-09-27
109 Ceodes umbellifera Manoa Cliff trail Preserved_Specimen Forbes, C.N. 2494 Oahu BISH 1917-05-01
110 Ceodes umbellifera Kalalau Trail, along first mile or so Preserved_Specimen Wagner, W.L. 5176 Kauai BISH 1983-12-21
111 Ceodes umbellifera R fork, Wailupe Valley Preserved_Specimen Garber, D.W. 167 Oahu BISH 1920-01-12
112 Ceodes umbellifera Manoa Cliffs Trail Preserved_Specimen Sohmer, S.H. 6113 Oahu BISH 1969-09-27
113 Ceodes umbellifera Kipahulu, L ridge Preserved_Specimen Forbes, C.N. 1722 Maui BISH 1919-11-26
114 Ceodes umbellifera Hanalei Valley Preserved_Specimen Forbes, C.N. 129 Kauai BISH 1909-08-05
115 Ceodes umbellifera High on Kamalii Ridge. Just below Makalcha Mt., E. Kauai Preserved_Specimen Hobdy, R.W. 11 Kauai BISH 1968-10-01
116 Ceodes umbellifera Palolo Valley Preserved_Specimen Rock, J.F.C. Oahu BISH 1914-10-01
117 Ceodes umbellifera Waiahuakua Valley, Hanakoa Preserved_Specimen St.John, H. 23151 Kauai BISH 1947-12-31
118 Ceodes umbellifera Hillebrands Glen Preserved_Specimen Forbes, C.N. Oahu BISH 1908-12-29
119 Ceodes umbellifera Waiakea Forest Reserve, above Hilo Preserved_Specimen Rock, J.F.C. Hawaii BISH 1957-03-01
120 Ceodes umbellifera Tantalus ridge Preserved_Specimen Rock, J.F.C. Oahu BISH 1909-09-05
121 Ceodes umbellifera base of Kaala Preserved_Specimen Kerr, M. Oahu BISH 1946-02-23
122 Ceodes umbellifera Foot of Pali Preserved_Specimen Forbes, C.N. Oahu BISH 1908-09-07
123 Ceodes umbellifera Maui Preserved_Specimen Forbes, C.N. Maui BISH 1920-01-01
124 Ceodes umbellifera Waikane-Schofield trail, Koolau Mts., ~100' above pump Preserved_Specimen Morley, H. Oahu BISH 1934-05-13
125 Ceodes umbellifera Nailiilihaili Gulch Preserved_Specimen Rock, J.F.C. Maui BISH 1908-12-27
126 Ceodes umbellifera Waiahole Trail Preserved_Specimen Kerr, M. Oahu BISH 1946-08-18
127 Ceodes umbellifera Kamehameha Schools Preserved_Specimen Pearsall, G.A. Oahu BISH 1952-01-18
128 Ceodes umbellifera Kanehoa, Waianae Mts Preserved_Specimen Kondo, Y. Oahu BISH 1941-12-07
129 Ceodes umbellifera Naalehu Preserved_Specimen Russ, G.W. Hawaii BISH 1927-07-01
130 Ceodes umbellifera Waiahole ditch trail Preserved_Specimen Meebold, A. Oahu BISH 1932-06-01
131 Ceodes umbellifera Palolo Valley Preserved_Specimen Rock, J.F.C. Oahu BISH 1914-10-01
132 Ceodes umbellifera Lanai Preserved_Specimen Lanai BISH
133 Ceodes umbellifera [Hawaiian Islands; U.S.A. Preserved_Specimen Mann, H. 347 BISH 1864-01-01
134 Ceodes umbellifera [Hawaiian Islands; U.S.A. Preserved_Specimen Mann, H. BISH 1864-01-01
135 Ceodes umbellifera Wailupe Preserved_Specimen Hillebrand, W.B. Oahu BISH
136 Ceodes umbellifera R fork, Wailupe Valley Preserved_Specimen Garber, D.W. 159 Oahu BISH 1920-01-12
137 Ceodes umbellifera Hii Mts Preserved_Specimen Forbes, C.N. 645 Kauai BISH 1916-10-20
138 Ceodes umbellifera Manoa Cliffs Trail Preserved_Specimen Sohmer, S.H. 6113 Oahu BISH 1969-09-27
139 Ceodes umbellifera Pole Line Trail (S side) Preserved_Specimen Herbst, D.R. 2856 Kauai BISH 1972-12-15
140 Ceodes umbellifera Manu Kani Reminent dryland forest on old weathered `a`a lava, in lee of ridge. Preserved_Specimen Tree 3 m ht, flowers pinkish. Kirch, P.V. 552 Maui BISH 1996-07-21
141 Ceodes umbellifera Waianae Mts., Puu Manawahua wooded gulch Preserved_Specimen 15' tree St.John, H. 9900 Oahu BISH 1929-09-29
