Melicope hawaiensis

(Wawra) T.G.Hartley & B.C.Stone (1989)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Melicope

alani, alani kuahiwi, manena, mokihana kūkae moa

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Description

Key Characters:

Growth Form: Shrubs or trees 3–10 m tall, bark smooth, pale brown, new growth usually yellowish brown or reddish brown puberulent, sometimes becoming glabrate.

Stems:

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly to broadly elliptic, oblong–elliptic, or elliptic–ovate, sometimes thinly so, 5.5–20 cm long, (2.3–)3–8 cm wide. Apex obtuse to rounded or sometimes subacute, ± emarginate. Base cuneate to truncate or weakly cordate. Upper surfaces glabrous or midrib sometimes sparsely puberulent; lower surfaces glabrate to sparsely puberulent, especially along midrib, or densely tomentose; coriaceous. Margins entire, flat or revolute. Primary lateral veins 7–18 pairs, somewhat prominent on lower surface, venation pinnate, lateral veins connected by an arched submarginal vein, higher order venation forming a reticulate pattern. Petioles 12–45 mm long. Stipules absent.

Flowers: Flowers usually 3–27 in sparsely to densely puberulent or densely short–pilose, axillary cymes, peduncles 3–30(–45) mm long, pedicels 4–6 mm long, bracteoles puberulent, bracts small, opposite. Calyx of 4 sepals; sepals ovate, 2.8–3.1 mm long, densely tomentulose, imbricate. Corolla of 4 petals; petals valvate, often reddish within, oblong–deltate, 4–4.5 mm long, tomentulose. Stamens 8, in 2 whorls, distinct, reduced but always present in functionally pistillate flowers. Ovary superior, densely yellowish brown or golden tomentulose; style hirtellous, ovary 4-celled and 4-lobed, carpels connate, only slightly distinct at apex to almost entirely distinct; ovules 2 per cell, hemitropous, 1 ascending; style 1, terminating in 4 short stigmatic branches.

Fruit: Fruit 16–34 mm wide; of distinct follicles 8–17 mm long; exocarp densely golden brown to reddish brown short–pilose; the hairs sometimes yellowish green; endocarp densely pilose.fruit; the carpels dehiscing along the upper suture; exocarp. Seeds 1–2 per follicle; 4–7 mm long; glossy black when ripe; crustaceous; ovoid; sometimes angled from compression; embryo straight in fleshy endosperm; radicle short; cotyledons compressed; ovate; putamen brown; warty.

Ploidy:

Habitat: Occurring in dry forest and sometimes mesic forest.

Elevation Range: 610–1,220 m.

Historical Distribution

Images

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Taxon 38: 121 (1989)

