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Melicope spathulata

A.Gray (1854)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Melicope

pilo kea

Description

Key Characters:

Growth Form: Shrubs or shrubby trees 2–4(–12) m tall.

Stems: Trunk up to 20 cm in diameter, branches spreading, leafy toward apex, glabrous or yellowish brown pubescent on new growth and inflorescences.

Roots:

Leaves: Leaves simple. Opposite. Blades spatulate to oblanceolate or elliptic–obovate, rarely elliptic, usually 10–50 cm long, 2.5–20 cm wide. Apex rounded to obtuse, acute, or sometimes acuminate. Base rounded to obtuse or acute. Surfaces glabrous or lower surfaces puberulent, especially on veins, usually paler on lower surface, coriaceous. Margins entire, sometimes slightly revolute. Lateral veins subopposite, ascending, straight ca. ¾ their length, then dividing to unite with the distinct submarginal vein, midrib sulcate on upper surface. Petioles (0–)10–95 mm long. Stipules absent.

Flowers: Flowers 3–5 in axillary cymes, peduncles 0–30 mm long, pedicels 6–30 mm long, bracts ovate–subulate or deltate. Flowers bisexual (perfect), actinomorphic. Calyx of 4 sepals, in imbricated pairs, broadly ovate, 8–10 mm long, subpersistent, ciliolate, otherwise glabrous, or sparsely to densely puberulent. Corolla of 4 petals, white or cream, slightly spreading at anthesis, oblong–ovate or oblong–obovate, 14–21 mm long, slightly imbricate. Stamens 8; filaments white, nearly completely connate into a staminal tube; anthers dithecal, subsessile on lobes of staminal tube, elongate, base subhastate. Ovary superior, 4-lobed, ± puberulent, usually becoming glabrate, carpels connate adaxially throughout their length, placentation axile; ovules 5–8 per carpel, pendulous on short, broad funiculi; style ca. 9 mm long, subclavate; stigmas linear.

Fruit: Fruit subglobose; 20–35 mm in diameter; 4–lobed; glabrous or minutely puberulent; carpels erect; connate throughout; separating at various places during development; adnate to summit; apex rounded. Seeds 5–8 per cell; funicles short; endocarp glabrous; embryo with thin; broad; rounded cotyledons; hypocotyl very short; embedded in white endosperm.

Ploidy:

Habitat: Mesic to wet forest.

Elevation Range: 450–1,420 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: U.S. Expl. Exped., Phan. 1: 352 (1854)

