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Touchardia sandwicensis

(Wedd.) Magnacca (2025)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Rosales Family: Urticaceae Genus: Touchardia

hona, hōpue [hopue], ōpuhe [opuhe]

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Description

Key Characters:

Growth Form: Shrubs or small trees 2–8 m tall.

Stems: Stems many-branched, glabrous, branches somewhat drooping, bark brown.

Roots:

Leaves: Leaves simple. Alternate. Blades deltate–lanceolate to ovate or narrowly elliptic–lanceolate, (9.5–)15–36 cm long, (3.2–)4–12 cm wide. Apex acuminate, sometimes bluntly acute. Base broadly cuneate to truncate, sometimes (hawaiʻi) cuneate. Upper surfaces glabrous; lower surfaces sparsely short–hirtellous along the veins or glabrous, sometimes (Maui and Hawai'i) pubescent over entire surfaces but more densely so along the veins; Leaves dark green, sometimes (Kauaʻi) with red veins and petioles, chartaceous to firm chartaceous, usually palmately, cystoliths usually prominent, especially on lower surface. Margins usually undulate and irregularly crenate, crenate–serrate, or sometimes subentire. Palmately 3-veined, 3-veined at base, with 12–15 additional pairs of pinnate veins. Petioles 2–14.5 cm long, glabrous or short-hirtellous. Stipules membranous, lanceolate and usually somewhat falcate, 3–4.5 cm long, glabrous.

Flowers: Flowers in dichotomous, widely spreading, paniculate cymes arising from the lower leaf axils or at nodes below the leaves. staminate cymes: 3–9 cm long, peduncles 0.5–2.5(–3.5) cm long, each inflorescence branch terminated by a glomerule of 5–15 (–20) sessile flowers; pistillate cymes more delicate than staminate ones, (2.5–)3–9(–11) cm long, peduncles 0.8–3(–4) cm long, each inflorescence branch terminated by a green glomerule of 5–10 subsessile flowers, the interior pair somewhat wider. Flowers unisexual (and the plants dioecious). Calyx iof staminate flowers greenish white, sometimes tinged violet or on Kauaʻi lavender to pink or red, depressed–globose, lobes 4–5, ovate, hooded, 1.7–2.1 mm long, apex ciliate. Calyx of pistillate flowers green, ovoid–ellipsoid, slightly compressed, lobes 4, elliptic–ovate, unequal, concave, 0.6–0.8 mm long, ciliate toward apex. Corolla (petals) absent. Stamens in staminate flowers white or on Kauaʻi purple to pink; filaments incurved in bud, elastically reflexed when pollen is shed; anthers dithecal, opening by longitudinal slits; pistillate cymes: staminodes absent. Ovary superior (pistillate cymes), psuedomonomerous, 1-celled; ovule 1, placentation basal; style 1; stigma large. Staminate flowers with ovoid pistillode, 0.3 mm long; ovary vestigial and sterile.

Fruit: Achenes ovoid; slightly compressed; 1.8–2.2 mm long; completely enclosed by the orange or yellow; enlarged;fleshy calyx 2.5–3 mm long; the surface with prominent cystoliths and scattered transparent glands. Seeds 1 per achene.

Ploidy:

Habitat: Slopes and gulch bottoms in mesic to wet and diverse mesic forests.

Elevation Range: 150–1700 m.

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Notes

  • 150-1700 m
  • Description digitized by Alyssa Larson
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • fibers of the inner bark were formerly used for fishnets and occasionally for kapa cloth

Bibliography

Name Published In: Bishop Mus. Occas. Pap. 163: 243 (2025)