Other References

Wagner et al. 1990:1189 (Mo, L, M, H [as Pelea hawaiensis]); Hartley & Stone 1989:121 (COMBNOV, Syn. P. hawaiensis = M. hawaiensis); Hank Oppenheimer pers. obs./Wood 12744 PTBG (L†, 2008)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Melicope hawaiensis Kaiholena Preserved_Specimen Forbes, C.N. 351 Lanai BISH 1917-09-01
2 Melicope hawaiensis Puuwaawaa hill, N. Kona Preserved_Specimen Rock, J.F.C. 3654 Hawaii BISH 1909-06-10
3 Melicope hawaiensis Haalelepaakai Preserved_Specimen Munro, G.C. 225 Lanai BISH 1915-02-16
4 Melicope hawaiensis Lanaihale, SW face Preserved_Specimen Munro, G.C. 542 Lanai BISH 1916-04-10
5 Melicope hawaiensis Kakapala [Kapapala?, Kau Preserved_Specimen 20 ft. tall, well-branched, rounded crown. Wood rather hard, very thick bark. Yamayoshi, H. Hawaii BISH 1939-05-10
6 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 132 Lanai BISH 1913-06-01
7 Melicope hawaiensis Kakapala [Kapapala?, Kau Preserved_Specimen 20 ft. tall, well-branched, rounded crown. Wood rather hard, very thick bark. Yamayoshi, H. Hawaii BISH 1939-05-10
8 Melicope hawaiensis Haalelepaakai and Lanaihale, between Preserved_Specimen Munro, G.C. 422 Lanai BISH 1915-04-08
9 Melicope hawaiensis Kawaihae-Uka Preserved_Specimen Tree 10 to 15 ft. high Rock, J.F.C. Hawaii BISH 1955-12-01
10 Melicope hawaiensis Kawaihae uka Preserved_Specimen Hillebrand, W.B. Hawaii BISH 1862-01-01
11 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 130 Lanai BISH 1913-06-01
12 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 130 Lanai BISH 1913-06-01
13 Melicope hawaiensis Ridge below Puu Kolekole Preserved_Specimen Forbes, C.N. 126 Molokai BISH 1912-07-01
14 Melicope hawaiensis Waiakeakua Preserved_Specimen Munro, G.C. 143 Lanai BISH 1927-04-18
15 Melicope hawaiensis Mountains at E end Preserved_Specimen Munro, G.C. 251 Lanai BISH 1913-06-01
16 Melicope hawaiensis Makawao Forest Reserve, along Borge Ridge road Preserved_Specimen 4 m tree Hobdy, R.W. 1375 Maui BISH 1982-04-30
17 Melicope hawaiensis East Maui, Auwahi Preserved_Specimen Rock, J.F.C. 8676 Maui BISH 1910-11-01
18 Melicope hawaiensis East Maui, Makawao Preserved_Specimen Rock, J.F.C. 8550 Maui BISH 1910-09-01
19 Melicope hawaiensis Puuwaawaa Preserved_Specimen Forbes, C.N. 18 Hawaii BISH 1911-06-03
20 Melicope hawaiensis Olowalu Valley, central ridge Preserved_Specimen Forbes, C.N. 2345 Maui BISH 1920-05-12
21 Melicope hawaiensis Lanai Preserved_Specimen Forbes, C.N. 362 Lanai BISH 1919-09-01
22 Melicope hawaiensis Waiapaa Preserved_Specimen Munro, G.C. 593 Lanai BISH 1916-09-18
23 Melicope hawaiensis W slope, main ridge Preserved_Specimen Munro, G.C. 467 Lanai BISH 1915-04-21
24 Melicope hawaiensis East Maui, Auwahi Preserved_Specimen Rock, J.F.C. 8676 Maui BISH 1910-11-01
25 Melicope hawaiensis Kilauea, Kipuka Puaulu Preserved_Specimen MacDaniels, L.H. 212 Hawaii BISH 1926-11-10
26 Melicope hawaiensis Mahana Preserved_Specimen Munro, G.C. 440 Lanai BISH 1915-05-29
27 Melicope hawaiensis Mahana Preserved_Specimen Munro, G.C. 254 Lanai BISH 1915-05-29
28 Melicope hawaiensis Puaulu, Kilauea, Volcano Preserved_Specimen Rock, J.F.C. 8775 Hawaii BISH 1911-07-01
29 Melicope hawaiensis Waiapaa Preserved_Specimen Munro, G.C. 468 Lanai BISH 1916-09-18
30 Melicope hawaiensis East Maui, Auwahi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2099 Maui BISH 1920-03-24
31 Melicope hawaiensis Olowalu Valley, central ridge Preserved_Specimen Forbes, C.N. 2345 Maui BISH 1920-05-12
32 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 105 Lanai BISH 1913-06-01
33 Melicope hawaiensis East Maui, Auwahi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2098 Maui BISH 1920-03-24
34 Melicope hawaiensis Hawaii National Park, Bird Park Preserved_Specimen Fagerlund, G.O. 573 Hawaii BISH 1943-05-24
35 Melicope hawaiensis Puuwaawaa, N. Kona Preserved_Specimen Rock, J.F.C. 10211 Hawaii BISH 1912-03-01
36 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 105 Lanai BISH 1913-06-01
37 Melicope hawaiensis Kalulu Preserved_Specimen Munro, G.C. 473 Lanai BISH 1915-11-30