Other References

Wagner et al. 1990:1211 (K, O, M, H [as Platydesma spathulata]); Appelhans et al 2017:132 (COMBNOV, Syn. Platydesma spathulata)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Melicope spathulata A.Gray Punaluu-Kaluanui Ridge, windward side PRESERVED_SPECIMEN MacDaniels, L.H. 382 Oahu BISH 1/1/1927
2 Melicope spathulata A.Gray Waipio-Waiawa Ridge, rain forest PRESERVED_SPECIMEN MacDaniels, L.H. 53 Oahu BISH 10/6/1926
3 Melicope spathulata A.Gray Mokuleia Moderate forest PRESERVED_SPECIMEN Morley, H. s.n. Oahu BISH 2/4/1934
4 Melicope spathulata A.Gray Hanalei District, Koke'e State Park, Kalalau rim W of Pu'u o Kila lookout, just past drainage below lookout Metrosideros polymorpha dominated wet forest PRESERVED_SPECIMEN Wagner, W.L. 6893 Kauai BISH 2/25/2000
5 Melicope spathulata A.Gray off Honopu trail, Kokee State Park in dense shade off Honopu trail. In native Metrosideros forest PRESERVED_SPECIMEN Ishikawa, S. 302 Kauai BISH 8/22/1973
6 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 5343 Kauai BISH 9/1/1909
7 Melicope spathulata A.Gray Poamoho Native rainforest PRESERVED_SPECIMEN Takeuchi, W.N. s.n. Oahu BISH 6/1/1986
8 Melicope spathulata A.Gray Koolau Mts., Moanalua Valley, summit crest ridge PRESERVED_SPECIMEN Perlman, S. 5942 Oahu BISH 6/7/1987
9 Melicope spathulata A.Gray kokee Moist lower forest, steep slope. PRESERVED_SPECIMEN j. f. rock s.n. Kauai BISH
10 Melicope spathulata A.Gray Waikane-Schofield Trail, Koolau Range, Waianae Uka, On wooded divide PRESERVED_SPECIMEN St. John, H. 12125 Oahu BISH 10/16/1932
11 Melicope spathulata A.Gray Glenwood PRESERVED_SPECIMEN Faurie, A. s.n. Hawaii BISH 5/1/1910
12 Melicope spathulata A.Gray Kilauea Forest Reserve PRESERVED_SPECIMEN Rock, J.F.C. 12999 Hawaii BISH 8/1/1917
13 Melicope spathulata A.Gray Koolau Range, Paalaa-Wahiawa, Poamoho Trail PRESERVED_SPECIMEN Cowan, R.S. 248 Oahu BISH 10/20/1946
14 Melicope spathulata A.Gray S. Kohala Distr., Kawaihae 1 Ahu., Tr 78(38); Kohala Mtns; SW from Kehena Ditch to pasture 1 km NE of Puu Ahia Wet `ohi`a forest with other native trees and a treefern and native shrub understory. PRESERVED_SPECIMEN Warshauer, F.R. FRW 2468 Hawaii BISH 7/30/1979
15 Melicope spathulata A.Gray N. Kohala, Kahua Ranch On wet wooded slope. Occasional PRESERVED_SPECIMEN Hosaka, E.Y. 2030 Hawaii BISH 5/8/1938
16 Melicope spathulata A.Gray S. Hilo Dist., Paukaa Ahu. Tr 21(45); E slope of Mauna Kea, on ridge betwn Nukupahu Gulch/Awehi Stream Wet `ohi`a forest with other native trees and a treefern and native shrub understory PRESERVED_SPECIMEN Warshauer, F.R. FRW 1389 Hawaii BISH 9/11/1977
17 Melicope spathulata A.Gray Wright Road, Olaa For. Res., Puna Dist Virgin Cibotium & Metrosideros forest. PRESERVED_SPECIMEN St. John, H. 25051 Hawaii BISH 1/29/1953
18 Melicope spathulata A.Gray Divide between head of Kipapa Gulch and head of Uwau Gulch PRESERVED_SPECIMEN Hosaka, E.Y. 1877 Oahu BISH 5/30/1937
19 Melicope spathulata A.Gray Waimea District. Nuololo, headwater just west of Kaunuohua Ridge. Metrosideros-Acacia mixed mesic forest w/ Coprosma waimeae, Kadua affinis, Psychotria mariniana, P. hexandra, P. greenwelliae, Chrysodracon aurea, Scaevola procera, Xylosma hawaiiense, Dodonaea viscosa, Syzygium sandwicensis, Polyscias wai PRESERVED_SPECIMEN Wood, K.R. 17190.01 Kauai BISH 11/13/2016
20 Melicope spathulata A.Gray Kohala Mts., Puu o Umi N.A.R., Transect 8, scattered 240 & 800 meters station, on slopes of Mt. Eke in Metrosideros-Cheirodendron-Cibotium spp. montane wet forest with Broussaisia, Sadleria, Cyrtandra, Myrsine, Vaccinium, Uncinia, Alyxia, Labordia, Psychotria, Melicope, Rubus hawaiensis, Dryopteris wallichiana, Astelia, Clermontia. About 20 plants seen PRESERVED_SPECIMEN Perlman, S. 14840 Hawaii BISH 8/9/1995
21 Melicope spathulata A.Gray East Maui, Hana District, Waihoi, Puu Hoolio near Labordia, Cheirodendron PRESERVED_SPECIMEN Harrison, B. 2551 Maui BISH 7/20/1972
22 Melicope spathulata A.Gray Kuia headwaters, Kuia Natural Area Reserve. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 18363 Kauai PTBG 1/28/2020
23 Melicope spathulata A.Gray oahu, waianae mts., slope of koala, sept. 25 o. selling. 3710. hawaiian bog survey 1938. PRESERVED_SPECIMEN selling, o. 3710 GB 9/25/1938
24 Melicope spathulata A.Gray oahu PRESERVED_SPECIMEN h. mann & w. t. brigham 94 GH
25 Melicope spathulata A.Gray gorge west of howokawenui, kohala mountains PRESERVED_SPECIMEN j. f. rock 8367 GH 6/6/1910
26 Melicope spathulata A.Gray PRESERVED_SPECIMEN o. degener 8643 MPM 6/26/1926
27 Melicope spathulata A.Gray PRESERVED_SPECIMEN urbain jean faurie; b.1847; d.1915; faurie 243 NHMUK
28 Melicope spathulata A.Gray olinda, maui, t. hawaii [territory of hawaii]. along pipe-line trail. In dark forest. PRESERVED_SPECIMEN o. degener 8615 NY 7/29/1927
29 Melicope spathulata A.Gray PRESERVED_SPECIMEN h. mann jr. 94 NY
30 Melicope spathulata A.Gray KAUAI: *Lihue District, upper Wahiawa drainage, *Metrosideros-Dicranopteris low statured wet forest with wind swept ridges and riparian vegetation, w/ Machaerina angustifolia, Gahnia beecheyi, Cheirodendron platyphylla, Psychotria mariniana, Antidesma platyphylla, Dubautia laxa, Broussaisia arguta, Syzygium sandwicensis, Lobelia kauaiensis, Melicope feddei, Sphenomeris chinensis, Sadleria pallida PRESERVED_SPECIMEN K. R. Wood 12723 Kauai PTBG 12/20/2007
31 Melicope spathulata A.Gray Awa'awapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Moderate to steep slope. PRESERVED_SPECIMEN Seana Walsh SKW209 Kauai PTBG 3/4/2016
32 Melicope spathulata A.Gray Nuololo, headwaters just west of Kaunuohua Ridge. Metrosideros-Acacia mixed mesic forest PRESERVED_SPECIMEN K. R. Wood 17185 Kauai PTBG 11/8/2016
33 Melicope spathulata A.Gray Alakai, west of Sincock Bog, headwaters of Koaie Metrosideros-Cheirodendron-Dicranopteris montane wet forest, 5-7 m tall canopy, 70-90% closed, with adjacent riparian vegetation PRESERVED_SPECIMEN K. R. Wood 15312 Kauai PTBG 12/13/2012
34 Melicope spathulata A.Gray Limahuli Valley: upper eastern drainage above falls and near Pali Eleele. Metrosideros Wet Forest with dissecting drainages and windswept ridges. PRESERVED_SPECIMEN K. R. Wood 7922 Kauai PTBG 7/15/1999
35 Melicope spathulata A.Gray Koaie, headwater gulches and stream, well above Koaie river, north of Mohihi-Waialae trail. Metrosideros transitional mesic to wet forest and riparian vegetation PRESERVED_SPECIMEN K. R. Wood 15964 Kauai PTBG 6/24/2014
36 Melicope spathulata A.Gray Wahiawa Mts., Lihue-Koloa Forest Reserve, gulch just behind Kapalaoa, subgulch on east facing side of ridge. Montane Wet Metrosideros-Cheirodendron-Dicranopteris Forest. PRESERVED_SPECIMEN Natalia Tangalin 2872 Kauai PTBG 10/25/2011
37 Melicope spathulata A.Gray Mohihi, slopes and drainage north of Mohihi-Waialae trail. Metrosideros mixed montane wet forest PRESERVED_SPECIMEN K. R. Wood 16836 Kauai PTBG 5/19/2016