Other References

Wagner et al. 1990:1312 (K, O, Mo, L, M, H [as Urera glabra]); Wells et al. 2021 (COMBNOV, NOMNOV Syn. U. glabra = T. oahuensis); Thomas, Magnacca, et al. 2025:243 (Syn. T. oahuensis [nom. illeg.] = T. sandwicensis)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Touchardia sandwicensis Hanakapiai, Napali Coast Moist woods. Preserved_Specimen 10 ft tall. St.John, H. 10932 Kauai BISH 1930-12-31
2 Touchardia sandwicensis Pia Valley, west branch Local concentration along the streambed, canopy of kukui. Preserved_Specimen Population in flower. Takeuchi, W.N. 1866 Oahu BISH 1984-11-24
3 Touchardia sandwicensis Head of Awaawapuhi Valley, Honopu By stream in deep wet wooded gulch. Preserved_Specimen Tree 8 m x 1.5 dm. Petioles and main veins reddish. St.John, H. 22889 Kauai BISH 1949-12-21
4 Touchardia sandwicensis Olokele Canyon with Pipturus albidus Preserved_Specimen Rock, J.F.C. 5497 Kauai BISH 1909-09-30
5 Touchardia sandwicensis Laupahoehoe Section, Hilo Forest Reserve Growing in a gulch. Associated plants include Metrosideros, Coprosma, and Cheirodendron. Preserved_Specimen Small tree 9 ft. tall. Anderson, S.J. 490 Hawaii BISH 1982-02-23
6 Touchardia sandwicensis East Maui, Mamuuaiuii gulch dry Preserved_Specimen Medeiros, A.C. 187 Maui BISH 1981-10-21
7 Touchardia sandwicensis Nualele [sp?] Valley shrubby rainforest Preserved_Specimen spreading lax 8' trunk Degener, O. 22217 Molokai BISH 1952-03-18
8 Touchardia sandwicensis Waianae Mts. Kamananui, Kaawa Gulch, Moist bottom of gulch, under Aleurites, with Pipturus Preserved_Specimen Tree 7m. x 15cm. fl. tinged with pale-violet. St.John, H. 24842 Oahu BISH 1952-10-18
9 Touchardia sandwicensis Near head of Kaumokuni culch, north side of Mt Kaala Growing in talus at base of very narrow canyon. Associate: Aleurites moluccana Preserved_Specimen Tree 25 ft. tall, 8 in. DBH. Milky sap. Hatheway, W.H. 167 Oahu BISH 1950-08-02
10 Touchardia sandwicensis Puuwaawaa Hot dry open mesic forest. (Average of 25 ft. between trees.) Urera, banana poka, ferns. Preserved_Specimen Tree 18ft. DBH 5 in. Ruffin, W. 22 Hawaii BISH 1971-07-03
11 Touchardia sandwicensis Laupahoehoe section of Hilo Forest Reserve. Growing on a streambank near a waterfall, a feeder stream of Kaiwilahilahi. Open 'ohi'a-koa forest with Cibotium, Cheirodendron, Ilex, Coprosma, and Rubus hawaiensis. Preserved_Specimen Rare tree in this forest. Height 15-20 ft, spread of 20 ft. Flowers pink and lavender. Davis, J. 681 Hawaii BISH 1982-02-02
12 Touchardia sandwicensis Kau District, Kilauea Forest Was located in a 20' deep depression with 4-5 other Urera trees. Scattered Acacia - Metrosideros forest, with a closed Cibotium understory. Preserved_Specimen Tree 20' tall having 4 basal diameter. All trees were in good shape. No other Urera plants were located. Clarke, G. 507 Hawaii BISH 1980-09-25
13 Touchardia sandwicensis Awaawapuhi - Nualolo crossover trail. Nualolo streambed In shade. Preserved_Specimen Hobdy, R.W. 163 Kauai BISH 1969-11-05
14 Touchardia sandwicensis Waianae Mts., Kawaihapai Gulch on steep sloping sides of gulch, in mesic-summer dry forest Preserved_Specimen Small tree with PTBG 71s1715. Herbst, D.R. 2247 Oahu BISH 1971-12-05
15 Touchardia sandwicensis Puuhualalai; Puuwaawaa Woods. Preserved_Specimen Tree 20 ft. x 6 inches. St.John, H. 11398 Hawaii BISH 1931-12-29
16 Touchardia sandwicensis Waimea district Koaie, headwater gulches and stream, well above Koaie river, northwest of Mohihi- Waialae trail Metrosideros transitional mesic to wet forest and riparian vegetation Cheirodendron, Perrottetia sandwicensis, Kadua affinis, Ilex anomala, Syzygium sandwicensis, Dodonaea viscosa, Coprosma waimeae, Cyrtandra kauaiensis, Melicope clusiifolia, Labordia degeneri, Cyanea f Preserved_Specimen herb, terrestrial along stream bank with P. kokeana, P. cookiana, P. remyi, erect stems green with orange-red tinge, common Wood, K.R. 15392 Kauai BISH 2013-02-06
17 Touchardia sandwicensis Kalaupapa, upper Waihanau, upper Mokomoko gulch above Kapuna springs mixed mesic, semi-closed canopy, native forest with Metrosideros polymorpha, Metrosideros waialealae, Alyxia oliviformis, Antidesma platyphylla, Myrsine lessertiana, Perrottetia sandwicensis, Pittosporum argentifolium, Urera glabra, Psychotria, Dicranopteris linearis, Charpentiera obovata, Dodonaea viscosa, Bobea elatior, Polygonum punctatum Preserved_Specimen 4 m shrub; many branched Wysong, M.L. 845 Molokai BISH 2005-12-08
18 Touchardia sandwicensis Lihue, Upper Waikoko headwaters, Lihue-Koloa Forest Reserve Lowland wet ohia forest: Metrosideros polymorpha, Pisonia umbellifera, Polyscias oahuensis, Sphaeropteris cooperi, Angiopteris evecta, Spathodea campanulata, Miconia crenata, Cyrtandra kamoolaensis, C. confertiflora, Psychotr... Preserved_Specimen A single tree, 5 m tall and wide with spreading crown, in the forest mid-story below emergent ohia. In full flower, inflorescences reddish-pink. In large axillary bunches below the clusters of leaves terminating each stem. Flowers appear to be female. A.M. Williams 861 Kauai BISH 2024-03-14
19 Touchardia sandwicensis Kulani region; Puu Makaola, 1.3 miles in Disappointment Road Rainforest. Preserved_Specimen St.John, H. 26947 Hawaii BISH 1980-06-02
20 Touchardia sandwicensis Pia valley, west beanch Under dense Kukui canopy Preserved_Specimen A limby 9' shrub with leaves clustered at the branch tips Takeuchi, W.N. 1858 Oahu BISH 1984-11-24
21 Touchardia sandwicensis South Kona, Honaunau Forest Reserve, near Kealakekua Ranch Scattered through closed Acacia - Metrosideros forest, mixed native understory. Preserved_Specimen Shrub 7'. Davis, J. 255 Hawaii BISH 1980-03-27
22 Touchardia sandwicensis Waianae Mountains: Puu Kaua, south fork of Ekahanui Gulch Woods at head of gulch. Preserved_Specimen Tree 10 meters in height x 2 dm. St.John, H. 18669 Oahu BISH 1938-02-27
23 Touchardia sandwicensis Napali Coast, W. branch of Hanakoa Valley Wet shaded pocket. Preserved_Specimen 3 m shrub. St.John, H. 23196 Kauai BISH 1947-12-31
24 Touchardia sandwicensis Makakupaia, Kupaia Gulch with Metrosideros, Buddleja, Coprosma, Gouldia, Dodonaea; next to stream Preserved_Specimen tree 3 m tall Lau, J. 1457 Molokai BISH 1985-09-24
25 Touchardia sandwicensis Upper Onini Gulch Branches hanging over bank above stream bed. Associated plants: Dissochondrus biflorus, Doodia kunthiana, Microlepia, Euphorbia celastroides, Pisonia, Psychotria, Myrsine lanaiensis, Pipturus, Viola tracheliifolia, Peperomia ligustrina var. oopuolana in open Metrosideros forest. Preserved_Specimen Large shrub 5' tall with 10' branches. Basal diameter 8. Milky watery sap. Sterile. Davis, J. 863 Molokai BISH 1982-05-28
26 Touchardia sandwicensis Napali Cliff Trail, near Makena dark gulch Preserved_Specimen Degener, O. 27463 Kauai BISH 1960-11-25
27 Touchardia sandwicensis Waianae Mountains: Waianaeuka; east ridge Puu Kaala Lower woods. Preserved_Specimen Tree. St.John, H. 9935 Oahu BISH 1929-10-13