38 Melicope hawaiensis Kau Desert, Kau Preserved_Specimen Forbes, C.N. 385 Hawaii BISH 1911-08-02
39 Melicope hawaiensis Lanaihale Preserved_Specimen Munro, G.C. 544 Lanai BISH 1916-04-10
40 Melicope hawaiensis Kau Preserved_Specimen Hillebrand, W.B. Hawaii BISH
41 Melicope hawaiensis Lanai Preserved_Specimen Hillebrand, W.B. Lanai BISH 1870-07-01
42 Melicope hawaiensis Kilauea, Kipuka Puaulu Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
43 Melicope hawaiensis Hookio Ridge Preserved_Specimen Munro, G.C. Lanai BISH 1915-05-06
44 Melicope hawaiensis Ocean View Estates, Kau Preserved_Specimen Montgomery, S.L. Hawaii BISH 1971-08-01
45 Melicope hawaiensis Lanai Preserved_Specimen Hillebrand, W.B. Lanai BISH
46 Melicope hawaiensis Puaulu Preserved_Specimen Russ, G.W. Hawaii BISH 1927-08-01
47 Melicope hawaiensis Lanai Preserved_Specimen Hillebrand, W.B. Lanai BISH 1870-07-01
48 Melicope hawaiensis Kaupo, Planch. Ridge Preserved_Specimen Holt, R.A. Maui BISH 1981-06-24
49 Melicope hawaiensis Bird Park, Kilauea, Kau Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-08-09
50 Melicope hawaiensis Kawaihaeiuka Preserved_Specimen Hillebrand, W.B. Hawaii BISH
51 Melicope hawaiensis Mahana Preserved_Specimen Munro, G.C. Lanai BISH 1915-05-15
52 Melicope hawaiensis Kaupo, Planch. Ridge Preserved_Specimen Holt, R.A. Maui BISH 1981-06-24
53 Melicope hawaiensis S slope of Haleakala Preserved_Specimen Forbes, C.N. Maui BISH 1920-03-01
54 Melicope hawaiensis Kaiholena Preserved_Specimen Munro, G.C. Lanai BISH 1915-10-23
55 Melicope hawaiensis Bird Park (Kipuka Puaulu) Preserved_Specimen Morley, H. Hawaii BISH 1935-08-01
56 Melicope hawaiensis unknown Preserved_Specimen Gaudichaud-Beaupré, C. BISH 1819-01-01
57 Melicope hawaiensis unknown Preserved_Specimen Morley, H. BISH 1934-06-01
58 Melicope hawaiensis unknown Preserved_Specimen Morley, H. BISH 1934-06-01
59 Melicope hawaiensis East Maui, Auwahi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2115 Maui BISH 1920-11-27
60 Melicope hawaiensis Waiakeakua Preserved_Specimen Munro, G.C. 459 Lanai BISH 1929-10-24
61 Melicope hawaiensis Mahana Preserved_Specimen Munro, G.C. 466 Lanai BISH 1915-05-19
62 Melicope hawaiensis Puuwaawaa, N. Kona Preserved_Specimen Rock, J.F.C. 3561 Hawaii BISH 1909-06-06
63 Melicope hawaiensis Mountains at E end Preserved_Specimen Munro, G.C. 251 Lanai BISH 1913-06-01
64 Melicope hawaiensis Waiakeakua Preserved_Specimen Munro, G.C. 82 Lanai BISH 1927-06-14
65 Melicope hawaiensis Puaulu, Kilauea, Volcano Preserved_Specimen Rock, J.F.C. 8775 Hawaii BISH 1911-07-01
66 Melicope hawaiensis Lanai Preserved_Specimen Mann, H. 371 Lanai BISH 1864-01-01
67 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 132 Lanai BISH 1913-06-01
68 Melicope hawaiensis Kipuka Puaulu, Kilauea Preserved_Specimen Degener, O. 8587 Hawaii BISH 1922-12-22
69 Melicope hawaiensis Puuwaawaa, N. Kona Preserved_Specimen Rock, J.F.C. 12966 Hawaii BISH 1917-08-01
70 Melicope hawaiensis Kohala Range Preserved_Specimen Hillebrand, W.B. 247 Hawaii BISH
71 Melicope hawaiensis Olowalu Valley, central ridge Preserved_Specimen Forbes, C.N. 2345 Maui BISH 1920-05-12
72 Melicope hawaiensis Puaulu, Kilauea Preserved_Specimen Rock, J.F.C. 10210 Hawaii BISH 1911-07-01
73 Melicope hawaiensis Kipuka Puaulu Preserved_Specimen Russ, G.W. Hawaii BISH 1930-06-01
74 Melicope hawaiensis S. Kona Preserved_Specimen Hillebrand, W.B. Hawaii BISH 1868-01-01
75 Melicope hawaiensis Pahilai Forest, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2082 Maui BISH 1920-03-23
76 Melicope hawaiensis Gulches west of Kamalo Canyon Preserved_Specimen Hobdy, R.W. 3036 Molokai BISH 1989-03-30
77 Melicope hawaiensis Valley on SE ridge of mountains on N end of Puhielelu Ridge, on telephone line ridge Preserved_Specimen Munro, G.C. 545 Lanai BISH 1916-04-10
78 Melicope hawaiensis Kalulu Preserved_Specimen Munro, G.C. 472 Lanai BISH 1915-10-30
79 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 132 Lanai BISH 1913-06-01
80 Melicope hawaiensis unknown Preserved_Specimen Hillebrand, W.B. BISH