38 Melicope spathulata A.Gray Awa'awapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Moderate to steep slope. PRESERVED_SPECIMEN Seana Walsh SKW210 Kauai PTBG 3/4/2016
39 Melicope spathulata A.Gray Nuololo, north facing slopes above northern branch, Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 15104 Kauai PTBG 7/19/2012
40 Melicope spathulata A.Gray Kokee, end of Camp 10 road, to Mohihi Stream, up ridge above stream. On hunter trail on ridge line. Wet Metrosideros-Dicranopteris Forest PRESERVED_SPECIMEN Natalia Tangalin 2588 Kauai PTBG 4/13/2013
41 Melicope spathulata A.Gray Kohala mts., Puu Ahia area, north of Puu on drainage of Waipahoehoe stream. In Metrosideros- Cibotium wet forest. PRESERVED_SPECIMEN Steve Perlman 17831 Hawaii PTBG 12/19/2001
42 Melicope spathulata A.Gray Hono o Na Pali NAR, upper Hanakoa via Pihea. Metrosideros-Cheirodendron-Dicranopteris montane wet forest and riparian vegetation PRESERVED_SPECIMEN K. R. Wood 16164 Kauai PTBG 12/10/2014
43 Melicope spathulata A.Gray Awaawapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Groundcover of ferns. PRESERVED_SPECIMEN Seana Walsh SKW207 Kauai PTBG 3/4/2016
44 Melicope spathulata A.Gray Kokee, End of Camp 10 Road, Trail to Circle Bog Metrosideros-Dicranopteris Wet Forest PRESERVED_SPECIMEN Natalia Tangalin 4477 Kauai PTBG 12/15/2015
45 Melicope spathulata A.Gray Near the rim of Kalalau, in the Puu o Kila area. Predominately native, wet forest. Covered canopy. PRESERVED_SPECIMEN Seana Walsh SKW420 Kauai PTBG 2/26/2017
46 Melicope spathulata A.Gray Kalalau rim, 480 m southwest of Pihea. Metrosideros - Cheirodendron montane wet forest with matting ferns of Dicranopteris linearis, Diplopterygium pinnatum, Sticherus owhyhensis PRESERVED_SPECIMEN K. R. Wood 17300 Kauai PTBG 3/6/2017
47 Melicope spathulata A.Gray Awa'awapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Flat (no slope). PRESERVED_SPECIMEN Seana Walsh SKW213 Kauai PTBG 3/4/2016
48 Melicope spathulata A.Gray Kauaikinana, headwaters east of Pihea. Metrosideros-Cheirodendron montane wet forest PRESERVED_SPECIMEN K. R. Wood 17486 Kauai PTBG 7/27/2017
49 Melicope spathulata A.Gray central Alakai, southwestern portion of Halehaha proposed fence unit Metrosideros-Cheirodendron montane wet forest with Dicranopteris-Diplopterygium-Sticherus matting ferns PRESERVED_SPECIMEN K. R. Wood 15504 Kauai PTBG 5/8/2013
50 Melicope spathulata A.Gray Kauaikinana headwaters below Pihea. Metrosideros-Cheirodendron-Dicranopteris montane wet, summer mesic forest PRESERVED_SPECIMEN K. R. Wood 16065 Kauai PTBG 9/11/2014
51 Melicope spathulata A.Gray Koaie, North facing slopes well above Koaie river. Metrosideros transitional mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 15384 Kauai PTBG 2/6/2013
52 Melicope spathulata A.Gray Nuololo, forested hills to north of Nuololo trail, Metrosideros-Acacia montane mesic forest, PRESERVED_SPECIMEN K. R. Wood 15561 Kauai PTBG 7/10/2013
53 Melicope spathulata A.Gray Nualolo, MEL SPA NUA A 753. Acacia koa-Metrosideros polymorpha var. glaberrima mesic forest PRESERVED_SPECIMEN Seana Walsh SKW753 Kauai PTBG 5/20/2020
54 Melicope spathulata A.Gray Hono O Na Pali NAR, Hanakapiai headwaters via Pihea, central branch, Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation PRESERVED_SPECIMEN K. R. Wood 15286 Kauai PTBG 11/8/2012
55 Melicope spathulata A.Gray Mohihi, slopes and drainage north of Mohihi-Waialae trail. Metrosideros mixed montane wet forest, 7 m canopy, 90% closed native cover PRESERVED_SPECIMEN K. R. Wood 16784.01 Kauai PTBG 4/5/2016
56 Melicope spathulata A.Gray Mohihi, slopes and drainage north of Mohihi-Waialae trail. Metrosideros mixed montane wet forest, 7 m canopy, 90% closed native cover PRESERVED_SPECIMEN K. R. Wood 17345 Kauai PTBG 4/11/2017
57 Melicope spathulata A.Gray Nuololo, forests above headwater drainage. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 18307 Kauai PTBG 9/19/2019
58 Melicope spathulata A.Gray Nuololo, north of trail, headwaters of central Nuololo stream, Metrosideros-Acacia montane mesic to wet forest PRESERVED_SPECIMEN K. R. Wood 15321 Kauai PTBG 1/2/2013
59 Melicope spathulata A.Gray Limahuli upper valley, Hanakapiai side drainage, on east side of stream about 100 ft elevation above streambed about 100 ft. before waterfall. PRESERVED_SPECIMEN Natalia Tangalin 813 Kauai PTBG 6/29/2006
60 Melicope spathulata A.Gray Awaawapuhi, north facing slopes SW of Kainamanu. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 16597 Kauai PTBG 9/8/2015
61 Melicope spathulata A.Gray Upper Limahuli Preserve Wet Forest, Uluhe ridge PRESERVED_SPECIMEN J.C. Watson JCW008 Kauai PTBG 8/22/2013
62 Melicope spathulata A.Gray OAHU: Koolau mts. Manana trail to summit, next ridge north of Manana in Metrosideros wet forest with Platydesma spathulata, Cibotium, Syzygium sandwicense, Dubautia, Scaevola mollis, Pritchardia martii, Pipturus, Antidesma, Sadleria, Wikstroemia oahuensis, Coprosma, Melicope clusiifolia, Psychotria hathewayi, Phyllostegia grandiflora, Labordia , Plantago, Lobelia oahuensis, Cyrtandra, Tetraplasandra PRESERVED_SPECIMEN K. R. Wood 8514 Oahu PTBG 6/10/2000
63 Melicope spathulata A.Gray Awaawapuhi Mesic, mixed native and non-native forest (predominantly native). Canopy cover 80-90%. In gulch. PRESERVED_SPECIMEN Seana Walsh SKW215 Kauai PTBG 3/4/2016
64 Melicope spathulata A.Gray Koolau Mts., Moanalua Valley, summit crest ridge. In Metrosideros-Dicranopteris linearis lowland wet forest. PRESERVED_SPECIMEN Steve Perlman 5942 Oahu PTBG 6/7/1987
65 Melicope spathulata A.Gray Mohihi. About 2 miles up on right side of trail. On edge of trail. About 75% canopy cover. No slope. PRESERVED_SPECIMEN Seana Walsh SKW217 Kauai PTBG 3/17/2016
66 Melicope spathulata A.Gray Koolau Mts.,east side of Puu Pauao,between Poamoho and Schofield trail,drainages of Punaluu valley. South side subgulch. In Metrosideros mixed lowland wet forest. PRESERVED_SPECIMEN Steve Perlman 14712 Oahu PTBG 3/12/1995
67 Melicope spathulata A.Gray Below Kawaiiki Ridge along the S fork of Twin Falls. Five ft above gulch bottom. Mixed Mesic Forest. PRESERVED_SPECIMEN Natalia Tangalin 2351 Kauai PTBG 6/2/2010
68 Melicope spathulata A.Gray Waiakoali, Na Pali Kona Forest Reserve Metrosideros-Cheirodendron montane wet slopes and drainage PRESERVED_SPECIMEN K. R. Wood 15213 Kauai PTBG 9/13/2012
69 Melicope spathulata A.Gray Kahuamaa Flat, Kaunuohua Ridge. Metrosideros-Cheirodendron-Dicranopteris montane wet forest PRESERVED_SPECIMEN K. R. Wood 18134 Kauai PTBG 3/14/2019
70 Melicope spathulata A.Gray Kalalau, below Pihea. Metrosideros - Cheirodendron montane wet forest with matting ferns of Dicranopteris linearis, Diplopterygium pinnatum, Sticherus owhyhensis. PRESERVED_SPECIMEN K. R. Wood 15816 Kauai PTBG 2/2/2014
71 Melicope spathulata A.Gray Kuia, upper southern headwater gulch. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 16606 Kauai PTBG 9/13/2015
72 Melicope spathulata A.Gray Kaapahu, Haleakala National Park land. Acacia koa- Metrosideros mesic forest. PRESERVED_SPECIMEN Steve Perlman 21660 Maui PTBG 5/27/2009