28 Touchardia sandwicensis Nahiku, Kuhiwa Trail, Koolau Forest Reserve Moist woods. Preserved_Specimen 9 m tree, flowers green, fruits white. Female flowers. St.John, H. 17907 Maui BISH 1937-01-02
29 Touchardia sandwicensis Kaholuamano, stream bed of Opaiwela Stream bed Preserved_Specimen Rock, J.F.C. 9006 Kauai BISH 1911-10-01
30 Touchardia sandwicensis Kilauea, Bird Park Nat. Park, open woods. Preserved_Specimen 25 ft tree. St.John, H. 11255 Hawaii BISH 1931-12-22
31 Touchardia sandwicensis East Maui, Kipahulu Valley, lower dogleg mesic to wet koa forest, with Perrottetia sandwicensis, Psychotria hawaiiensis, Broussaisia arguta, Hedyotis acuminata Preserved_Specimen shrub ~2 m; pistillate Wagner, W.L. 5913 Maui BISH 1988-03-09
32 Touchardia sandwicensis Kipuka Puaulu (Bird Park), Kilauea Dry, open forest Preserved_Specimen Tree 4m. tall. Fosberg, F.R. 10136 Hawaii BISH 1933-08-30
33 Touchardia sandwicensis Wright Road, Olaa Forest Reserve, Puna District Virgin Cibotium and Metrosideros forest Preserved_Specimen Tree 8 m x 4 dm. Leaf veins red below. St.John, H. 25089 Hawaii BISH 1953-01-29
34 Touchardia sandwicensis Small canyon above Kaala trail, Waianae Mts., Waianaeulea Wet canyon bottom Preserved_Specimen Tree 7 m. tall Fosberg, F.R. 9110 Oahu BISH 1933-01-08
35 Touchardia sandwicensis Waianae Mountains: Waianae Kai; Puu Kaslena Wooded gulch. Preserved_Specimen 25 foot tree. St.John, H. 12974 Oahu BISH 1933-03-12
36 Touchardia sandwicensis Laupahoehoe section of Hilo Forest Reserve, North Hilo District Growing near jeep road in `ohi`a - hapu`u forest with planted exotic trees. Preserved_Specimen Small tree. Corn, C. 329 Hawaii BISH 1981-11-02
37 Touchardia sandwicensis Olokele Canyon with Pipturus albidus Preserved_Specimen Rock, J.F.C. 5496 Kauai BISH 1909-09-30
38 Touchardia sandwicensis Pia Valley, west branch Local concentration along the streambed, canopy of kukui. Preserved_Specimen Population in flower. Takeuchi, W.N. 1866 Oahu BISH 1984-11-24
39 Touchardia sandwicensis Kilauea, Bird Park Nat. Park, open woods. Preserved_Specimen 30 ft tree. St.John, H. 11258 Hawaii BISH 1931-12-22
40 Touchardia sandwicensis Kailua: first main fork north of main S fork of Kahanaiki Stream In gully at base of mountain. Preserved_Specimen Rather rare small tree; sparsely-branched. Stone, B.C. 1167 Oahu BISH 1955-09-26
41 Touchardia sandwicensis Ukumehame Gulch Rain forest, bottom of wet gulch. Preserved_Specimen Tree 6 m tall. St.John, H. 25692 Maui BISH 1955-12-28
42 Touchardia sandwicensis Hoolulu Valley, Haena-Kalalau Trail, Na Pali Kona Coast shady, rocky stream bottom; with Asplenium nidus, Pisonia umbellifera, Cyrtandra, Cordyline terminalis, Aleurites Preserved_Specimen Carlquist, S. 1652 Kauai BISH 1964-12-05
43 Touchardia sandwicensis Nualele [sp?] Valley shrubby rainforest Preserved_Specimen spreading lax 8' trunk Degener, O. 22217 Molokai BISH 1952-03-18
44 Touchardia sandwicensis Waianae Mountains: 3.5 miles SE of Puu Hapapa at head of Kanehoa In valley, on stream bank, moderately dry. Preserved_Specimen Pearsall, G.A. 224 Oahu BISH 1958-10-27
45 Touchardia sandwicensis S. Kona Forest Preserved_Specimen Trees 15-20 ft. high. Leaves about 12x6 cm. Male flowers. Bryan, L.W. Hawaii BISH 1957-02-19
46 Touchardia sandwicensis Makaleha Mts, ridge just SE of Puu Eu, Kealia Forest Reserve Windswept shurbland and patches of lowland wet forest W/ Metrosideros, Dicranopteris, Ilex anomala, Psychotria wawrae, P. mariniana, Vaccinium calycinum, Syzygium sandwicensis, Pneumatopteris, Machaerina angustifolia, Cheirodendron spp, Kadua affinis, Br Preserved_Specimen tree, 4 m tall, well branched, staminate plant, stems light brown, leaves glossy green with red tinge on main nerves, uncommon Wood, K.R. 14535 Kauai BISH 2011-02-24
47 Touchardia sandwicensis Waianae Mountains: Puu Kaala Lower moist woods. Preserved_Specimen Small tree, 20 feet tall. St.John, H. 10644 Oahu BISH 1930-11-30
48 Touchardia sandwicensis Kau District, Kilauea Forest Scattered Acacia - Metrosideros forest, with a closed Cibotium understory. It was located in a 20' deep depression. Preserved_Specimen Tree 6' tall having 3 basal diameter. Clarke, G. 506 Hawaii BISH 1980-09-25
49 Touchardia sandwicensis N side of Mt. Haupu in shade of Aleurites and Psidium cattleianum Preserved_Specimen Bergau, M. 7 Kauai BISH 1981-10-17
50 Touchardia sandwicensis Upper Awaawapuhi Large but isolated thicket on moist banks along the streambed. Preserved_Specimen Flowering profusely, leaf midveins conspicuously red. Takeuchi, W.N. 3694 Kauai BISH 1987-10-25
51 Touchardia sandwicensis Haupu (Hoary Head Mts) Kipu Moist forest. Preserved_Specimen Bush 10 ft tall. St.John, H. 13651 Kauai BISH 1933-12-25
52 Touchardia sandwicensis Waiapa'a Vegetation: Metrosideros, Xylosma, Dodonaea, Pisonia, Antidesma, Syzygium, Wikstroemia, Leptecophylla, Cinnamomum, Psidium, Morella, Schinus, Tetraplasandra, Nestegis, Diospyros, Psychotria, Pouteria, Freycinetia, Perrottetia, Pipturus, Myrsine, Coprosma, Pleomele, Melicope, Cyrtomium, Adiantum, Blechnum, Selaginella, Sadleria, Doodia. Nephrolepis, Pittosporum, Dicranopteris. Preserved_Specimen Occasional large shrubs or small trees to 7m. Perlman, S. 111228 Lanai BISH 2012-11-28
53 Touchardia sandwicensis S. Kona Forest Preserved_Specimen Trees 15-20ft. high; leaves about 12x6 cm; male flowers. Bryan, L.W. Hawaii BISH 1957-02-19
54 Touchardia sandwicensis Olokele Canyon with Pipturus albidus Preserved_Specimen Rock, J.F.C. 5497 Kauai BISH 1909-09-30
55 Touchardia sandwicensis South Kona, Honaunau Forest Reserve near Kealakekua Ranch Found scattered throughout closed Acacia - Metrosideros forest. Mixed native understory, Coprosma, Myrsine, Pipturus, Psychotria, Cheirodendron, Pelea. Preserved_Specimen Shrub, 6' tall, female flowers pink. Corn, C. Hawaii BISH 1980-03-27
56 Touchardia sandwicensis Kukuiala Valley In moist rainforest, in deep gulches Preserved_Specimen Small tree 4-6 m. tall; flower: yellow green Bryan Jr., E.H. 822 Oahu BISH 1934-10-09
57 Touchardia sandwicensis Anahola River Wet forested slope. Preserved_Specimen Tree 7 m x 20 cm petioles and main veins red, flowers cauliflorous. St.John, H. 26773 Kauai BISH 1971-04-01
58 Touchardia sandwicensis Mt. Kalena Waianae District Preserved_Specimen 8ft. shrub Grant, M.L. 7373 Oahu BISH 1934-09-30
59 Touchardia sandwicensis Waiahole, weedy streambed Preserved_Specimen ten feet highly branched shrub with Cyrtandrae 24242427; only a few plants seen, all sterile, species should be coming into flower within 2-3 months. Takeuchi, W.N. 2423 Oahu BISH 1985-08-31
60 Touchardia sandwicensis East Maui, Hana District, Kipahulu, E part of valley, at base of central pali Preserved_Specimen tree 5 m; milky latex Lamoureux, C.H. 3907 Maui BISH 1967-08-07