81 Melicope hawaiensis unknown Preserved_Specimen Gaudichaud-Beaupré, C. BISH
82 Melicope hawaiensis Kauaula, SE rim mesic forest, steep to vertical rocky slopes; 30% Metrosideros, 5-7 m, 10% Dodonaea viscosa 3-4 m, 10% Coprosma foliosa, 2 m, 5% Pipturus albidus, 3 m, Leptecophylla tameiameiae, Dianella sandwicensis, Dicranopteris linearis, Bidens micrantha, Neraudia melastomifolia Preserved_Specimen 2 m tree; abundant fruit Wood, K.R. 12102 Maui BISH 2005-08-24
83 Melicope hawaiensis Waiakuilani Gulch, Kamalo forest Preserved_Specimen 6 m x 8 cm tree; leaves subcoriaceous, above dark green, shiny, below cinereous; cocci cinereous; petals cream colored St.John, H. 19875 Molokai BISH 1938-12-28
84 Melicope hawaiensis Olinda, edge of forest reserve slopes Preserved_Specimen 12' tree, compact, shapely Degener, O. 22112 Maui BISH 1952-03-26
85 Melicope hawaiensis East Maui, Makawao Dist., Kahakapao drainage basin on sleep (sic) slope at edge of narrow ridge. Vegetation: Mesic Forest with Acacia, Metrosideros, Fraxinus, Psidium, Dicranopteris, Coprosma, Freycinetia, Canavalia. Preserved_Specimen Single 5 m. tree. Oppenheimer, H.L. 100503 Maui BISH 2005-10-08
86 Melicope hawaiensis Puehuehunui, S of Kauaula rim small drainage, mesic forest; ferns, Metrosideros polymorpha, Cheirodendron trigynum, Dodonaea viscosa, Nestegis sandwicensis, Hedyotis acuminata, Cyrtandra grayi, Melicope hawaiiense Preserved_Specimen 4-5 m tree; leaves dark green, below grayish scruf; fruit yellow hairs on exocarp Wood, K.R. 11938 Maui BISH 2006-06-21
87 Melicope hawaiensis East Maui, Makawao Forest Reserve ridge tops, upper gulch slopes Preserved_Specimen 4-6 m tree Hobdy, R.W. 1882 Maui BISH 1983-12-20
88 Melicope hawaiensis Makolelau, below and west of Puu Kolekole, above 2nd gulch to east of Contour road gulch Preserved_Specimen Small tree about 8 ft. tall. Lvs. Clustered towards apes, down flexed w/ bm-grey pubescence below. Top of leaf dk. Green w/ sparse pubescence. Capsuels dense pubescence, yellow, rugose. Calyxes green, overlapping in bud. Calyx connate in flw. New growth grey-green w/ brn pubescence, stem dark brown w/ pubescence Tangalin, N. 650 Molokai BISH 2005-12-27
89 Melicope hawaiensis Pohakuloa Military Training Area Vegetation: Open metrosideros forest. Growing on aa lava substrate Preserved_Specimen Warshauer, F.R. 811 Hawaii BISH 1977-01-11
90 Melicope hawaiensis E Makolelau Ridge, Puu Kolekole S to coast, road dry; Diospyros, Pouteria, Canthium, Nestegis, Pleomele Preserved_Specimen 3 m tree; bud and fruit green Lorence, D.H. 5673 Molokai BISH 1987-10-23
91 Melicope hawaiensis Hulopoe Gulch, head of, at Munro Trail dark woods Preserved_Specimen 12' tree, few branches Degener, O. 28600 Lanai BISH 1963-08-01
92 Melicope hawaiensis East Maui, Makawao District, Makawao Forest Reserve, Borges Ridge, on E rim of W tributary of Waiohiwi Stream Vegetation: Metrosideros polymorpha var. glaberrima, Cheirodendron trigynum, Acacia koa, Scaevola chamissoniana, Coprosma foliosa, Dianella sandwicensis, Smilax melastomifolia, Alyxia oliviformis, Cinchona pubescens, Setaria palmifolia, Dryopteris glabra. Preserved_Specimen Oppenheimer, H.L. 60145 Maui BISH 2001-06-18
93 Melicope hawaiensis E side, W fork, Kawela Gulch Styphelia, Dodonaea, Exocarpos Preserved_Specimen 8' x 4 tree Anderson, S.J. 553 Molokai BISH 1982-05-29
94 Melicope hawaiensis Puuwaawaa cone, on transect T9 Native remnant forest in north facing slopes. Growing among Pittosporum hosmeri and Nestegis sandwicensis. Preserved_Specimen 25' tree, most inds. in fruit. Takeuchi, W.N. 5820 Hawaii BISH 1989-04-01
95 Melicope hawaiensis East Maui, E of Makawao woods Preserved_Specimen Forbes, C.N. 234 Maui BISH 1910-07-01
96 Melicope hawaiensis head of Hulopoe Gulch forest Preserved_Specimen sprawling tree Degener, O. 28523 Lanai BISH 1963-08-13