73 Melicope spathulata A.Gray Awa'awapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Moderate slope. PRESERVED_SPECIMEN Seana Walsh SKW211 Kauai PTBG 3/4/2016
74 Melicope spathulata A.Gray Nuololo, trail to exclosure at 3520 ft and head north. Metrosideros-Acacia montane mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 11096 Kauai PTBG 1/2/2005
75 Melicope spathulata A.Gray Awa'awapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Groundcover of ferns. PRESERVED_SPECIMEN Seana Walsh SKW208 Kauai PTBG 3/4/2016
76 Melicope spathulata A.Gray headwaters of Koaie stream near Wainiha Pali Metrosideros-Cheirodendron-Dicranopteris mixed wet forest, 8-12 m tall canopy, 50-90% closed, PRESERVED_SPECIMEN K. R. Wood 15296 Kauai PTBG 11/16/2012
77 Melicope spathulata A.Gray Honopu forests adjacent to trail. Acacia koa - Metrosideros polymorpha - Dicranopteris linearis montane mesic PRESERVED_SPECIMEN K. R. Wood 16273 Kauai PTBG 1/23/2015
78 Melicope spathulata A.Gray Makaha, forests around dividing ridge between upper north and south fork. Metrosideros-Acacia-Alphitonia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 16743 Kauai PTBG 2/18/2016
79 Melicope spathulata A.Gray Waiakoali, upper flats to the south, Na Pali Kona Forest Reserve. Metrosideros-Cheirodendron montane wet forest PRESERVED_SPECIMEN K. R. Wood 16278 Kauai PTBG 1/27/2015
80 Melicope spathulata A.Gray Hono O Na Pali NAR, Hanakapiai headwaters, western branch, north of Moaalele. Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation PRESERVED_SPECIMEN K. R. Wood 16060 Kauai PTBG 9/3/2014
81 Melicope spathulata A.Gray ridge just south below Kawaikini, Iliiliula headwaters Dubautia-Sadleria shrubland/fernland, high wind summit slopes PRESERVED_SPECIMEN K. R. Wood 15467.01 Kauai PTBG 4/3/2013
82 Melicope spathulata A.Gray Puu Makaala NAR, Dissappointment Road, along NAR survey transect station 1.0. Metrosideros polymorpha-Cibotium splendens- Montane Wet Forest. PRESERVED_SPECIMEN Steve Perlman 11465 Hawaii PTBG
83 Melicope spathulata A.Gray Kohua Ridge, North facing slopes above Mohihi Stream. Metrosideros-Acacia montane mesic forest. PRESERVED_SPECIMEN K. R. Wood 15091 Kauai PTBG 7/12/2012
84 Melicope spathulata A.Gray KAUAI: upper Honopu, forested region below and west of Kainamanu, Acacia koa-Metrosideros polymorpha montane mesic forest w/ Psychotria mariniana, Melicope anisata, M. barbigera, Pouteria sandwicensis, Dodonaea viscosa, Dianella sandwicensis PRESERVED_SPECIMEN K. R. Wood 8288 Kauai PTBG 3/3/2000
85 Melicope spathulata A.Gray KAUAI: Waimea District; Mohihi-Waialae Trail; Mohihi stream near trailhead; Metrosideros Montane Wet Forest w/ Cheirod- endron, Hedyotis terminalis, Psychotria mariniana, Sadleria, Sticherus, Diploterygium pinnatum. Threats: Rubus argutus, Pigs. Elev. ca. 3500 ft. PRESERVED_SPECIMEN K. R. Wood 2547 Kauai PTBG 5/10/1993
86 Melicope spathulata A.Gray Hono O Na Pali NAR, Hanakapiai headwaters, western branch, east of Moaalele. Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation PRESERVED_SPECIMEN K. R. Wood 16404 Kauai PTBG 4/10/2015
87 Melicope spathulata A.Gray KAUAI: Makaha valley behind Keo Keo shelter in Acacia koa- Metrosideros mesic forest with Myrsine alyxifolia, Bobea brevipes, Pouteria sandwicensis, Pleomele aurea, Dodonaea viscosa, Psychotria greenwelliae, Xylosma hawaiiense, Perrottetia sanwicensis, Hedyotis terminalis, Tetraplasandra, Alphitonia ponderosa, Rubus argutus, Diplazium sandwichianum, Passiflora mollissima PRESERVED_SPECIMEN Steve Perlman 18660 Kauai PTBG 10/22/2003
88 Melicope spathulata A.Gray Awaawapuhi valley, west aspect. Acacia koa-Metrosideros-Dicranopteris linearis mesic forest PRESERVED_SPECIMEN K. R. Wood 5391 Kauai PTBG 12/20/1996
89 Melicope spathulata A.Gray Aiea trail, near summit, between Kaivipoo hill and large power relay. Metrosideros- Dicranopteris linearis lowland wet forest. PRESERVED_SPECIMEN Steve Perlman 23644 Oahu PTBG 8/20/2013
90 Melicope spathulata A.Gray Mohihi. About 1.25 miles down trail, on right side about 3 feet away from trail. Mesic forest on upper, wide ridge. Canopy cover about 65%. PRESERVED_SPECIMEN Seana Walsh SKW102 Kauai PTBG 9/4/2015
91 Melicope spathulata A.Gray Waiakoali headwaters, South branch. Metrosideros-Cheirodendron montane wet, steep north facing closed to open forest with rich fern and moss ground cover. PRESERVED_SPECIMEN K. R. Wood 14346 Kauai PTBG 10/8/2010
92 Melicope spathulata A.Gray Kohala Mts.; Puu Ahia. Metrosideros-Cheirodendron Montane Wet PRESERVED_SPECIMEN K. R. Wood 4782 Hawaii PTBG 11/21/1995
93 Melicope spathulata A.Gray Honopu, NW of Kainamanu, Acacia koa - Metrosideros polymorpha - Dicranopteris linearis montane mesic PRESERVED_SPECIMEN K. R. Wood 15150 Kauai PTBG 8/8/2012
94 Melicope spathulata A.Gray KAUAI: Wainiha Valley, back of valley, on left (N side) fork (not Hinalele side) at very back of valley, subgulches and near falls; Metrosideros lowland wet forest with Perrottetia Boehmeria, Diplazium, Cyrtandra spp., Tetraplasandra, Psychotria, Hedyotis terminalis, Cibotium spp., and Peperomin. ELev. 2500 ft. PRESERVED_SPECIMEN Steve Perlman 13291 Kauai PTBG 1/29/1993
95 Melicope spathulata A.Gray Nuololo, headwaters just west of Kaunuohua Ridge. Metrosideros-Acacia mixed mesic forest PRESERVED_SPECIMEN K. R. Wood 17190.01 Kauai PTBG 11/13/2016
96 Melicope spathulata A.Gray Nuololo, northern headwater gulches, ca. 350 m south of Awaawapuhi trail. Metrosideros-Acacia montane mesic forest transitioning into Metrosideros-Dicranopteris wet forest PRESERVED_SPECIMEN K. R. Wood 16779 Kauai PTBG 3/31/2016
97 Melicope spathulata A.Gray Koaie, headwater gulches and stream, well above Koaie river, north of Mohihi-Waialae trail. Metrosideros transitional mesic to wet forest and riparian vegetation PRESERVED_SPECIMEN K. R. Wood 15963 Kauai PTBG 6/24/2014
98 Melicope spathulata A.Gray Kokee, Na Pali-Kona Forest Reserve, Alakai Swamp Trail. Mesic to wet Metrosideros polymorpha forest. PRESERVED_SPECIMEN Marc Appelhans MA672 Kauai PTBG 10/5/2016
99 Melicope spathulata A.Gray Hono O Na Pali NAR, Pohakea region above Hanakapiai. Metrosideros-Dicranopteris-montane mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 14374 Kauai PTBG 11/18/2010
100 Melicope spathulata A.Gray Puumakaala NAR; off Dissappointment Road, in fenced exclosure. Metrosideros-Cibotium Wet Forest. PRESERVED_SPECIMEN Natalia Tangalin 637 Hawaii PTBG 12/19/2005
101 Melicope spathulata A.Gray Wainiha Pali, eastern boundary of Hono O Na Pali Natural Area Reserve. Metrosideros-Cheirodendron montane wet forest with Dicranopteris, on side stream above cliffs PRESERVED_SPECIMEN K. R. Wood 16025 Kauai PTBG 7/31/2014
102 Melicope spathulata A.Gray Nuololo, north facing slopes above drainage, Metrosideros-Acacia montane mesic forest, 70-80% closed, up to 20 m tall canopy, PRESERVED_SPECIMEN K. R. Wood 14551b Kauai PTBG 2/28/2011