142 Ceodes umbellifera Kahuna Stream above intake, moist cliffs Preserved_Specimen 5 m x 1 dm tree St.John, H. 17671 Maui BISH 1936-12-19
143 Ceodes umbellifera Lihue District. Hoary Head Range. North side of range between Haupu and Kamaulele In secondary forest below sheer rock face dominated by Aleurites, Pisonia, Cordyline and Asplenium nidus. Preserved_Specimen Common tree of 20 ft. Leave dark, glossy green above, dull, paler below; stems pale brown. Flowers: in bud green at apex with reddish base and pedicel becoming yellow-green and finally opening pale peach to whitish. Photinia-like odour. Flynn, T.W. 2939 Kauai BISH 1988-04-21
144 Ceodes umbellifera Hanalei District. Awaawapuhi, Na Pali coast. Lowland dry non-native herbland-grassland-shrubland. Preserved_Specimen Tree, 10 meters tall, 2 meter wide trunk, in flower, single tree in valley. Wood, K.R. 17949 Kauai BISH 2018-10-04
145 Ceodes umbellifera Manoa Cliffs Trail moist; Psidium guajava Preserved_Specimen 4 m shrub Takeuchi, W.N. 75 Oahu BISH 1984-02-26
146 Ceodes umbellifera Wahiawa Botanic Garden partial shade Preserved_Specimen 6 m tree, multi-trunked; flower whitish, strong odor, afternoon Lau, J. 2302 Oahu BISH 1986-05-02
147 Ceodes umbellifera Located on the Mauna Loa Strip Road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 ° north of road. (plot 19) Acacia - Sapindus savanna community Preserved_Specimen Tree Newell, C.L. 152 Hawaii BISH 1966-07-26
148 Ceodes umbellifera Waiahole, base of Koolau divide exotic riparian Preserved_Specimen large tree Takeuchi, W.N. 2521 Oahu BISH 1985-10-26
149 Ceodes umbellifera Moanalua Valley, power line trail, ~0.2 mi from jeep road wet; Diospyros, Clidemia hirta, Phaius tankervillae, Elaeocarpus bifidus, Adenophorus, Peperomia Preserved_Specimen 15' tree Gustafson, R. 2006 Oahu BISH 1980-12-09
150 Ceodes umbellifera Waianae Mountains: Kuaokala, Haili Gulch Closed forest. Preserved_Specimen Egler, F.E. 37184 Oahu BISH 1937-07-17
151 Ceodes umbellifera Hana, Puu Ki woods Preserved_Specimen 8 m x 30 cm tree St.John, H. 17888 Maui BISH 1936-12-31
152 Ceodes umbellifera Waianae Mountains: Kuaokala, Haili Gulch Closed forest. Preserved_Specimen Egler, F.E. 37183 Oahu BISH 1937-07-17
153 Ceodes umbellifera Tantalus, W of crater forest Preserved_Specimen 4 m tree Bryan Jr., E.H. 1425 Oahu BISH 1940-06-23
154 Ceodes umbellifera 2nd N fork, Kipapa Gulch, Waipio, Koolau Mts along stream bed Preserved_Specimen 30' tree Hosaka, E.Y. 833 Oahu BISH 1932-11-13
155 Ceodes umbellifera Manoa Cliffs trail, near Tantalus/Pauoa Flats jct beneath Metrosideros tremuloides, Hibiscus arnottianus Preserved_Specimen 3-4 m shrub Takeuchi, W.N. 74 Oahu BISH 1984-02-26
156 Ceodes umbellifera Hana, Haleakala, E fork of Oheo stream forest with Aleurites, Psidium Preserved_Specimen up to 12' tree; leaves dark green above with yellowish green midrib, below paler; petals white within, pinkish brown without with white margin; pedicels pale green to brownish green; flowers with odor of semen. Flynn, T.W. 2418 Maui BISH 1987-10-13
157 Ceodes umbellifera Wailuku, Iao Valley Narrow stream gorge. Preserved_Specimen St.John, H. 10282 Maui BISH 1930-02-09
158 Ceodes umbellifera Keanae Valley, Koolau Forest Reserve Pocket of Native Plants; 3-4 mi. into valley along jeep road Preserved_Specimen Sohmer, S.H. 6584 Maui BISH 1970-08-18