97 Melicope hawaiensis west Maui, Lahaina District, Manawainui Gulch, ridge between west & east forks. Vegetation: Metrosideros, Diospyros, Nestegis, Dodonaea, Leptecophylla, Myrsine, Wikstroemia, Osteomeles, Bidens, Eragrostis, Pteridium, Asplenium, Panicum, Cocculus, Casuarina, Grevillea, Psidium, Erigeron, Senecio, Andropogon, Setaria. Preserved_Specimen Rare, one mature 1.5m tall and two immature plants observed on ridgetop. Plants in deep gulches nearby are trees with large leaves that are densely pubescent on lower leaf surface. Oppenheimer, H.L. 101312 Maui BISH 2013-10-17
98 Melicope hawaiensis Makolelau gulch, below Puu Kolekole Dodonaea, Styphelia, Vaccinium, Psychotria, Diospyros, Sophora chrysophylla, Neraudia sericea, Lipochaeta rockii, Bidens menziesii, Myoporum, Wikstroemia, Metrosideros, Pleomele, Pittosporum, Chamaesyce, Lysimachia, Viola, Pipturus, Scaevola, Urera glabra, Lantana, Melinis minutiflora, Schiedea Preserved_Specimen 10' tree; fruit Perlman, S. 6671 Molokai BISH 1987-10-23
99 Melicope hawaiensis Kipuka Puaulu Closed mixed forest. Preserved_Specimen Clarke, G. 169 Hawaii BISH 1978-08-16
100 Melicope hawaiensis East Maui, Makawao Forest Reserve, Borge Ridge Road, West side of road along edge of gulch Aspect: N. Patch of trees along edge of gulch. Mesic forest: Acacia koa, Metrosideros polymorpha, Psidium, Hedychium, Setaria palmifolia, Rubus argutus, Dicranopteris linearis Preserved_Specimen 2-5 m tall, puberulent exocarp. Mostly glabrous endocarp except along sutures Z. Pezzillo 1215 Maui BISH 2025-07-20
101 Melicope hawaiensis Puuwaawaa cone; N-facing slopes; transect T9 native remnant forest; among Pittosporum hosmeri and Nestegis sandwicensis; locally common, most inds. in fruit Preserved_Specimen 25 ft. tree Takeuchi, W.N. 5820 Hawaii BISH 1989-04-01
102 Melicope hawaiensis W Maui, Lahaina District, Puehuehunui, 3rd gulch SE of Kauaula Valley, 2nd Gulch NW of Luakoi Ridge Growing on steep, leeward facing slope. Vegetation: Metrosideros, Dodonaea, Nestegis, Santalum, Myrsine, Wikstroemia, Leptecophylla, Coprosma, Myoporum, Psydrax, Diospyros, Pleomele, Pisonia, Melicope sp, Freycinetia, Alyxia, Microlepia, Doodia, Dryopteris, Bidens, Osteomeles, Carex. Preserved_Specimen Single 6m tall, multi-trunked emergent tree. Oppenheimer, H.L. 80630 Maui BISH 2006-08-22
103 Melicope hawaiensis Manuka NAR;, nature trail up to Kipuka Kilioe Metrosideros lowland mesic forest ca 25 - 30 m tall. W/ understory of Nestegis, Pisonia, Charpentiera, Psychotria hawaiiensis, Claoxylon, Myrsine lessertiana, Dodonaea viscosa, Alyxia oliviformis, Pipturus albidus, Antidesma platyphyllum & A. pulvinatum. Preserved_Specimen tree, 5 - 10 m tall, moderately branched, trunk gray-brown with slight orange tinge, occasional with Gouania vitifolia. Wood, K.R. 13203 Hawaii BISH 2008-08-07
104 Melicope hawaiensis Puuwaawaa cone; N. facing slopes on transect T9 native remnant forest; locally common, most inds. in fruit Preserved_Specimen 25 ft. tree among Pittosporum hosmeri and Nestegis sandwicensis Takeuchi, W.N. 5820 Hawaii BISH 1989-04-01
105 Melicope hawaiensis Keauhou Ranch, Kau District Growing in disturbed Metrosideros - Acacia forest with Myrsine, Myoporum, Rubus hawaiensis, Clermontia lindseyana, and ground cover of Dryopteris paleacea and Microlaena stipoides. Preserved_Specimen Tree 25' tall, d.b.h. 10, spread 30'. Green fruit and dried fruit and young inflorescence seen. Davis, J. 726 Hawaii BISH 1982-05-05
106 Melicope hawaiensis Kau Forest Reserve Rough lava Preserved_Specimen Tree, height 7 m. MacDaniels, L.H. 257 Hawaii BISH 1926-11-16
107 Melicope hawaiensis Hulopoe Gulch, head of, at Munro Trail Dark, rich woods. Preserved_Specimen Tree: 12 feet in height; few branched. Degener, O. 28600 Lanai BISH 1963-08-01
108 Melicope hawaiensis Laupahoehoe 1 Ahupuaa, Kaimu Stream, North Kohala District, Puu O Umi Natural Area Reserve. On steep slope below the Pali Kai Fence unit going down into Kaimu Gulch. Growing on steep slope with METPOL, RAUSAN, ANTPUL, SCHTER, PSICAT, PLUCAR, OSTANT, CHACEL, CHAOVA. Preserved_Specimen Small Tree about 4 m tall. Flowers greenish white, fruit green ManDeMark, J. Hawaii BISH 2017-12-12