103 Melicope spathulata A.Gray Upper Limahuli Preserve, in drainage in SW Bowl Metrosideros-Cheirodendron-Dicranopteris Wet Forest PRESERVED_SPECIMEN Natalia Tangalin 3954 Kauai PTBG 11/17/2014
104 Melicope spathulata A.Gray Pohakea. About 5 feet off side of trail, 30 feet away from NARS weather port. PLA SPA WAI A 0001. In understory with bare ground on flat surface. PRESERVED_SPECIMEN Seana Walsh SKW236 Kauai PTBG 4/6/2016
105 Melicope spathulata A.Gray Mohihi. About 2 miles down trail, then down to stream off left side of trail. Continued down gulch to opening at larger stream and continued down stream, on right side. Mesic-wet forest alongside stream. Canopy cover about 50% PRESERVED_SPECIMEN Seana Walsh SKW107 Kauai PTBG 9/4/2015
106 Melicope spathulata A.Gray Milolii. NW side Predominately non-native vegetation. Mostly bare ground. Canopy cover about 90%. PRESERVED_SPECIMEN Seana Walsh SKW99 Kauai PTBG 9/3/2015
107 Melicope spathulata A.Gray Kalalau Rim, just West of Puu o Kila. Metrosideros-Cheirodendron-Dicranopteris transitional mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 15187 Kauai PTBG 8/16/2012
108 Melicope spathulata A.Gray Nahiku, E. Maui PRESERVED_SPECIMEN C. N. Forbes 240.M Maui PTBG 7/1/1910
109 Melicope spathulata A.Gray Kohala Mts.,Puu o Umi N.A.R.,Transect 8,scattered 240 & 800 meters station, on slopes of Mt. Eke. In Metrosideros-Cheirodendron-Cibotium spp. montane wet forest. PRESERVED_SPECIMEN Steve Perlman 14840 Hawaii PTBG 8/9/1995
110 Melicope spathulata A.Gray KAUAI: Kawai`iki, off Kaluakaulu ridge, upper forest and drainage to the south of Koai’e and north of Waialae, Metrosideros polymorpha mixed mesic forest w/ Dubautia laevigata, Hedyotis terminalis, Cheirodendron trigynum, Dianella sandwicensis, Poa sandwicensis, Bidens cosmoides, Peperomia membranacea PRESERVED_SPECIMEN K. R. Wood 9002 Kauai PTBG 6/27/2001
111 Melicope spathulata A.Gray West Hanakapiai, northeast of Pihea. Metrosideros-Cheirodendron forest 5-8 m tall, steep slopes of Dicranopteris, Diplopterygium, & Sticherus, dissected by riparian vegetation. PRESERVED_SPECIMEN K. R. Wood 14327 Kauai PTBG 9/28/2010
112 Melicope spathulata A.Gray Awa'awapuhi Mesic, mixed native and non-native forest (predominantly native). In understory with about 80-90% canopy cover. Flat (no slope). PRESERVED_SPECIMEN Seana Walsh SKW212 Kauai PTBG 3/4/2016
113 Melicope spathulata A.Gray Below Waialeale Summit Bogs. Eastern wind-swept ridges, montane shrubs with forest patches on flanks. PRESERVED_SPECIMEN K. R. Wood 12289 Kauai PTBG 3/6/2007
114 Melicope spathulata A.Gray Kumuwela, gulch just South of Maluapopoki Stream. Metrosideros-Acacia mesic forest. PRESERVED_SPECIMEN K. R. Wood 15222 Kauai PTBG 9/19/2012
115 Melicope spathulata A.Gray Koolau mts., Manana valley, south side of valley, on ridge top, scattered plants in area, north facing slope. In Metrosideros - Dicranopteris linearis windswept low veg. PRESERVED_SPECIMEN Steve Perlman 17177 Oahu PTBG 6/10/2000
116 Melicope spathulata A.Gray Nuololo, trail to exclosure at 3520 ft and head north. Metrosideros-Acacia montane mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 11093 Kauai PTBG 1/2/2005
117 Melicope spathulata A.Gray Waiakoali, Na Pali Kona Forest Reserve Metrosideros-Cheirodendron montane wet slopes and drainage PRESERVED_SPECIMEN K. R. Wood 15207 Kauai PTBG 9/13/2012
118 Melicope spathulata A.Gray Wahiawa, Metrosideros-Dicranopteris low statured wet forest with wind swept ridges and riparian vegetation, w/ Machaerina angustifolia, Gahnia beecheyi, Cheirodendron platyphylla, Psychotria mariniana, Dubautia laxa, Broussaisia arguta, Syzygium sandwicensis, Lobelia kauaiensis, Melicope feddei, Sphenomeris chinensis, Sadleria pallida PRESERVED_SPECIMEN K. R. Wood 14213 Kauai PTBG 5/12/2010
119 Melicope spathulata A.Gray Kokee State Park, Pihea, south east (behind) the back of Kalalau Valley rim. PLA SPA ALP A 0001 Ohia (Metrosideros) Montane Wet Mixed Community PRESERVED_SPECIMEN Seana Walsh SKW548 Kauai PTBG 10/11/2017
120 Melicope spathulata A.Gray Honopu, slopes around northern branch. Acacia koa-Metrosideros polymorpha-Dicranopteris linearis mesic to transitional wet forest and drainage PRESERVED_SPECIMEN K. R. Wood 16437 Kauai PTBG 5/2/2015
121 Melicope spathulata A.Gray Nuololo, forests west of road. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 18310 Kauai PTBG 9/26/2019
122 Melicope spathulata A.Gray KAUAI: Kawaiiki valley in Metrosideros mixed mesic forest with Dodonaea viscosa, Alphitonia, Perrottetia , Cheirodendron, Hedyotis terminalis, Peperomia, Rubus argutus, Lantana, Bidens cosmoides, Diplazium sandwichianum, Psidium cattleianum, Charpentiera elliptica, Acacia koa, Diellia erecta, Cyanea fissa PRESERVED_SPECIMEN K. R. Wood 9019 Kauai PTBG 6/27/2001
123 Melicope spathulata A.Gray Mohihi. About 2.25 miles up, about 10 feet off right side of trail. On south facing, open, moderate slope covered in Dicranopteris linearis. PRESERVED_SPECIMEN Seana Walsh SKW218 Kauai PTBG 3/17/2016
124 Melicope spathulata A.Gray Kauaikinana headwaters east of Pihea trail. Metrosideros-Cheirodendron-Dicranopteris montane wet, summer mesic forest PRESERVED_SPECIMEN K. R. Wood 17459 Kauai PTBG 6/28/2017
125 Melicope spathulata A.Gray Wainiha, upper northeastern fork. Closed Metrosideros forest, 8-12 m canopy. PRESERVED_SPECIMEN K. R. Wood 13262 Kauai PTBG 9/10/2008
126 Melicope spathulata A.Gray Nuololo, headwaters just west of Kaunuohua Ridge. Metrosideros-Acacia mixed mesic forest PRESERVED_SPECIMEN K. R. Wood 17186 Kauai PTBG 11/8/2016
127 Melicope spathulata A.Gray Kokee State Park, Pihea, south east (behind) the back of Kalalau Valley rim. PLA SPA ALP A 0002 Ohia (Metrosideros) Montane Wet Mixed Community. PRESERVED_SPECIMEN Seana Walsh SKW550 Kauai PTBG 10/11/2017
128 Melicope spathulata A.Gray Kahili mountain region. Metrosideros-Dicranopteris lowland to montane wet habitat PRESERVED_SPECIMEN K. R. Wood 16382 Kauai PTBG 3/27/2015
129 Melicope spathulata A.Gray Upper Limahuli Preserve, Wainiha or East side drainage between Pali Eleele & Hono O Na Pali. Off of spur referred to as the Express Way. Metrosideros-Dicranopteris-Diplopterygium-Cheirodendron Wet Forest PRESERVED_SPECIMEN Natalia Tangalin 2544 Kauai PTBG 3/6/2012
130 Melicope spathulata A.Gray KAUAI: Border of Lihue and Kawaihau District; Blue hole; below Waialeale and Kawaikini; Headwaters of N Wailua River; Metrosideros lowland Wet Forest surrounded by verticle cliffs of Machaerina angustifolia and associated w/ Bobea, Perrottetia, Gunnera, numerous Cyrtandra, Cyanea, small stunted trees; pteridophytes are common. Major Threats; Passiflora edulis, landslides. Elev. 2000-2200 ft. PRESERVED_SPECIMEN K. R. Wood 786 Kauai PTBG 5/7/1991
131 Melicope spathulata A.Gray KAUAI: North facing slope of upper Awaawapuhi valley. Diverse mesic forest. Growing with Melicope haupuensis, Dodonea viscosa, Psychotria greenwellii, Metrosideros polymorpha, Cryptocaria mannii and Nothocestrum peltatum. Area being invaded by Rubus argutus PRESERVED_SPECIMEN Clay Trauernicht 605 Kauai PTBG 11/11/2008