61 Touchardia sandwicensis Kikipua Pt. Preserved_Specimen Warshauer, F.R. 3516 Molokai BISH 1981-06-17
62 Touchardia sandwicensis Along Awaawapuhi Stream bed, just below Kokee Road Preserved_Specimen Small shrubby plant with transluscent, pale-yellow fruit. Herbst, D.R. 2238 Kauai BISH 1971-11-10
63 Touchardia sandwicensis Gulch between Kaua and Kanehoa, in localized stand of native canopy; common in are also grows well in weedy communities at lower elevations Preserved_Specimen Most plants fertile Takeuchi, W.N. 2880 Oahu BISH 1986-10-05
64 Touchardia sandwicensis unknown Preserved_Specimen Rémy, J. 200 Oahu BISH 1851-01-01
65 Touchardia sandwicensis Kaholuamanu Preserved_Specimen Rock, J.F.C. 1724 Kauai BISH 1909-03-03
66 Touchardia sandwicensis Kohala, Waiakamali Gulch Preserved_Specimen 5m tall. Warshauer, F.R. 3110 Hawaii BISH 1980-11-30
67 Touchardia sandwicensis East Maui. Station 15, transect 6 Preserved_Specimen 3-4m tall. Warshauer, F.R. 2577 Maui BISH 1980-05-31
68 Touchardia sandwicensis Koolau Mountains, Pia Gulch in Niu Valley. Collections made in left-hand gulch between junction with right-hand gulch and the large waterfall Preserved_Specimen Tree 5 meters tall with whitish-green flowers in spreading clusters at the axils. Sohmer, S.H. 11946 Oahu BISH 1983-11-27
69 Touchardia sandwicensis Near Huliwai; field no. Waianae 535 b Preserved_Specimen 5m. tree in cocl gulch floor Takeuchi, W.N. 2895 Oahu BISH 1986-10-12
70 Touchardia sandwicensis Gulch between Kaua and Kanehoa Also grows well in weedy communities at lower elevations. Preserved_Specimen Takeuchi, W.N. 2880 Oahu BISH 1986-10-05
71 Touchardia sandwicensis 2nd north fork, Kipapa gulch, Waipio, Koolay Mts Preserved_Specimen 15 ft. treesalong woode stream bed. Hosaka, E.Y. 1072 Oahu BISH 1933-07-11
72 Touchardia sandwicensis Kilauea Puaulu Preserved_Specimen Tree 12m high, male (staminate) form. MacDaniels, L.H. 215 Hawaii BISH 1926-11-10
73 Touchardia sandwicensis Along Awaawapuhi Stream bed, just below Kokee Road Preserved_Specimen Small shrubby plant with transluscent, pale-yellow fruit. Herbst, D.R. 2232 Kauai BISH 1971-11-10
74 Touchardia sandwicensis Haiku, top of talus band resting against the Divide, Preserved_Specimen Weak stemmed plant to 4m. height, unusually glabrous in comparison to Waianae populations Takeuchi, W.N. 3019 Oahu BISH 1986-11-11
75 Touchardia sandwicensis Haiku, steep talus apron near the proposed H-3 corridor Preserved_Specimen Fruits yellow-orange Takeuchi, W.N. 3013 Oahu BISH 1986-11-11
76 Touchardia sandwicensis Maunalui [sp?] Preserved_Specimen female Rock, J.F.C. 6145 Molokai BISH 1910-03-21
77 Touchardia sandwicensis North of Puuwaawaa, Waihou Preserved_Specimen Male. Rock, J.F.C. 3786 Hawaii BISH 1909-06-18
78 Touchardia sandwicensis Hiulani forest, W. of Olaa Preserved_Specimen Male plant. Skottsberg, C.J.F. 446 Hawaii BISH 1922-09-08
79 Touchardia sandwicensis Haiku, top of talus bank resting against the divide, Preserved_Specimen Weeak stemmed plant to 4m. hight, unusually glabrous incomparison to Waianae population. Takeuchi, W.N. 3019 Oahu BISH 1986-11-11
80 Touchardia sandwicensis Forest of Waihoa, north of Puuwaawaa Preserved_Specimen Male. Rock, J.F.C. 3792 Hawaii BISH 1909-06-18
81 Touchardia sandwicensis Waihou forest. Puuwaawaa, north Kona Preserved_Specimen Male. Rock, J.F.C. 3785 Hawaii BISH 1909-06-18
82 Touchardia sandwicensis Kaluaa gulch; Field no. Waianae_165a Preserved_Specimen 7-8ft. female ind. streambedf Takeuchi, W.N. 1841 Oahu BISH 1984-11-18
83 Touchardia sandwicensis Waianae Mts. Kupehau gulch beyond Pipeline, Honouliuli,in woods abundant Preserved_Specimen Tree 7m.x5cm. St.John, H. 22260 Oahu BISH 1946-10-13
84 Touchardia sandwicensis Hawaii National Park; S.W. wall, Makaopuhi Preserved_Specimen Tree 15 ft high; leaves with green veins. Fagerlund, G.O. 584 Hawaii BISH 1943-05-25
85 Touchardia sandwicensis Near Puu Hapapa, vicinity of Alectryon 2518 Dry streambed. Common along drainage channels. Preserved_Specimen Takeuchi, W.N. 2516 Oahu BISH 1985-10-13
86 Touchardia sandwicensis Mt.Kalena; Waianae district Preserved_Specimen 8ft. shrub Grant, M.L. 7373 Oahu BISH 1934-09-30
87 Touchardia sandwicensis Kapapaka, Kau Preserved_Specimen Trees 15 ft tall Forbes, C.N. 404 Hawaii BISH 1911-08-08
88 Touchardia sandwicensis Kaholuamauo Preserved_Specimen Male plant. Rock, J.F.C. 5240 Kauai BISH 1909-08-01
89 Touchardia sandwicensis Kiluaea, Kipuka Puaulu Preserved_Specimen Tree 8 m high; leaves small; female (pistillate) form. MacDaniels, L.H. 216 Hawaii BISH 1926-11-10
90 Touchardia sandwicensis West branch, east Makaleha stream, in small side gulch. Mokuleia, Waianae range Preserved_Specimen tree 10ft. tall, 2in. DBH Hatheway, W.H. 401 Oahu BISH 1950-10-04
91 Touchardia sandwicensis Kohala Preserved_Specimen Male. Rock, J.F.C. 8369 Hawaii BISH 1910-06-01
92 Touchardia sandwicensis 17 miles from Kohala along main road toward Waimea. In forest mauka of road Preserved_Specimen Male plant. Degener, O. 8711 Hawaii BISH 1926-08-14
93 Touchardia sandwicensis Haiku, steep talus apron near the proposed H-3 corridor Preserved_Specimen flowers whith an atupical pinkish cast growing nest to urera 3013 Takeuchi, W.N. 3018 Oahu BISH 1986-11-11
94 Touchardia sandwicensis Kauai Preserved_Specimen female Rock, J.F.C. 5494 Kauai BISH 1909-09-30
95 Touchardia sandwicensis near Huliwai; Field no. Waianae 535c Preserved_Specimen 5m tree in cool gulch floor Takeuchi, W.N. 2895 Oahu BISH 1986-10-12
96 Touchardia sandwicensis South Kona district; Kiilai forest Preserved_Specimen Tree 20 ft.high;lvs.pale green; male flowers pinkish-red, petals pale-lilac. Rock, J.F.C. 25709 Hawaii BISH 1957-03-01
97 Touchardia sandwicensis East Maui, Hana District, Waihoi, base of N wall of valley ~25' up side of bank with Eupatorium adenopherum, Antidesma and Athyrium Preserved_Specimen Harrison, B. 67 Maui BISH 1972-07-14
98 Touchardia sandwicensis Nuuanu Pali: east ascent from lookout, Lanihuli Ridge to 2000 ft altitude Preserved_Specimen Sterile specimen. Petioles and veins, and stipules, reddish. Stone, B.C. 2786 Oahu BISH 1959-05-03
99 Touchardia sandwicensis [Hawaiian Islands; U.S.A.; O`ahu; Hawai`i; Honolulu] Preserved_Specimen Rémy, J. 200 Oahu BISH 1855-01-01
100 Touchardia sandwicensis Olokele Canyon Preserved_Specimen female Rock, J.F.C. 5490 Kauai BISH 1909-09-30
101 Touchardia sandwicensis Hawaii Preserved_Specimen Tree, 30 ft tall, extraordinary size, diameter 2 ft. Judd Jr., A.F. 1192 Hawaii BISH 1923-08-06
102 Touchardia sandwicensis Hawaii Fern forest. Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-22
103 Touchardia sandwicensis Olokele Canyon Preserved_Specimen female Rock, J.F.C. 5494 Kauai BISH 1909-09-30
104 Touchardia sandwicensis East Maui, Kaopilopilo and Kapakahawai, S slope of Haleakala in a gulch Preserved_Specimen 12 ft tree, wide branching, berries orange, leaves pubsecent below. Female flowers. Forbes, C.N. 1945 Maui BISH 1920-03-12