109 Melicope hawaiensis Keauhou Ranch, Kaudistrict. USFS study area Disturbed Acacia koa - Metrosideros closed forest. Preserved_Specimen Tree, approx. 30' tall. Flowers light green. DBH 9, spread 15-20 ft. Clarke, G. 79.272 Hawaii BISH 1979-12-12
110 Melicope hawaiensis Kipuka Puaulu (Bird Park), Kilauea Occasional in open park-like woodland with lush grass, deep soil Preserved_Specimen Spreading, unbrella-crowned tree, 5 m. tall; flowers greenish; fruit dry Fosberg, F.R. 33275 Hawaii BISH 1951-02-11
111 Melicope hawaiensis West Maui, Pohakea mesic; Styphelia, Dodonaea, Metrosideros, Coprosma foliosa, Myrsine lanaiensis, Nestegis sandwicensis, Pittosporum glabrum, Pleomele auwahiensis, Microlepia strigosa, Dicranopteris linearis, Carex meyenii, C. wahuensis, Morelotia gahniiformis, Trisetum inaequale, Alyxia oliviformis, Smilax melastomifolia Preserved_Specimen tree; stems upright, moderately branched; leaves dark green, below soft pubescent tan brown; exterior carpels soft pilose golden brown Wood, K.R. 11563 Maui BISH 2005-10-11
112 Melicope hawaiensis Waiakuilani Gulch on steep windward facing slope in mesic forest. Vegetation: Metrosideros, Diospyros, Dodonaea, Coprosma, Alyxia, Pouteria, Wikstroemia, Pleomele, Myrsine, Dryopteris, Doryopteris, Leptecophylla, Pteridium, Peperomia, Viola, Chamaesyce, Dubautia, Streblus, Sophora Preserved_Specimen Uncommon, 4 trees observed in this location to 3 m. tall Oppenheimer, H.L. 40714 Molokai BISH 2007-04-05
113 Melicope hawaiensis Slope of Hualalai mauka of road leading from Huehue to Puuwaawaa moderately dry forest Preserved_Specimen Degener, O. 2331 Hawaii BISH 1926-08-24
114 Melicope hawaiensis Waiakeakua, hanging valley SW, W of divide Patch of dryish, shrubby forest. Preserved_Specimen Degener, O. 28805 Lanai BISH 1964-02-01
115 Melicope hawaiensis Hulopoe Gulch, head of, at Munro Trail dark woods Preserved_Specimen 12' tree, few branches Degener, O. 28600 Lanai BISH 1963-08-01
116 Melicope hawaiensis head of Hulopoe Gulch forest Preserved_Specimen sprawling tree Degener, O. 28523 Lanai BISH 1963-08-13
117 Melicope hawaiensis USA, Hawaiian Island, Lanai, Lanai'ihale, headwaters of Lopa Guch below Ha'alelepa'akai, 2900ft, Aspect: N Reminant mesic forest: Morella faya, Psidium cattleianum, Notelaea sandwincensis, Odontosoria chinesis, Pipturus albidus, Dicanopteris linearis, Diplopterygium pinnatum, Cibotium menziesii, Psychotria sp., Scaevola chamasoniana, Christella parasiticia, Doodia kunthiana, Adianthum hispidulum, Selaginella arbuscula, Blechnum appendiculatum Preserved_Specimen Single plant on upper half of steep drainage slope. Henderson, R. 73 Lanai BISH 2025-02-21
118 Melicope hawaiensis Waiakeakua, hanging valley SW, W of divide Patch of dryish, shrubby forest. Preserved_Specimen Tree 6 foot tall, erect, symmetrically branched. Degener, O. 28805 Lanai BISH 1964-02-01
119 Melicope hawaiensis East Maui, Makawao FR, Kahakapao drainage Moderate slope near top of gulch, windward, in shade. Associated vegetation: Metrosideros, Acacia, Nestegis, Pleomele, Ochrosia, Xylosma, Psychotria, Perrottetia, Cyrtandra, Freycinetia, Smilax, Ipomoea, Alyxia, Diplazium, Doodia, Asplenium, Dryopteris, Rubus, Fraxinus, Psidium, Hedychium. Preserved_Specimen To 5 m tree. Oppenheimer, H.L. 100646 Maui BISH 2006-10-25
120 Melicope hawaiensis East Maui, Hana District, SW Haleakala, Auwahi, 0.5 km SE of Puu Ouli dry forest, exotic grass and shrubs, native trees and shrubs Preserved_Specimen 2.75 m; in flower Higashino, P.K. 9262 Maui BISH 1980-06-25
121 Melicope hawaiensis west Maui, Lahaina District, Kaua`ula Valley, south slope. Vegetation: Myrsine, Diospyros, Metrosideros, Planchonella, Nestegis, Leptecophylla, Chrysodracon, Dodonaea, Charpentiera, Euphorbia, Myoporum, Santalum, Coprosma, Psychotria, Alyxia, Dryopteris, Ipomoea, Cocculus, Trisetum, Cyrtomium, Microlepia, Carex, Doodia, Bidens, Peperomia, Ageratina, Rubus, Oplismenus, Bromus, Erigeron. Preserved_Specimen Rare, a few scattered in this area. Tree 6m tall. Oppenheimer, H.L. 41607 Maui BISH 2016-04-13