132 Melicope spathulata A.Gray Kohua Ridge, north facing slopes above Awini Stream Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 14975 Kauai PTBG 5/16/2012
133 Melicope spathulata A.Gray Nuololo, forests above headwater drainage. Metrosideros-Acacia montane mesic, high canopy forest PRESERVED_SPECIMEN K. R. Wood 18420.04 Kauai PTBG 5/20/2020
134 Melicope spathulata A.Gray Kawaikoi, western headwaters. Metrosideros-Cheirodendron-Dicranopteris montane wet forest and riparian habitat PRESERVED_SPECIMEN K. R. Wood 16117 Kauai PTBG 10/3/2014
135 Melicope spathulata A.Gray Koaie, headwater gulches and forested slopes, north of Mohihi-Waialae trail Metrosideros-Cheirodendron montane wet forest with Dicranopteris-Diplopterygium-Sticherus matting ferns PRESERVED_SPECIMEN K. R. Wood 15722 Kauai PTBG 11/13/2013
136 Melicope spathulata A.Gray KAUAI: Honopu valley, around Kainamanu in Acacia koa- Metrosideros- Dicranopteris linearis wet forest with Claoxylon sandwicensis, Psychotria mariniana, Coprosma kauaiensis, Hillebrandia sandwicensis, Ranunculus mauiensis, Passiflora mollissima, Rubus argutus, Pouteria, Platydesma spathulata, Cheirodenron, Tetraplasandra, Perrottetia, Labordia degeneri, Erigeron karvinskianus PRESERVED_SPECIMEN K. R. Wood 8831 Kauai PTBG 1/5/2001
137 Melicope spathulata A.Gray Nuololo, northern branch. Metrosideros-Acacia montane mesic forest transitioning into Metrosideros-Dicranopteris wet forest. PRESERVED_SPECIMEN K. R. Wood 15681 Kauai PTBG 10/6/2013
138 Melicope spathulata A.Gray KAUAI: Hanalei District; Wainiha; drainage south of Mahina- kehau; Metrosideros-Dicranopteris Lowland Wet Forest w/ saturated stream walls; Cyrtandra limahuliensis, C.oenobarba C.cyaniodes, C.pickeringii, Bidens spp., Cyanea spp., Hedy- otis spp., pteridophytes are abundant. Threats: Rubus rosi- folius, Erigeron, Pigs, Landslides. Elev. ca. 700-800 m. PRESERVED_SPECIMEN K. R. Wood 2330 Kauai PTBG 1/29/1993
139 Melicope spathulata A.Gray on the lower slopes of konahuanui, above manoa PRESERVED_SPECIMEN a. a. heller 2373 Oahu US 1895-05-28
140 Melicope spathulata A.Gray Hawaiian Islands; Kauai. East rim of Kalalau Valley. Sunny, shrubby rain forest. PRESERVED_SPECIMEN O. Degener; I. Degener, W. Cadenhead 27150 SFV 11/16/1960
141 Melicope spathulata A.Gray Oahu Is., koolau Mts., laie-waimea divide wet forest PRESERVED_SPECIMEN M. Chong s.n. UBC 4/15/1933
142 Melicope spathulata A.Gray PRESERVED_SPECIMEN s. carlquist 2197 Maui US
143 Melicope spathulata A.Gray waimea district, kohua ridge, north facing slopes above mohihi stream. Montane mesic forest. PRESERVED_SPECIMEN k. r. wood, m. query & m. kirkpatrick 15091 Kauai US 7/12/2012
144 Melicope spathulata A.Gray mohihi river, waimea PRESERVED_SPECIMEN h. st. john 19999 Kauai US 2/12/1939
145 Melicope spathulata A.Gray beyond summit of piko trail, above makua valley PRESERVED_SPECIMEN o. degener, k. park & y. nitta 8630 Oahu US 7/10/1932
146 Melicope spathulata A.Gray land of olaa, olaa forest reserve, 2/3 mile wsw of pu`u maka`ala. PRESERVED_SPECIMEN d. r. herbst & m. mull 5507 Hawaii US 10/25/1975
147 Melicope spathulata A.Gray kaholuamano PRESERVED_SPECIMEN a. s. hitchcock 15344 Kauai US 10/20/1916
148 Melicope spathulata A.Gray e bank of honomanu gulch PRESERVED_SPECIMEN c. n. forbes 2568.m. Maui US 6/21/1920
149 Melicope spathulata A.Gray beyond cabin at top of pig-god trail, punaluu PRESERVED_SPECIMEN o. degener, k. park & y. nitta 8633 Oahu US 6/22/1932
150 Melicope spathulata A.Gray behind honolulu. PRESERVED_SPECIMEN h. mann & w. t. brigham 94 Oahu US
151 Melicope spathulata A.Gray waimea drainage basin, w side PRESERVED_SPECIMEN c. n. forbes 782.k Kauai US 7/3/1917
152 Melicope spathulata A.Gray off honopu trail, kokee state park PRESERVED_SPECIMEN s. s. ishikawa 302 Oahu US 8/22/1973
153 Melicope spathulata A.Gray kalalau trail, above kokee PRESERVED_SPECIMEN f. r. fosberg 12717 Kauai US 12/29/1935
154 Melicope spathulata A.Gray forests of mauna kea. PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US 1838-08-08
155 Melicope spathulata A.Gray kaholuamano PRESERVED_SPECIMEN a. s. hitchcock 15326 Kauai US 10/20/1916
156 Melicope spathulata A.Gray kipapa gulch, s ridge, koolau mts PRESERVED_SPECIMEN e. y. hosaka 970 Oahu US 4/16/1933
157 Melicope spathulata A.Gray PRESERVED_SPECIMEN s. carlquist 1995 Maui US
158 Melicope spathulata A.Gray ewa forest reserve, koolau range PRESERVED_SPECIMEN e. y. hosaka 2031 Oahu US 5/8/1938
159 Melicope spathulata A.Gray olinda, along pipe line trail PRESERVED_SPECIMEN o. degener 2367 Maui US 6/14/1927
160 Melicope spathulata A.Gray konahuanui PRESERVED_SPECIMEN c. n. forbes s.n. Oahu US 8/8/1908
161 Melicope spathulata A.Gray awaawapuhi valley, north- facing slope near kainamanu PRESERVED_SPECIMEN s. p. perlman & l. m. dunn 18660 Kauai US 10/22/2003
162 Melicope spathulata A.Gray vicinity of wahiawa swamp PRESERVED_SPECIMEN c. n. forbes 231.k Kauai US
163 Melicope spathulata A.Gray PRESERVED_SPECIMEN w. b. hillebrand 1801 Hawaii US
164 Melicope spathulata A.Gray main divide, crest of koolau mts, at head of n branch of kahana valley PRESERVED_SPECIMEN f. r. fosberg & e. y. hosaka 13934 Oahu US 5/30/1937
165 Melicope spathulata A.Gray valley s of e ridge of puu kalena, waianae mts, waianaeuka PRESERVED_SPECIMEN f. r. fosberg 12994 Oahu US 3/22/1936
166 Melicope spathulata A.Gray wahiawa stream PRESERVED_SPECIMEN s. p. perlman 11679 Kauai US
167 Melicope spathulata A.Gray waianae mts, puu kanehoa and kaluaa gulch PRESERVED_SPECIMEN s. p. darwin 1032 Hawaii US 6/22/1975
168 Melicope spathulata A.Gray poamoho trail, near top PRESERVED_SPECIMEN o. degener 27328 Oahu US 3/9/1961
169 Melicope spathulata A.Gray waimea drainage basin, w side PRESERVED_SPECIMEN c. n. forbes 836.k. Kauai US 7/3/1917
170 Melicope spathulata A.Gray kaholuamano PRESERVED_SPECIMEN a. s. hitchcock 15342 Kauai US 10/20/1916
171 Melicope spathulata A.Gray hanalei district, koke`e state park, kalalau rim w of pu`u o kila lookout, just past drainage below lookout. PRESERVED_SPECIMEN w. l. wagner & k. r. wood 6893 Kauai US 2/25/2000
172 Melicope spathulata A.Gray Waimea District, Kohua Ridge, north facing slopes above Mohihi Stream. Montane mesic forest. PRESERVED_SPECIMEN K. R. Wood, M. Query & M. Kirkpatrick 15091 Kauai US 7/12/2012
173 Melicope spathulata A.Gray along pipe line trail, olinda PRESERVED_SPECIMEN o. degener 2367 Maui US 6/14/1927
174 Melicope spathulata A.Gray koolau range, between punaluu and kaipapau PRESERVED_SPECIMEN c. m. cooke & c. n. forbes s.n. Oahu US 5/3/1909
175 Melicope spathulata A.Gray Waimea District, Nuololo, headwaters just west of Kaunuohua Ridge Mixed mesic forest PRESERVED_SPECIMEN K. R. Wood & M. Query 17190 Kauai US 11/13/2016