105 Touchardia sandwicensis Papaia valley, 6th valley east ofMakaliha valley, Waianae Mts, Waialua wooded slope Preserved_Specimen Judd Jr., A.F. Oahu BISH 1931-12-02
106 Touchardia sandwicensis Kaholuamano, stream bed of Opaiwela Stream bed. Preserved_Specimen Rock, J.F.C. 9006 Kauai BISH 1911-10-01
107 Touchardia sandwicensis Kaluaa gulch Preserved_Specimen Takeuchi, W.N. 1842 Oahu BISH 1984-12-18
108 Touchardia sandwicensis Haiku, steep talus apron near the proposed H-3 corridor Preserved_Specimen fruits yellow-orange; Takeuchi, W.N. 3013 Oahu BISH 1986-12-11
109 Touchardia sandwicensis East Maui, Koolau Forest Reserve deeply cut, shaded streambank, with Perrottetia, Dicranopteris, Athyrium, Dryopteris, Tetraplasandra dipyrena Preserved_Specimen Harrison, B. 347 Maui BISH 1973-07-22
110 Touchardia sandwicensis Second gulch east of Puu Kaupakuhale; Mokuleia, Puu Kaala, Waianae Mts Preserved_Specimen Small tree, 8m. tall, female. Fosberg, F.R. 8903 Oahu BISH 1932-10-23
111 Touchardia sandwicensis Hawaii Volcanoes National Park, Olaa Tract, Wright Road Rainforest. Preserved_Specimen St.John, H. 26951 Hawaii BISH 1980-06-02
112 Touchardia sandwicensis Head of Waikolu Valley, near Hanalilolilo Preserved_Specimen Tree 20 ft high. Lamoureux, C.H. 47 Molokai BISH 1953-12-24
113 Touchardia sandwicensis East Maui, Koolau Forest Reserve deeply cut, shaded streambank, with Perrottetia, Dicranopteris, Athyrium, Dryopteris, Tetraplasandra dipyrena Preserved_Specimen Harrison, B. 387 Maui BISH 1973-07-29
114 Touchardia sandwicensis Kilauea, Kipuka Puaulu Preserved_Specimen Tree 10m high; female (pistillate) form. MacDaniels, L.H. 214 Hawaii BISH 1926-11-10
115 Touchardia sandwicensis Ekahanui; field no. Waianae 517 a Preserved_Specimen Tall shrub in stream bed; with Nothocestrum longifolium, phullostegia mollis, and garnotiana Takeuchi, W.N. 2875 Oahu BISH 1986-10-05
116 Touchardia sandwicensis Waikolu Valley; steep valley wall below Puu Kaeo Wet woods. Preserved_Specimen St.John, H. 23463 Molokai BISH 1948-12-22
117 Touchardia sandwicensis Kilauea Preserved_Specimen Rock, J.F.C. Hawaii BISH
118 Touchardia sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 45 Lanai BISH 1913-10-01
119 Touchardia sandwicensis Na Pali Trail, half way between Hanakapiai and Haena Preserved_Specimen Inflorescence open, fls light lavender. Judd, C.S. Kauai BISH 1938-12-21
120 Touchardia sandwicensis Hawaii Preserved_Specimen Female inflorescence Orley, H. Hawaii BISH 1934-06-01
121 Touchardia sandwicensis Puuaulu, Kilauea Volcano Preserved_Specimen Male plant. Rock, J.F.C. Hawaii BISH 1911-05-01
122 Touchardia sandwicensis Kaeaune Gap, Haleakala Crater Preserved_Specimen Forbes, C.N. 1042 Maui BISH 1919-08-03
123 Touchardia sandwicensis Hawaii Preserved_Specimen Male inflorescence Morley, H. Hawaii BISH 1934-06-01
124 Touchardia sandwicensis Kilauea Volcano Kuala Preserved_Specimen Female plant. Rock, J.F.C. Hawaii BISH 1911-07-01
125 Touchardia sandwicensis Puaulu, Kilauea Vocano Preserved_Specimen Female plant. Rock, J.F.C. Hawaii BISH 1911-07-01
126 Touchardia sandwicensis East Maui, Upper Shelf of Kipahulu Valley just below USFWS Tr #16 stn 6 on north facing slope w/ Akala and Ulu Preserved_Specimen Welton, P. 1914 Maui BISH 1997-04-08
127 Touchardia sandwicensis E Maui betw Transect 27-28 Preserved_Specimen Caespetose. Warshauer, F.R. 2689 Maui BISH 1980-06-24
128 Touchardia sandwicensis Kilauea, Kipua Puaulu Preserved_Specimen Skottsberg, C.J.F. 526 Hawaii BISH 1922-09-15
129 Touchardia sandwicensis Streambed of Kaluaa gulch Preserved_Specimen 7-8ft. female ind. Takeuchi, W.N. 1841 Oahu BISH 1984-11-18
130 Touchardia sandwicensis Olokele Canyon Preserved_Specimen female Rock, J.F.C. 5494 Kauai BISH 1909-09-30
131 Touchardia sandwicensis Kaeauue (?) Gap, Halehaku, Crater of Haleakala Preserved_Specimen Veins of leaves red, bracts burnt orange. Female flowers. Forbes, C.N. 1015 Maui BISH 1919-08-02
132 Touchardia sandwicensis Kamoku, Kaiholena Preserved_Specimen Munro, G.C. 242 Lanai BISH 1927-11-10
133 Touchardia sandwicensis unknown Preserved_Specimen Rémy, J. 202 Oahu BISH 1851-01-01
134 Touchardia sandwicensis Along watercourses, Mts of Kohala Preserved_Specimen Male plant. Rock, J.F.C. 8369 Hawaii BISH 1910-06-01
135 Touchardia sandwicensis Hanalei Preserved_Specimen Mann, H. 398 Kauai BISH 1864-01-01
136 Touchardia sandwicensis 2nd north fork, Kipapa gulch, Waipio, koolau Mts Preserved_Specimen 15ft. tree, along wooded stream bed Hosaka, E.Y. 1072 Oahu BISH 1933-07-11
137 Touchardia sandwicensis Punaluu; Field no. Koolau_374, PRC 880 Preserved_Specimen 8ft. ind. in streambed crossed by Castle Trail Takeuchi, W.N. 1804 Oahu BISH 1984-10-28
138 Touchardia sandwicensis Near Huliwai, cool gulch floor; growing beside Urera 2895 Preserved_Specimen Takeuchi, W.N. 2896 Oahu BISH 1986-10-12
139 Touchardia sandwicensis Wailupe Valley Preserved_Specimen Meinecke, W.H. 4 Oahu BISH 1927-01-09
140 Touchardia sandwicensis unknown Preserved_Specimen Male plant. Rock, J.F.C. BISH
141 Touchardia sandwicensis Kaholuamanoa Preserved_Specimen Forbes, C.N. 388 Kauai BISH 1909-09-01
142 Touchardia sandwicensis Puna District, Olaa Forest Reserve; Wright Rd., 1.7 miles north of junction with Laukapu Ave Preserved_Specimen Ozaki, E.T. 1293 Hawaii BISH 1954-12-22
143 Touchardia sandwicensis Mapulehu Valley Preserved_Specimen Forbes, C.N. 596 Molokai BISH 1912-09-01
144 Touchardia sandwicensis N.Kona, Puuwaawaa, Poohohoo, at water tank Preserved_Specimen Kondo, Y. 88 Hawaii BISH 1946-07-21
145 Touchardia sandwicensis Mapulehu-Punaula Ridge Preserved_Specimen Usinger, R.L. 1 Molokai BISH 1936-08-01
146 Touchardia sandwicensis S Kohala Distr, Puukapu ahu. Kohala Mtns, Kehena DitchTransect 79 betw s-21 & 22 Preserved_Specimen Warshauer, F.R. 2476 Hawaii BISH 1979-07-31
147 Touchardia sandwicensis Waianae Mts., Kupehau Gulch Preserved_Specimen Cowan, R.S. 158 Oahu BISH 1946-10-13
148 Touchardia sandwicensis East Maui, Hana, Kahikinui, S Haleakala, 0.1-0.5 km W of Manawainui Gulch. Transect 25 ext. trail Preserved_Specimen Higashino, P.K. 9235 Maui BISH 1980-06-20
149 Touchardia sandwicensis Haupu range, right side of Kipu Kai Gap Preserved_Specimen Forbes, C.N. 741 Kauai BISH 1916-11-03
150 Touchardia sandwicensis Waialae Iki Valley Preserved_Specimen Lyon, H.L. 28 Oahu BISH 1925-01-02
151 Touchardia sandwicensis Ainahou valley on W side of Koolau Gap, Haleakala Crater Preserved_Specimen Gillett, G.W. 1920 Maui BISH 1966-07-05
152 Touchardia sandwicensis Kaluaa gulch Preserved_Specimen Takeuchi, W.N. 1842 Oahu BISH 1984-12-18
153 Touchardia sandwicensis E of Kaholuamano, along stream leading to Opaiwela Preserved_Specimen Rock, J.F.C. 5238 Kauai BISH 1909-09-04
154 Touchardia sandwicensis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2605 Kauai BISH 1895-09-10