159 Ceodes umbellifera Waiahuakua Valley, Hanakoa Wet, wooded slope Preserved_Specimen Tree 5 m. x 12 cm. Flowers pink without, white within St.John, H. 23151 Kauai BISH 1947-12-31
160 Ceodes umbellifera Kalalau Valley. Upper reaches of stream in East side of Valley Riparian forest along valley floor dominated by Psidium guayava and Aleurites moluccana with vriable canopy 10-15 m tall. Occasional remnant patches of Psydrax oderata. Weedy, open understory with Christella dentata and Blechnum, Lantana camara and patches of Nephrolepis exaltata. Pipturis sp. occasional. Preserved_Specimen Tree 7 m tall, 25 cm dbh. Obovate lvs to 30 cm long. Frts on panicles, 6 cm long, 4-angled with sticky, resinous exudate along corners. Seed accessionaed with NTBG Living Collections (acc#090256) Trauernicht, C. 757 Kauai BISH 2009-02-26
161 Ceodes umbellifera Manoa Cliffs Trail moist; Psidium guajava Preserved_Specimen 4 m shrub Takeuchi, W.N. 75 Oahu BISH 1984-02-26
162 Ceodes umbellifera Waiahole, base of Koolau divide exotic riparian Preserved_Specimen large tree Takeuchi, W.N. 2521 Oahu BISH 1985-10-26
163 Ceodes umbellifera Na Pali Coast, Awa awapuhi Valley, Growing with Aleurites moluccana. Preserved_Specimen Single medium-size tree, flowers white. Corn, C. 390 Kauai BISH 1979-07-19
164 Ceodes umbellifera Huliwai Valley, below Honouliuli FR, Waianae Mts Osmanthus, Metrosideros, Eucalyptus Preserved_Specimen 15' x 10 tree; leaves 4 in a whorl; obovate, 17x7 cm, glabrous; inflorescence axillary, subumbellate; flower with greenish to yellow perigone Spence, G. 37 Oahu BISH 1972-03-26
165 Ceodes umbellifera Kauai Wet countryside Preserved_Specimen 40-50 ft. tree Thropp, J. Kauai BISH 1965-08-20
166 Ceodes umbellifera NE Kahaluu, head Coffea, Cordyline Preserved_Specimen fertile; cream colored perigones; odor similar to mango, heavy, oppressive Takeuchi, W.N. 2933 Oahu BISH 1986-10-25
167 Ceodes umbellifera Ekahanui Urera kaalae Preserved_Specimen in flower Takeuchi, W.N. 2642 Oahu BISH 1986-05-25
168 Ceodes umbellifera Haena, Waiahuakua Valley mesic forest; Aleurites, Diospyros, Nestegis, Pisonia umbellifera, Wikstroemia oahuensis, Freycinetia arborea, Ochrosia kauaiensis, Psydrax odorata Preserved_Specimen tree; trunks 50-100 cm diam., brown gray, mottled green white gray Wood, K.R. 12159 Kauai BISH 2006-09-24
169 Ceodes umbellifera Napali coast; Hanakapiai, Maunapuluo On cliff. Preserved_Specimen St.John, H. 10891 Kauai BISH 1930-12-30
170 Ceodes umbellifera Palolo, Waiomao Valley, Koolau Mts moist Preserved_Specimen 12' tree, erect Nishida, T. Oahu BISH 1943-12-05
171 Ceodes umbellifera Haupu (Hoary Head Mts.,) Kipu Moist forest Preserved_Specimen Slender tree 10 ft. tall; leaves on other plants nearly up to 15 in. long) St.John, H. 13673 Kauai BISH 1933-12-25
172 Ceodes umbellifera East Maui, Hana District, Hana, Puu Ki woods Preserved_Specimen 8 m x 30 cm tree St.John, H. 17888 Maui BISH 1936-12-31
173 Ceodes umbellifera Waikane, Schofield trail woods Preserved_Specimen 15' tree St.John, H. 10167 Oahu BISH 1930-01-19
174 Ceodes umbellifera Waianae Mountains: Kuaokala, Haili Gulch Closed forest. Preserved_Specimen Root fleshy. Egler, F.E. 37185 Oahu BISH 1937-07-17
175 Ceodes umbellifera NW Haleakala, Koolau Forest Reserve, Kailua Stream wet ohia forest with native trees, shrub, treefern Preserved_Specimen 4 m x 10 cm Warshauer, F.R. 2781 Maui BISH 1980-07-11