176 Melicope spathulata A.Gray Hanalei District, Koke`e State Park, Kalalau rim W of Pu`u o Kila lookout, just past drainage below lookout. PRESERVED_SPECIMEN W. L. Wagner & K. R. Wood 6893 Kauai US 2/25/2000
177 Melicope spathulata A.Gray waimea district, nuololo, headwaters just west of kaunuohua ridge Bank of a stream PRESERVED_SPECIMEN k. r. wood & m. query 17190 Molokai US 11/13/2016
178 Melicope spathulata A.Gray Kokee State Park, along Nualolo Trail about 1 mile from bigining at the Kokee Lodge PRESERVED_SPECIMEN Sohmer, S.H. 6507 Kauai BISH 7/16/1970
179 Melicope spathulata A.Gray Hanalei, pole line trail PRESERVED_SPECIMEN Herbst, D.R. 2462 Kauai BISH 5/31/1972
180 Melicope spathulata A.Gray Koolau Mountains, along crest, ca. 1.6 km (1 mi) W of 'Aiea trail summnit; Off a grassy mound on the leeward side PRESERVED_SPECIMEN Obata, J.K. 85-603 Oahu BISH 8/11/1985
181 Melicope spathulata A.Gray Koolau Range, Kipapa Gulch Trail in moist area along trail PRESERVED_SPECIMEN Cowan, R.S. 700 Oahu BISH 10/5/1947
182 Melicope spathulata A.Gray kipapa gulch, s ridge of koolau mts PRESERVED_SPECIMEN e. y. hosaka 617 Oahu US 7/4/1932
183 Melicope spathulata A.Gray vicinity forest ranger station at kokee, pipeline trail PRESERVED_SPECIMEN w. l. stern & s. carlquist 1307 Kauai US
184 Melicope spathulata A.Gray Kumuwela ridge Below Kokee PRESERVED_SPECIMEN Cranwell, L.M. 2841 Kauai BISH 8/13/1938
185 Melicope spathulata A.Gray Koolau Range, between Punaluu and Kaipapau PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 5/3/1909
186 Melicope spathulata A.Gray Kohua Ridge, north facing slopes above Mohihi Stream. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 17335 Kauai PTBG 4/6/2017
187 Melicope spathulata A.Gray Wahiawa Mts 80' from stream bed; east exposure PRESERVED_SPECIMEN Perlman, S. 490 Kauai BISH 9/1/1979
188 Melicope spathulata A.Gray Maakua-Papali ridge Kaipapau Forest Reserve Hauula In rain forest PRESERVED_SPECIMEN Yoshioka, T. s.n. Oahu BISH 10/15/1933
189 Melicope spathulata A.Gray Waimea District. Kohua Ridge, north facing slopes above Mohihi Stream. Metrosideros-Acacia montane mesic forest with 80% closed canopy, 10-12 meters tall, with Cheirodendron trigynum, Kadua affinis, Coprosma waimeae, Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Dodonaea viscosa, Myrsine lanaiensis. PRESERVED_SPECIMEN Wood, K.R. 17335 Kauai BISH 4/6/2017
190 Melicope spathulata A.Gray Kipapa Gulch, So. ridge Koolau Mts on wooded ridge PRESERVED_SPECIMEN Hosaka, E.Y. 617 Oahu BISH 7/4/1932
191 Melicope spathulata A.Gray Honopu, northern slopes and drainage below Kainamanu. Acacia koa - Metrosideros polymorpha - Dicranopteris linearis montane mesic PRESERVED_SPECIMEN K. R. Wood 17533 Kauai PTBG 9/21/2017
192 Melicope spathulata A.Gray Waimea Highlands, Kokee, near Ranger Station, in rainforest in rainforest PRESERVED_SPECIMEN Stone, B.C. s.n. Kauai BISH 8/8/1961
193 Melicope spathulata A.Gray Koolau Mts., Punaluu PRESERVED_SPECIMEN 459 Oahu BISH 12/24/1908
194 Melicope spathulata A.Gray East Maui PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH 5/1/1911
195 Melicope spathulata A.Gray Koolau Mts. between Punaluu & Kaipapau PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 2/14/1908
196 Melicope spathulata A.Gray Honopu, slopes around northern branch. Acacia koa-Metrosideros polymorpha-Dicranopteris linearis mesic to transitional wet forest and drainage PRESERVED_SPECIMEN K. R. Wood 16415 Kauai PTBG 4/28/2015
197 Melicope spathulata A.Gray East Maui, Nohiku PRESERVED_SPECIMEN Forbes, C.N. 240.M Maui BISH 7/1/1910
198 Melicope spathulata A.Gray pupukea. Slopes rocky with plenty moisture PRESERVED_SPECIMEN w. n. takeuchi koolau_518b Maui BISH 2/18/1985
199 Melicope spathulata A.Gray Trail between Konahuanui & Olympus PRESERVED_SPECIMEN Forbes, C.N. 1637 O Oahu BISH 2/14/1911
200 Melicope spathulata A.Gray KAUAI: *Waioli, hanging valley on north face of Namolokama, prominent ridge to east of main falls, * Metrosideros-Cheirodendron wind swept forest and shrubland with Sticherus, Diplopterygium, and Dicranopteris matting ferns, associated with Psychotria mariniana, P. hexandra, Dubautia laxa, D. knudsenii, Bidens campylotheca, Freycinetia arborea, Sadleria pallida, Hedyotis tryblium PRESERVED_SPECIMEN K. R. Wood 12186 Kauai PTBG 10/18/2006
201 Melicope spathulata A.Gray KAUAI: Awaawapuhi valley, north facing steep slope near Kainamanu in Acacia koa - Metrosideros forest with Pouteria, Pittosporum, Nestegis, Cheirodendron, Scaevola procera, Coprosma kauaiensis, Bobea brevipes, Pleomele aurea, Dicranopteris linearis, Melicope anisata, Ilex, Xylosma hawaiiense, Elaeocarpus, Perrottetia, Passiflora mollissiama, Myrica faya, Rubus argutus, Hedychium gardnerianum PRESERVED_SPECIMEN Steve Perlman 18660 Kauai PTBG 10/22/2003
202 Melicope spathulata A.Gray Kohala, Honokanenui PRESERVED_SPECIMEN Rock, J.F.C. 8367 Hawaii BISH 6/1/1910
203 Melicope spathulata A.Gray Below Kawaiiki Ridge along the S fork of Twin Falls. 5 ft above gulch bottom. Mixed Mesic Forest. Canopy of A koa and Metrosideros polymorpha, Dicranopteris, w/ Pittosporum, Psychotria, Coprosma, Cyanea leptostegia, Asplenium, Cheirodendron trigynum, Dianella sandwicensis, Poa sandwicensis, Perrottetia PRESERVED_SPECIMEN Tangalin, N. 2351 Kauai BISH 6/2/2010
204 Melicope spathulata A.Gray KAUAI: boundary of Lihue & Kawaihau Districts; below the "Blue Hole"; headwaters of the N fork of the Wailua River, just N. of Mt. Kawaikini, & SE of Mt. Waialeale; deep, narrow valley surrounded by vertical cliffs laced w/ waterfalls low, undisturbed wet forest of small, stunted trees and shrubs with groundcover of pteridophytes, Gunnera, Cyrtandra and Cyanea over saturated rocky ground. PRESERVED_SPECIMEN K. R. Wood 1123 Kauai PTBG 8/7/1991
205 Melicope spathulata A.Gray Halii Valley PRESERVED_SPECIMEN Degener, O. 30247 Kauai BISH 6/11/1926
206 Melicope spathulata A.Gray Wahiawa, Waikane-Schofield Trail, Koolau Mts Wooded, upper divide PRESERVED_SPECIMEN St. John, H. 10161 Oahu BISH 1/19/1930
207 Melicope spathulata A.Gray Waimea Highlands, Kokee Camp, Near Ranger Station, in rainforest in rainforest. PRESERVED_SPECIMEN Stone, B.C. s.n. Kauai BISH 8/8/1961
208 Melicope spathulata A.Gray Within a kipuka near the Puu Makaala access road, Upper Waiakea Forest Feserve, S. Hilo District Open to closed Metrosideros-Cibotium forest PRESERVED_SPECIMEN Clarke, G. Collector Number: 636 Hawaii BISH 10/27/1981
209 Melicope spathulata A.Gray Kokee State Park, Kalalau Valley Rim W. of Puu O Kila lookout, just past drainage below lookout. Metrosidros polymorpha dominated wet forest. PRESERVED_SPECIMEN W. L. Wagner 6893 Kauai PTBG 2/25/2000
210 Melicope spathulata A.Gray ridge east of Kukuiala Mokuleia Forest Preserve Gulch Waianae Mts PRESERVED_SPECIMEN Morley, H. 105 Oahu BISH 2/4/1934
211 Melicope spathulata A.Gray central Alakai, southwest of Sincock bog, southern boundary of Halehaha proposed fence unit Metrosideros-Cheirodendron montane wet forest with Dicranopteris-Diplopterygium-Sticherus matting ferns PRESERVED_SPECIMEN K. R. Wood 15490 Kauai PTBG 4/25/2013