155 Touchardia sandwicensis East Maui W of Transect 27, ext trail Preserved_Specimen Higashino, P.K. 9273 Maui BISH 1980-06-24
156 Touchardia sandwicensis Kaumalumalu, Kona District along jeep trail Preserved_Specimen Sohmer, S.H. 6483 Hawaii BISH 1970-07-02
157 Touchardia sandwicensis Kipuka Puaulu (Puu a Ulu) Preserved_Specimen Forbes, C.N. 1027 Hawaii BISH 1915-07-01
158 Touchardia sandwicensis Hii Mts Preserved_Specimen Forbes, C.N. 621 Kauai BISH 1916-10-20
159 Touchardia sandwicensis Mts. near Koele Preserved_Specimen Forbes, C.N. 128 Lanai BISH 1918-06-01
160 Touchardia sandwicensis Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 13049 Hawaii BISH 1917-08-01
161 Touchardia sandwicensis Kaiholena Preserved_Specimen Munro, G.C. 135 Lanai BISH 1913-10-01
162 Touchardia sandwicensis Olokele Preserved_Specimen Rock, J.F.C. 5496 Kauai BISH 1909-09-30
163 Touchardia sandwicensis E of Kaholuamano, along stream leading to Opaiwela Preserved_Specimen Rock, J.F.C. 5236 Kauai BISH 1909-09-04
164 Touchardia sandwicensis Waiahole; recollection from 2423 (Koolau 743) Preserved_Specimen Takeuchi, W.N. 2520 Oahu BISH 1985-10-26
165 Touchardia sandwicensis E of Kaholuamano, along stream leading to Opaiwela Preserved_Specimen Rock, J.F.C. 5237 Kauai BISH 1909-09-04
166 Touchardia sandwicensis East Maui, woods near Ukulele, above Olinda Preserved_Specimen Forbes, C.N. 189 Maui BISH 1910-07-01
167 Touchardia sandwicensis E of Kaholuamano, along stream leading to Opaiwela Preserved_Specimen Rock, J.F.C. 5243 Kauai BISH 1909-09-04
168 Touchardia sandwicensis Upper Waiakea Forest Reserve; Trail to 358-20 Preserved_Specimen Adee, K.T. Hawaii BISH
169 Touchardia sandwicensis Hawaii Volcanoes National Park Preserved_Specimen Warshauer, F.R. Hawaii BISH 1976-08-03
170 Touchardia sandwicensis Kauai Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
171 Touchardia sandwicensis Makaha Valley, Kooa Range Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
172 Touchardia sandwicensis Puuawaawaa Preserved_Specimen Munro, G.C. Hawaii BISH 1923-06-07
173 Touchardia sandwicensis Waianae Mountains: Mokuleia; NE slope of Puu Kaala Preserved_Specimen Swezey, O.H. Oahu BISH 1932-10-23
174 Touchardia sandwicensis Koolau Range, Waikane-Schofield Trail Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
175 Touchardia sandwicensis Kilauea Preserved_Specimen Rock, J.F.C. Hawaii BISH
176 Touchardia sandwicensis Mauna Loa, Puaulu Preserved_Specimen Rock, J.F.C. Hawaii BISH 1911-08-01
177 Touchardia sandwicensis Nuuanu Preserved_Specimen (Not on sheet) Oahu BISH 1985-01-01
178 Touchardia sandwicensis Waianae Mts., Palehua, in Valley Preserved_Specimen Student at University of Hawaii Oahu BISH 1931-11-22
179 Touchardia sandwicensis Hoolulu, next to stream by trail to Kalalau Preserved_Specimen Corn, C. Kauai BISH 1980-04-10
180 Touchardia sandwicensis Makaha Valley, Koola Range Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
181 Touchardia sandwicensis Wet forest on ridge ro kaala Mt Preserved_Specimen Furmidge, G. Oahu BISH 1933-01-08
182 Touchardia sandwicensis Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
183 Touchardia sandwicensis Bird Park, Kilauea, Kau Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-08-09
184 Touchardia sandwicensis Near Glenwood Preserved_Specimen Rock, J.F.C. Hawaii BISH 1918-07-01
185 Touchardia sandwicensis Manuka Kau Preserved_Specimen Russ, G.W. Hawaii BISH
186 Touchardia sandwicensis [Hawaiian Islands; U.S.A.; O`ahu; Hawai`i; Honolulu] Preserved_Specimen Lay, G.T. Oahu BISH 1826-01-01
187 Touchardia sandwicensis Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
188 Touchardia sandwicensis Kauai Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
189 Touchardia sandwicensis Upper Waiakea Forest Reserve; Trail to 358-20 Preserved_Specimen Adee, K.T. Hawaii BISH
190 Touchardia sandwicensis Konahuanui, Oahu Preserved_Specimen Mann, H. Oahu BISH
191 Touchardia sandwicensis Kaumalumalu, Kona District along jeep trail Preserved_Specimen Sohmer, S.H. 6483 Hawaii BISH 1970-07-02
192 Touchardia sandwicensis Makaleha Valley Preserved_Specimen Rock, J.F.C. 17047 Oahu BISH 1918-05-01
193 Touchardia sandwicensis Transect 78, 1/4mi E of S-1[?] Preserved_Specimen Warshauer, F.R. 2464 Hawaii BISH 1979-07-30
194 Touchardia sandwicensis S Kula slope of Haleakala Preserved_Specimen Forbes, C.N. 2168 Maui BISH 1920-04-07
195 Touchardia sandwicensis unknown Preserved_Specimen Rock, J.F.C. BISH
196 Touchardia sandwicensis Waianae Mountains: Puu Kalena; Waianae Kai Preserved_Specimen Suehiro, A. 65 Oahu BISH 1933-03-12
197 Touchardia sandwicensis Kaiholena, bottom Preserved_Specimen Hobdy, R.W. 236 Lanai BISH 1971-06-07
198 Touchardia sandwicensis Hoalulu, Napali coast. Kalalau Trail Preserved_Specimen Hobdy, R.W. 179 Kauai BISH 1970-04-09
199 Touchardia sandwicensis unknown Preserved_Specimen Rock, J.F.C. BISH
200 Touchardia sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
201 Touchardia sandwicensis unknown Preserved_Specimen Rock, J.F.C. BISH
202 Touchardia sandwicensis Near the base of Hualalai In a dense forest. Preserved_Specimen Tree of considerable size. Rock, J.F.C. 3890 Hawaii BISH 1909-06-18
203 Touchardia sandwicensis Makaha Valley, waianae Mts Moist forest Preserved_Specimen Tree9m.x2cnm; berries yellow; dark sample St.John, H. 24857 Oahu BISH 1952-11-30
204 Touchardia sandwicensis near Puu O Wahaula rainforest Preserved_Specimen sap watery milky Degener, O. 8704 Molokai BISH 1928-04-17
205 Touchardia sandwicensis Olaa Tract, along Wright Road Metrosideros, Cibotium forest Preserved_Specimen 20' x 3.5 tree; veins purple on undersides of leaves; young stem red; peduncle red; corolla pink; sap milky-watery Nagata, K.M. 1888 Hawaii BISH 1979-06-02
206 Touchardia sandwicensis Makaleha Mts, ridge just SE of Puu Eu, Kealia Forest Reserve Windswept shurbland and patches of lowland wet forest W/ Metrosideros, Dicranopteris, Ilex anomala, Psychotria wawrae, P. mariniana, Vaccinium calycinum, Syzygium sandwicensis, Pneumatopteris, Machaerina angustifolia, Cheirodendron spp, Kadua affinis, Br Preserved_Specimen tree, 4 m tall, well branched, staminate plant, stems light brown, leaves glossy green with red tinge on main nerves, uncommon Wood, K.R. 14535 Kauai BISH 2011-02-24
207 Touchardia sandwicensis Honaunau Forest Reserve A transitional ohia - koa forest with Perrottetia sandwicensis, Clermontia coerulea, Stenogyne macrantha, Psychotria, Gouldia, Alyxia, Ilex anomala, Pittosporum hosmeri, Nothocestrum, Vaccinium calycinum. Preserved_Specimen Occasional tree. Clarke, G. 311 Hawaii BISH 1980-03-28
208 Touchardia sandwicensis Boundary Kipuda, East Rift, Hawaii Volcanoes National Park Plants collected in Kipuka and Cinder Cone area undisturbed by pigs. Closed Metrosideros, subcanopy native trees understory consisting of tree ferns and native shrubs. Preserved_Specimen Higashino, P.K. Hawaii BISH 1982-10-20