122 Melicope hawaiensis East Maui, Hana District, SW Haleakala, Auwahi, 0.5 km SE of Puu Ouli dry forest, exotic grass and shrubs, native trees and shrubs Preserved_Specimen 4 m x 16 cm tree; in flower and fruit Higashino, P.K. 9269 Maui BISH 1980-06-25
123 Melicope hawaiensis Waiakeakua, SW, W of divide hanging valley, patch of dryish, shrubby forest. Preserved_Specimen 10 foot tall erect forest tree, long sprawling branches Degener, O. 28806 Lanai BISH 1964-02-01
124 Melicope hawaiensis Puuwaawaa; On ridge between gully 55 & 6, 10 ft. from barbwire fence above Open forest. Associated species: (hoawa, olive), pilo, Annona, Ricinus, (Cibotium), orchard grass, ohia Preserved_Specimen Old tree, 30 ft. tall. Fruit. Bark: blotched grays. Diameter breast high: 11 inches. Montgomery, S.L. 66 Hawaii BISH 1971-07-25
125 Melicope hawaiensis Makolelau gulch, below Puu Kolekole Dodonaea, Styphelia, Vaccinium, Psychotria, Diospyros, Sophora chrysophylla, Neraudia sericea, Lipochaeta rockii, Bidens menziesii, Myoporum, Wikstroemia, Metrosideros, Pleomele, Pittosporum, Chamaesyce, Lysimachia, Viola, Pipturus, Scaevola, Urera glabra, Lantana, Melinis minutiflora, Schiedea Preserved_Specimen 10' tree; fruit Perlman, S. 6671 Molokai BISH 1987-10-23
126 Melicope hawaiensis North Kona, Puuwaawaa, near Huehue Ranch Lava beds, in company with mamani and uhiuhi Preserved_Specimen Rock, J.F.C. 3561 Hawaii BISH 1909-06-06
127 Melicope hawaiensis Keauhou Ranch, Kau District Growing in disturbed Metrosideros - Acacia forest with Myrsine, Myoporum, Rubus hawaiensis, Clermontia lindseyana, and ground cover of Dryopteris paleacea and Microlaena stipoides. Preserved_Specimen Tree 25' tall, d.b.h. 10, spread 30'. Green fruit, dried fruit, and younf infl. seen. Davis, J. 726 Hawaii BISH 1982-05-05
128 Melicope hawaiensis East Maui, Haleakala, E. Kaupo gap steep ridge with dry forest Preserved_Specimen Crivellone, C. Maui BISH 1983-06-10
129 Melicope hawaiensis Olinda, edge of forest reserve slopes Preserved_Specimen 12' tree, compact, shapely Degener, O. 22112 Maui BISH 1952-03-26
130 Melicope hawaiensis East Maui, Makawao open forest Preserved_Specimen 29' tree Fagerlund, G.O. 1001 Maui BISH 1945-02-25
131 Melicope hawaiensis W Maui, ridge north of Pohakea Gulch rain forest, top of ravine, afternoon fog Preserved_Specimen Small tree. Degener, O. 2375 Maui BISH 1927-07-23
132 Melicope hawaiensis USA, Hawaiian Islands, Maui, West Maui, Pana'ewa section West Maui NAR, first ridge N of Kaua'ula Valley, N slope in mesic forest N. 8 short stature trees in cluster, mid slope, fruit and flower buds.Mesic: Metrosideros, Psychotria, Schinus, Notelaea, Dodonaea, Planchonella, Xylosma, Dracaena, Freycinettia, Ageratina, Nephrolepis, Sadleria, Blechnum, Microlepia Preserved_Specimen Pezzillo, Z. 1143 Maui BISH 2025-05-02
133 Melicope hawaiensis East Maui, woods above Makawao Woods Preserved_Specimen Mann, H. 377 Maui BISH 1864-01-01
134 Melicope hawaiensis Makolelau gulch, below Puu Kolekole Dodonaea, Styphelia, Vaccinium, Psychotria, Diospyros, Sophora chrysophylla, Neraudia sericea, Lipochaeta rockii, Bidens menziesii, Myoporum, Wikstroemia, Metrosideros, Pleomele, Pittosporum, Chamaesyce, Lysimachia, Viola, Pipturus, Scaevola, Urera glabra, Lantana, Melinis minutiflora, Schiedea Preserved_Specimen 10' tree; fruit Perlman, S. 6671 Molokai BISH 1987-10-23
135 Melicope hawaiensis W Maui, Lahaina District, Kauaula Valley, on SE wall of Valley Growing on steep, rocky slope. Associated vegetation: Metrosideros, Antidesma, Urera, Coprosma, Diospyros, Dubautia, Pipturus, Hedyotis, Melicope sp, Alyxia, Dodonaea, Cocculus, Pleomele, Zanthoxylum, Achyranthes, Microlepia, Cyrtomium, Pteris. Preserved_Specimen 5m tall tree. Oppenheimer, H.L. 80639 Maui BISH 2006-08-24
136 Melicope hawaiensis head of Hulopoe Gulch Wind-protected shrubby forest. Preserved_Specimen Sprawling tree Degener, O. 28523 Lanai BISH 1963-08-13
137 Melicope hawaiensis Mahana Preserved_Specimen Munro, G.C. 440 Lanai BISH 1915-05-29
138 Melicope hawaiensis Olinda, edge of forest reserve slopes Preserved_Specimen 12' tree, compact, shapely Degener, O. 22112 Maui BISH 1952-03-26