176 Ceodes umbellifera East Maui, Nahiku, about .3 km upstream from outlet of Hanawi stream, on E bank of stream Lowland wet forest with mixed native and alien species: Pandanus, Pisonia, Cyrtandra, Syzygium malaccense, Aleurites, etc. Preserved_Specimen Tree 4m tall Price, J.P. 16 Maui BISH 2000-08-26
177 Ceodes umbellifera woods of Hilo wet woods Preserved_Specimen tree 15 ft. angular Rock, J.F.C. Hawaii BISH 1958-10-05
178 Ceodes umbellifera Waianae Mts., Puu Hapapa, Honouliuli, Popouela ridge moist ridge Preserved_Specimen 20' tree St.John, H. 13995 Oahu BISH 1934-01-07
179 Ceodes umbellifera Kaluaa, streambed Pipturus, Streblus Preserved_Specimen 4 m tree Takeuchi, W.N. 2383 Oahu BISH 1985-08-10
180 Ceodes umbellifera Waiahole, base of Koolau divide exotic riparian Preserved_Specimen large tree Takeuchi, W.N. 2521 Oahu BISH 1985-10-26
181 Ceodes umbellifera Koolau Mts., Kawaiiki Ditch trail open Preserved_Specimen 10-15 trees, in flower Herat, T. 398 Oahu BISH 1973-07-02
182 Ceodes umbellifera Kawaihau District. Lihue-Koloa Forest Reserve. Kuilau trail from Keahua Arboretum Highly disturbed forest of Metrosideros, Pisonia, Psidium and Albizia with Dicranopteris covered slopes. Preserved_Specimen Tree of 25 ft. f Leaves dark green semi-glossy above, dull, paler green below. Perianth in bud brownish-pink, whiter near apex. Open flowers dirty white. flowers semen scented. Flynn, T.W. 3306 Kauai BISH 1989-04-07
183 Ceodes umbellifera Kalalau Valley, hanging valley abovce Keanapuka falls with jajor headwater springs. Psidium guajava on banks with tall Aleurites in gulch & understory dominated by Lantana camara-relic Metrosideros, Diospyros, Pisonia umbellifera, Psydrax, Psychotria mariniana, Syzygium sandwicensis. Preserved_Specimen Tree 8 m tall, trunk dark brown with white & gray lichen, in fl and some frt. Wood, K.R. 13419 Kauai BISH 2009-01-21
184 Ceodes umbellifera Central Lualualei, below Kanehoa, valley 5 forest grove Preserved_Specimen 6 m tree; buds Christophersen, E. 3704 Oahu BISH 1932-06-15
185 Ceodes umbellifera Located on the Mauna Loa Strip Road about 3.6 miles from intersection with Kona-Hilo Highway, into the first section of Kipuka Ki. Plant was about 30 meters into forest at right of road. (plot 39) Acacia - Sapindus - Microlepia forest community Preserved_Specimen Tree Newell, C.L. 200 Hawaii BISH 1966-07-28
186 Ceodes umbellifera Lower Nanawale Forest Reserve, Near begining of transect D on border of Forest Reserve and fallow papaya fields On well drained a`a, in moderate to full shade. Preserved_Specimen Tree height approx. 10m. Croft, L.K. 790038 Hawaii BISH 1979-02-22
187 Ceodes umbellifera Ekahanui gulch Euphorbia forbesii, Claoxylon, Psychotria hathewayi, Alectryon, Urera glabra Preserved_Specimen Takeuchi, W.N. 2061 Oahu BISH 1985-03-10
188 Ceodes umbellifera Koolau Mountains; Waipio, ridge S of Kipapa Gulch Wooded slope. Preserved_Specimen Tree 20' tall, 1' diam.; leaves fleshy. St.John, H. 10020 Oahu BISH 1929-11-20
189 Ceodes umbellifera Lower Nanawale Forest Reserve, near beginning of transect D on border of Forest Reserve On border of fallow papaya fields. In association with Psidium guajava, Aleurites and Rubus rosifolius. On well drained `a`a, in moderate to full shade. Preserved_Specimen Tree height approxiamately 10 m. Croft, L.K. 790038 Hawaii BISH 1979-02-22
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