209 Touchardia sandwicensis Waianae Mountains: Puu Hapapa In forest. Preserved_Specimen Hosaka, E.Y. 318 Oahu BISH 1930-10-19
210 Touchardia sandwicensis Manuka NAR, Olopua Iki Kipuka mesic forest; Acacia koa, Metrosideros, Nestegis sandwicensis, Myrsine lanaiensis, Coprosma cymosa, Dodonaea viscosa, Pouteria sandwicensis, Claoxylon sandwicensis, Wikstroemia oahuensis, Cheirodendron trigynum, Psychotria marinianum, Pipturus albidus Pittosporum, Psydrax odorata, Pisonia, Urera glabra, Psidium guajava, Passiflora Preserved_Specimen 15' tree; flowering Perlman, S. 20517 Hawaii BISH 2007-07-12
211 Touchardia sandwicensis Kukuiala Valley; in deep gulch in moist rainforest Preserved_Specimen small tree hight 4-6m. Bryan Jr., E.H. 822 Oahu BISH 1934-10-09
212 Touchardia sandwicensis East Maui, Koolau Forest Reserve deeply cut, shaded streambank, with Perrottetia, Dicranopteris, Athyrium, Dryopteris, Tetraplasandra dipyrena Preserved_Specimen Harrison, B. 347 Maui BISH 1973-07-22
213 Touchardia sandwicensis Honaunau Forest Reserve A transitional ohia-koa forest with Perrottetia sandwicensis, Clermontia coerulea, Stenogyne macrantha, Psychotria, Gouldia, Alyxia, Ilex anomala, Pittosporum hosmeri, Nothocestrum, Vaccinium calycinium. Preserved_Specimen Occasional trees. Clarke, G. 5 Hawaii BISH 1980-03-28
214 Touchardia sandwicensis S. Kona Forest Preserved_Specimen Trees 15-20 ft. high. Leaves 12x6 cm, male flowers. Bryan, L.W. Hawaii BISH 1957-02-19
215 Touchardia sandwicensis Kalalau; Kilohana Lookout Mountain Woods. Preserved_Specimen 7 m. tree; leaves thick firm chartaceous; above is dark shiny and green. St.John, H. 19984 Kauai BISH 1913-02-11
216 Touchardia sandwicensis Waianae Mtn, Waianae Kai, Makaha valley side, in rich area, belwo Kamaileunu ridge In Oahu diverse mesic forest with Alyxia, Nestegis, Hibiscus, Alectryon, Diospyros, Antidesma, Schinus, Tetraplasandra, Aleurites, Pleomele, Claoxylon, Urera, Nothocestrum longiflorum, Zanthoxylum, Embelia, Pisonia, Bobea, Psychotria, Scaevola, Melicope, Astelia, Labordia, Perrottetia. Preserved_Specimen Perlman, S. 7050 Oahu BISH 1987-12-06
217 Touchardia sandwicensis upper Awaawapuhi valley At top of stream just below road. Emerging from thick Hedychium cover along stram. Under 40% canopy cover of Metrosideros polymorpha, Perrottetia and Coprosma. Athyrium ferns in understory. Preserved_Specimen Large shrub 5 m tall. Canopy 6 m across. Lvs to 30 cm long with conspicuous red venetation. Newly flushed lvs reddish-pink. Trauernicht, C. 749 Kauai BISH 2009-02-19
218 Touchardia sandwicensis Second gulch east of Puu Kaupakuhale Mts Wet forest at head of gulch Preserved_Specimen Small tree 6m tall, male. Most of inflorescences fallen. Fosberg, F.R. 8904 Oahu BISH 1932-10-23
219 Touchardia sandwicensis Kaluaa gulch Occasional along sides of gulch, on the streambed Preserved_Specimen most plants flowering, with watery milky sap. Takeuchi, W.N. 1842 Oahu BISH 1984-11-18
220 Touchardia sandwicensis Wailupe Str. E branch; a long streambed along streambed Preserved_Specimen a common species, populations flowering Takeuchi, W.N. 1949 Oahu BISH 1985-01-27
221 Touchardia sandwicensis Hawaii Volcanoes National Park, Olaa Tract, Wright Road Rainforest. Preserved_Specimen St.John, H. 26951 Hawaii BISH 1980-06-02
222 Touchardia sandwicensis N. of Puuawaawaa, Waihou Dense forest Preserved_Specimen Male plant; about 40ft. tall and 12-14 in. thick. Rock, J.F.C. 3790 Hawaii BISH 1909-06-18
223 Touchardia sandwicensis Olaa Tract, Wright Road Rainforest. Preserved_Specimen St.John, H. 26950 Hawaii BISH 1980-06-02
224 Touchardia sandwicensis Keauhou Kona, Mauka Found in association with kopiko, puahanui, ohia, & `ie`ie. Preserved_Specimen Tree-like shrub to 15 ft tall, gray-brown bark with prominent leaf scars, ripe fruit is yellow-orange. Rozett, J.R. 1 Hawaii BISH 1981-12-11
225 Touchardia sandwicensis Kupaia Gulch, Kamakou Preserve bottom; with Coprosma, Metrosideros, Nestegis, Buddleia, Polygonum Preserved_Specimen tree 7'; flowering profusely Imada, C. 108 Molokai BISH 1985-09-24
226 Touchardia sandwicensis East Maui, Mamuuaiuii gulch dry Preserved_Specimen Medeiros, A.C. 187 Maui BISH 1981-10-21
227 Touchardia sandwicensis Headwaters of S fork, Kaunakakai Gulch North of jeep trail to Pepeopae bog. Growing on bank of small stream, ca. 1000' upstream of tunnel. Growing with Nothocestrum sp., Gouldia, Scaevola, Coprosma, Cibotium, and Sticherus in closed Metrosideros forest. Preserved_Specimen Large spreading shrub about 12' tall; sterile. Watery milky sap. Davis, J. 876 Molokai BISH 1982-05-29
228 Touchardia sandwicensis Waiakea, back of Hilo Forest. Preserved_Specimen Rock, J.F.C. Hawaii BISH 1955-12-01
229 Touchardia sandwicensis Limahuli Valley. Along trail up valley on eastern side of stream Along river in Kukui grove. Preserved_Specimen Single shrub of 6 ft. Flynn, T.W. 1420 Kauai BISH 1985-12-13
230 Touchardia sandwicensis 3 mi NW of Hawaii National Park Fern forest Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-22
231 Touchardia sandwicensis Kalalau trail, 2nd gulch from E stream banks and seeps, with Cyrtandra, Pisonia, Charpentiera. Preserved_Specimen 12' tall Christensen, C. 62 Kauai BISH 1976-09-25
232 Touchardia sandwicensis Wailupe str. east reanch Dominated canopy with large population of Athyrium macraei in understory Preserved_Specimen an 8ft. ind. on slippery slopes above flowing stream Takeuchi, W.N. 1959 Oahu BISH 1985-01-27
233 Touchardia sandwicensis N.of Puuwaawaa, Waihou Forest. Preserved_Specimen Male. Rock, J.F.C. 3817 Hawaii BISH 1909-06-18
234 Touchardia sandwicensis East Maui, Haleakala, Koolau Gap rainforest Preserved_Specimen tree of 6 m; flower light-purple; fruit yellow Wilbur, R.L. 943 Maui BISH 1948-07-02
235 Touchardia sandwicensis Honaunau Forest Reserve A transitional ohia - koa forest with Perrottetia sandwicensis, Clermontia coerulea, Stenogyne macrantha, Psychotria, Gouldia, Alyxia, Ilex anomala, Pittosporum hosmeri, Nothocestrum, Vaccinium calycinum. Preserved_Specimen Tree. Clarke, G. 5 Hawaii BISH 1980-03-28
236 Touchardia sandwicensis head of Waikolu Valley, Hanalilolilo moist woods on headwall Preserved_Specimen 20' tree St.John, H. 12367 Molokai BISH 1932-12-21
237 Touchardia sandwicensis Kaapilopilo & Kapakahawai In a gulch. Preserved_Specimen Wide branching tree 3.66 m high; berries orange; lvs pubescent below. Forbes, C.N. 1945 Maui BISH 1920-03-12
238 Touchardia sandwicensis Olokele Canyon with Pipturus albidus Preserved_Specimen Rock, J.F.C. 5487 Kauai BISH 1909-09-30
239 Touchardia sandwicensis Kalalau trail, 2nd gulch from E stream banks and seeps, with Cyrtandra, Pisonia, Charpentiera. Preserved_Specimen 12' tall Christensen, C. 62 Kauai BISH 1976-09-25
240 Touchardia sandwicensis Waianae Mountains: Waianaeuka; east ridge Puu Kaala Lower woods. Preserved_Specimen St.John, H. 9936 Oahu BISH 1929-10-13