139 Melicope hawaiensis East Maui, Makawao, Kahakapao drainage basin E facing upper gulch slope; dense shade; Acacia, Metrosideros, Nestegis, Pisonia, Ochrosia, Pleomele, Coprosma, Kadua, Leptecophylla, Psychotria, Pouteria, Alyxia, Smilax, Doodia, Asplenium, Cyrtomium, Dryopteris. Preserved_Specimen Single 3 m tree, sprawling. Oppenheimer, H.L. 60725 Maui BISH 2007-06-26
140 Melicope hawaiensis In Kipuka Puaulu In grassy open woods Preserved_Specimen Degener, O. 8491 Hawaii BISH 1929-07-09
141 Melicope hawaiensis unamed gulch- 1st gulch east of road to Puu Kolekole on east side of East Kawela Gulch. lower, rocky gulch slope in remnant mesic forest patches. Vegetation: Dodonaea, Metrosideros, Diospyros, Schiedea, Pleomele, Pouteria, Wikstroemia, Leptecophylla, Bidens, Cyperus, Lipochaeta, Pouteria, Scaevola, Ageratina, Ochrosia, Pittosporum, Nestegis, Pisonia, Anthoxylum, Ipomoea, Melinis, Asplenium, Coprosma, Kadua, Rumex, Doryopteris, Chamaesyce, Carex, Osteomeles, Myrsine, Myoporum, Pteridium, Cocculus, Ricinis, Luzula. Preserved_Specimen Three trees to 4m tall. Oppenheimer, H.L. 10914 Molokai BISH 2009-01-13
142 Melicope hawaiensis Mts. near Koele Preserved_Specimen Forbes, C.N. 105 Lanai BISH 1913-06-01
143 Melicope hawaiensis Makawao Forest Reserve, along Borge Ridge road Preserved_Specimen 4 m tree Hobdy, R.W. 1375 Maui BISH 1982-04-30
144 Melicope hawaiensis West Maui, Hanaula Iki mesic; Styphelia, Dodonaea, Metrosideros, Coprosma foliosa, Myrsine lanaiensis, Nestegis sandwicensis, Pittosporum glabrum, Pleomele auwahiensis, Microlepia strigosa, Dicranopteris linearis, Carex meyenii, C. wahuensis, Morelotia gahniiformis, Trisetum inaequale, Alyxia oliviformis, Smilax melastomifolia Preserved_Specimen 2 m tree; moderately branched; leaves dull dark green, yellow green costa, below soft dull tan brown pubescence Wood, K.R. 11559 Maui BISH 2005-10-10
145 Melicope hawaiensis Makolelau, ridge S of Puu Kolekole dry; native shrubs and trees Preserved_Specimen Stemmermann, R.L. 3866 Molokai BISH 1979-06-20
146 Melicope hawaiensis Puu Waawaa, road west of corral and water tank lowland dry forest, aa lava with large surface boulders and rich brown soil, with Myrsine lanaiensis, Sophora chrysophylla, Nothocestrum breviflorum, Myoporum sandwicensis, Metrosideros polymorpha Preserved_Specimen tree, 3 meters tall, trunk gray-brown, 15 cm. diameter at base, with flower and old fruit carpels, near fence, rare in area Wood, K.R. 7877 Hawaii BISH 1999-06-29
147 Melicope hawaiensis West Maui, Lahaina District, Kauaula valley, below Helu summit, on west facing slope. Disopyros sadwicensis-Metrosidero mesic forest Dodonaea viscosa, Myrsine lanaiensis, Bidens micrantha, Coprosma foliosa, Myoporum, Nestegis sandwicensis, Chrysodracon, Leptecophylla, Remya mauiensis, Achyranthes splendens, Urera glabra. Preserved_Specimen Small tree 5 meter ht with flowers and fruit. Perlman, S. 24920 Maui BISH 2016-04-13
148 Melicope hawaiensis Kehewai Ridge North-facing slope, 50 m below Eucalyptus in Diospyros-Nestegis lowland dry forest with Nesoluma, Myoporum, Bobea, Lantana, Dodonaea, Pleomele, Rauvolfia, Melicope, Wikstroemia. Preserved_Specimen With fruit. Perlman, S. 6343 Lanai BISH 1987-08-25
149 Melicope hawaiensis Keauhou Ranch, Kau District In a disturbed Metrosideros - Acacia forest. Preserved_Specimen A tree of 6' with a 1 basal diameter. Clarke, G. 143 Hawaii BISH 1979-11-21
150 Melicope hawaiensis Mountains at E end Preserved_Specimen Munro, G.C. 251 Lanai BISH 1913-06-01
151 Melicope hawaiensis East Maui, Auwahi Preserved_Specimen Rock, J.F.C. 8676 Maui BISH 1910-11-01
152 Melicope hawaiensis East Maui, Auwahi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2097 Maui BISH 1920-03-24
153 Melicope hawaiensis Puuwaawaa Preserved_Specimen Skottsberg, C.J.F. 2114 Hawaii BISH 1922-09-26
154 Melicope hawaiensis Kilauea, Volcano Preserved_Specimen Rock, J.F.C. 8774 Hawaii BISH 1911-07-01
155 Melicope hawaiensis S of Puu Kolekole Preserved_Specimen small tree Pratt, T.K. 79 Molokai BISH 1972-01-06