241 Touchardia sandwicensis S.Kona, Honaunau Forest Reserve near Kealakekua ranch Scattered through closed Acacia-Metrosideros forest, mixed native understory Preserved_Specimen Shrub 7' tall. Davis, J. 255 Hawaii BISH 1980-03-27
242 Touchardia sandwicensis Kauai Waimea District. Waialae, headwater strealet just south of Kaluahaulu Ridge Trail. Acacia-Metrosideros montane mesic forest with Alphitonia ponderosa, Pouteria sandwicensis, Diospyros sandwicensis, Santalum pyrularum, Chrysodracon aurea, Pittosporum glabrum, Cheirodendron trigynum, Ilex anomala, Wikstroemia oahuensis, Claox Preserved_Specimen Tree, 5 meters tall, well-branched, stems gray-brown, in flower, inflorescense pink-red, small colony along stream near spring. Wood, K.R. 17524 Kauai BISH 2017-09-05
243 Touchardia sandwicensis W Maui, Haelaau forest, side of ridge Preserved_Specimen Ewart III, G.R. 31 Maui BISH 1928-12-19
244 Touchardia sandwicensis Kaumokunui Gulch, North side of Mt. Kaala Moist open forest of steep talus slope. Preserved_Specimen Tree 25 ft. tall, 8 in. DBH. Hatheway, W.H. 315 Oahu BISH 1950-08-02
245 Touchardia sandwicensis Kapuna Gulch Below the Mokuleia trail in a gulch with large Pisonia umbellifera; Alectryon mahoe; Diplazium sandwichianum; and Pipturus albidus. Preserved_Specimen Welton, P. 399 Oahu BISH 1990-04-07
246 Touchardia sandwicensis Waimea District. Honopu, northern slopes and drainage below Kainamanu. Acacia koa-Metrosideros polymorpha-Dicranopteris linearis montane mesic forest with Bobea brevipes, Ilex anomala, Wikstroemia furcata, Claoxylon sandwicensis, Polyscias waimeae, Melicope anisata, Melicope barbigera, Kadua affinis, Psychotria greenwelliae, Planchonella sandwicensis, Brou Preserved_Specimen Tree, 3 meters tall, stems green or brown, male flowers with pink-red tinge, preferring banks of stream, occasional. Wood, K.R. 17145 Kauai BISH 2016-10-18
247 Touchardia sandwicensis Above Huehue Ranch, Puu Hualalai In lava tube sink hole in Ohia-Myrsine-Sadleria fern forest. Preserved_Specimen Heed, W.B. 903 Hawaii BISH 1966-07-01
248 Touchardia sandwicensis South Kona, Honaunau Forest Reserve, near Kealakekua Ranch Scattered throughout closed Acacia - Metrosideros forest, mixed native understory, Coprosma, Myrsine, Pipturus, Psychotria, Cheirodendron, Pelea. Preserved_Specimen Shrub, 6' tall, female flowers pink. Corn, C. 158 Hawaii BISH 1980-03-27
249 Touchardia sandwicensis Mapulehu Valley, trail on ridge E of rainforest Preserved_Specimen 15' tree St.John, H. 12877 Molokai BISH 1932-12-29
250 Touchardia sandwicensis Wailupe Str. E branch; a long streambed along streambed Preserved_Specimen a common species, populations flowering Takeuchi, W.N. 1949 Oahu BISH 1985-01-27
251 Touchardia sandwicensis Wailupe Stream east branch In middle of streambed. Preserved_Specimen 9 foot individual. Takeuchi, W.N. 1957 Oahu BISH 1985-01-27
252 Touchardia sandwicensis Lihue District. Blue Hole, headwaters of Wailua River, below Waialeale and Kawaikini. Metrosideros lowland wet forest and shrubland surrounded by ca. 1000 m cliffs w/ Psychotria mariniana, P. hexandra, Antidesma platyphyllum var. hillebrandii, Pipturus albidus, P. ruber, Urera glabra, Kadua centranthoides, Touchardia latifolia, Dianella sandwicensis, Polyscias o Preserved_Specimen Tree, 3-4 m tall, many-branched, in flower, occasional. Wood, K.R. 18634 Kauai BISH 2021-01-28
253 Touchardia sandwicensis upper Lihue ditch rain forest Preserved_Specimen tree of 6 m MacDaniels, L.H. 843 Kauai BISH 1927-02-21
254 Touchardia sandwicensis East Maui, Hana District, Haleakala, Kipahulu, summit of W ridge of Kaukauai gulch koa woods Preserved_Specimen sparse shrub, 2-3 m tall; flower white St.John, H. 17812 Maui BISH 1936-12-28
255 Touchardia sandwicensis Kaholuamano, stream bed of Opaiwela Stream bed Preserved_Specimen Rock, J.F.C. 9006 Kauai BISH 1911-10-01
256 Touchardia sandwicensis Puu Makaala, Upper Olaa FR, Puna, Disappointment Rd. Metrosideros, Cibotium forest Preserved_Specimen 15' x 2 tree; male flowers pink Nagata, K.M. 1872 Hawaii BISH 1979-06-01
257 Touchardia sandwicensis W Maui, Haelaau rainforest Preserved_Specimen Bryan Jr., E.H. 670 Maui BISH 1928-12-21
258 Touchardia sandwicensis Waianae Mtn, Waianae Kai, S side ridge, Waianae Valley, to S of Hesperomania trail In Acacia koa-Metrosideros lowland mesic forest with Myrsine lessertiana, Melicope, Diospyros, Freycinetia, Scaevola gaudichaudii, Dicranopteris, Labordia, Pipturus, Zanthoxylum kauense, Psidium, Alyxia, Dodonaea, Rauvolfia, Psychotria, Elaeocarpus, Hibiscus. Pisonia, Aleurites, Lantana, Sophora chrysophylla. Preserved_Specimen Perlman, S. 6819 Oahu BISH 1987-11-08
259 Touchardia sandwicensis South Kona, kiilai forest forest Preserved_Specimen female flowers Bryan, L.W. Hawaii BISH 1957-03-31
260 Touchardia sandwicensis Upper Hamakua Ditch Trail, Alakahi stream, Puukapu Moist wooded gulch. Preserved_Specimen 5 m shrub, leaves bright green, buds pink. St.John, H. 25098 Hawaii BISH 1953-02-01
261 Touchardia sandwicensis Hoalulu Stream, Hanakapiai, W of headland Moist gulch in aleurites forest. Preserved_Specimen 15 ft shrub, berries yellow. St.John, H. 13860 Kauai BISH 1933-12-30
262 Touchardia sandwicensis Drive to end of Kulani Prison Road about 3 miles past Kulani Prison (Honor) Camp; or 19 miles from intersection with Kona-Hilo Highway near Hilo Mixed Acacia-Metrosideros-Cibotium community. Preserved_Specimen Shrub about 2 ft tall. Newell, C.L. 500 Hawaii BISH 1966-08-21
263 Touchardia sandwicensis Manuka State Park, South Kona Forest Reserve, Nature Trail. In collapsed lava tube of about 125' diameter. `A`a substrate. Growing with Cyrtandra sp. and Microlepia. Alyxia and Freycinetia also present. Preserved_Specimen Shrub 15' tall, 20' spread, 3 basal diameter. Excellent vigor, fertile. Clarke, G. 692 Hawaii BISH 1982-04-29
264 Touchardia sandwicensis Mokuleia Forest Reserve. Peacock Flats Metrosideros forest. Preserved_Specimen Small tree 20 feet; 3 inches diameter. Little Jr., E.L. 31033 Oahu BISH 1976-08-10
265 Touchardia sandwicensis Hanalei District. Waioli Valley, along first stream to West of main falls, near back of valley, Haleea Forest Reserve Riparian corridor through Lowland Wet Ohia Forest Riparian corridor through Lowland Wet Ohia Forest Touchardia latifolia, Boerhavia grandis, Pipturus kauaiensis, Syzyglum sandwicensis, Metrosideros polymorpha, Psidium guajava, Buddleja asiatica, Andropogon, Cyclosorus cyatheoides. Preserved_Specimen Round-headed tree 7 meters tall, growing on southeastern stream bank. Tree in full flower with cauliflorus inflorescences sprouting all throughtout the crowns interior. Leaves green with lighter green veins, inflorescences qall light green. Female. Williams, A.M. 701 Kauai BISH 2021-08-18
266 Touchardia sandwicensis Waioli valley along stream Preserved_Specimen tree of 5 m MacDaniels, L.H. 912 Kauai BISH 1927-02-27
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