212 Melicope spathulata A.Gray Waikamoi, below Entomology field station PRESERVED_SPECIMEN Gillett, G.W. 1879 Maui BISH 4/14/1966
213 Melicope spathulata A.Gray PRESERVED_SPECIMEN w. l. stern & s. carlquist 1307 Kauai US
214 Melicope spathulata A.Gray Pupukea on Clidemia infested gully slopes, ohia scrub forest at 1800' el. PRESERVED_SPECIMEN Takeuchi, W.N. 1997 Oahu BISH 2/18/1985
215 Melicope spathulata A.Gray Pupukea Trail PRESERVED_SPECIMEN Pearsall, G.A. s.n. Oahu BISH 4/24/1947
216 Melicope spathulata A.Gray Ridge S. of S. Opaeula Gulch, Paalaa, Koolau Mts Moist woods PRESERVED_SPECIMEN St. John, H. 13326 Oahu BISH 9/24/1933
217 Melicope spathulata A.Gray Wahiawa Mts PRESERVED_SPECIMEN Forbes, C.N. 231 K Kauai BISH 8/1/1909
218 Melicope spathulata A.Gray Alakai Swamp trail, Napali-Kona FR PRESERVED_SPECIMEN Crosby, M.R. 2015 Kauai BISH 8/12/1964
219 Melicope spathulata A.Gray Pupukea on Clidemia infested gully slopes, ohia scrub forest at 1850' el. PRESERVED_SPECIMEN Takeuchi, W.N. 1997 Oahu BISH 2/18/1985
220 Melicope spathulata A.Gray S Kohala District. Kawaihae 1 AHU. Tr 78 (38); Kohala Mtns; SW from Kehena Ditch to pasture 1km NE of Puu Ahia Wet `ohi`a forest with other native trees adn a treefern adn native shrub understory. PRESERVED_SPECIMEN Warshauer, F.R. FRW 2468 Hawaii BISH 7/30/1979
221 Melicope spathulata A.Gray Pihea, ravine, NE facing slopes PRESERVED_SPECIMEN Takeuchi, W.N. Alakai 142a Kauai BISH 7/30/1983
222 Melicope spathulata A.Gray Awaawapuhi trail. Honopu PRESERVED_SPECIMEN Woolford, H.A. 423 Kauai BISH 12/29/1956
223 Melicope spathulata A.Gray Halemanu PRESERVED_SPECIMEN Meebold, A. 26345 Kauai BISH 11/1/1938
224 Melicope spathulata A.Gray Alakai Swamp, Aipoinui bog, gully Pelea cruciata PRESERVED_SPECIMEN Takeuchi, W.N. 2460 Kauai BISH 9/21/1985
225 Melicope spathulata A.Gray Kaipapau Many along trail. PRESERVED_SPECIMEN Suehiro, A. 137 Oahu BISH 10/15/1933
226 Melicope spathulata A.Gray W Waimea drainage basin PRESERVED_SPECIMEN Forbes, C.N. 836 K Kauai BISH 7/3/1917
227 Melicope spathulata A.Gray Kaea PRESERVED_SPECIMEN Forbes, C.N. 2552.M Maui BISH 6/18/1920
228 Melicope spathulata A.Gray Kawailoa summit in Dicranopteris/Clidemia thicket PRESERVED_SPECIMEN Takeuchi, W.N. s.n. Oahu BISH 12/1/1985
229 Melicope spathulata A.Gray Oahu PRESERVED_SPECIMEN Mann, H. Brigham, W.T. Collector Number: 94 Oahu BISH
230 Melicope spathulata A.Gray Oahu PRESERVED_SPECIMEN Mann, H. 94 Oahu BISH 2/6/1905
231 Melicope spathulata A.Gray Wright Road, Olaa Forest Reserve, Puna District Virgin Cibotium & Metrosideros forest PRESERVED_SPECIMEN St. John, H. 25051 Hawaii BISH 1/29/1953
232 Melicope spathulata A.Gray East Maui, Makawao District, above Olinda, Ukulele woods PRESERVED_SPECIMEN Forbes, C.N. 205 M Maui BISH 7/1/1910
233 Melicope spathulata A.Gray Laie-Malaekahuna Ridge PRESERVED_SPECIMEN St. John, H. 11551 Oahu BISH 2/14/1932
234 Melicope spathulata A.Gray Honomanu Trail PRESERVED_SPECIMEN Rock, J.F.C. 10228 Maui BISH 4/1/1911
235 Melicope spathulata A.Gray Hana District, Kaapahu, Haleakala National Park land Acacia koa-Metrosideros mesic forest With Clermontia arborescens, Coprosma pubens, Broussaisia arguta, Cheirodentron trigynum, Myrsine lessertiana, Melicope clusifolia, Diplazium sandwhicianum, Melicope volcanica, Cyanea macrostegia, Cya PRESERVED_SPECIMEN Perlman, S. 21660 Maui BISH 5/27/2009
236 Melicope spathulata A.Gray Kokee, Waimea, Napali-Kona FR woods PRESERVED_SPECIMEN St. John, H. 10715 Kauai BISH 12/26/1930
237 Melicope spathulata A.Gray Kaunuohua Ecotone formation w/ mesic and wet forest elements PRESERVED_SPECIMEN Takeuchi, W.N. 3679 Kauai BISH 9/26/1987
238 Melicope spathulata A.Gray Kailua PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH 12/12/1908
239 Melicope spathulata A.Gray Awaawapuhi valley, Kainamanu N facing steep slope; Pouteria, Pittosporum, Nestegis, Cheirodendron, Scaevola procera, Coprosma kauaiensis, Bobea brevipes, Pleomele aurea, Dicranopteris linearis, Melicope anisata, Ilex, Xylosma hawaiiense, Elaeocarpus, Perrottetia, Passiflora mollissima, Myrica faya, Rubus argutus, Hedychium gardnerianum PRESERVED_SPECIMEN Perlman, S. 18660 Kauai BISH 10/22/2003
240 Melicope spathulata A.Gray S. Kona Distr., Kukuiopae Ahu., Tr 67(61); W slopes of Mauna Loa; 2.5-3 km. N. of Kaapuna 1950 lava flow Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Growing on aa. PRESERVED_SPECIMEN Warshauer, F.R. FRW 2027 Hawaii BISH 7/26/1978
241 Melicope spathulata A.Gray East Maui, Koolau Forest Reserve, below Kula pipeline PRESERVED_SPECIMEN Munro, G.C. 786 Maui BISH 2/25/1928
242 Melicope spathulata A.Gray Kaluanui, near stream, bottom of valley PRESERVED_SPECIMEN Hume, E.P. 76 Oahu BISH 1/11/1931
243 Melicope spathulata A.Gray Konahuanui PRESERVED_SPECIMEN Forbes, C.N. s.n. Oahu BISH 3/22/1905
244 Melicope spathulata A.Gray Mohihi River, Waimea Metrosideros PRESERVED_SPECIMEN St. John, H. 19999 Kauai BISH 2/12/1939
245 Melicope spathulata A.Gray Hono O Na Pali NAR, Pohakea region above Hanakapiai Metrosideros-Dicranopteris-montane mesic to wet forest. W/ Broussaisia arguta, Kadua affinis, Melicope spp, Psychotria mariniana, Coprosma waimeae, rich fern undersotyr with Asplenium insiticeum, A. polypodon, Athyrium microphyllum, Diplazium sandwichianum PRESERVED_SPECIMEN Wood, K.R. 14374 Kauai BISH 11/18/2010
246 Melicope spathulata A.Gray Puu Makaala, Puna District PRESERVED_SPECIMEN Nagata, R. ESP 304 Hawaii BISH 12/16/1980
247 Melicope spathulata A.Gray Puu Makaala, Puna District PRESERVED_SPECIMEN Nagata, R. Collector Number: ESP 304 Hawaii BISH 12/16/1980
248 Melicope spathulata A.Gray East Maui, Makawao District, Makawao FR, Waiohiwi stream headwaters Vegetation: Acacia, Metrosideros, Cheirodendron, Myrsine, Melicope, Broussaisia, Cyrtandra, Smilax, Alyxia, Asplenium spp, Athyrium, Diplazium, Marattia, Peperomia, Hedychium, Rubus. PRESERVED_SPECIMEN Oppenheimer, H.L. Nielsen, J. Collector Number: H20606 Maui BISH 2/7/2006
249 Melicope spathulata A.Gray SW ridge, Pihea, Waimea wet forest PRESERVED_SPECIMEN St. John, H. 22932 Kauai BISH 12/22/1947
250 Melicope spathulata A.Gray S. Hilo Dist., Paukaa Ahu. Tr 21(45); E slope of Mauna Kea, on ridge betwn Nukupahu Gulch/Awehi Stream PRESERVED_SPECIMEN Warshauer, F.R. FRW 1389 Hawaii BISH 9/11/1977
251 Melicope spathulata A.Gray Pahole Gulch PRESERVED_SPECIMEN Stemmermann, R.L. 1139 Oahu BISH 8/8/1976
252 Melicope spathulata A.Gray Glenwood forests PRESERVED_SPECIMEN Rock, J.F.C. s.n. Hawaii BISH 12/14/1914
253 Melicope spathulata A.Gray Kumuwela PRESERVED_SPECIMEN Rice, R.C.A. s.n. Kauai BISH 7/16/1976
254 Melicope spathulata A.Gray East Maui, Olinda PRESERVED_SPECIMEN Rock, J.F.C. 8534 Maui BISH 9/1/1910
255 Melicope spathulata A.Gray Waimea PRESERVED_SPECIMEN Faurie, A. 245 Kauai BISH 2/1/1910
256 Melicope spathulata A.Gray Alakai Swamp Trail, 1/4 mile past junction with Pihea Trail Single tree, PRESERVED_SPECIMEN Flynn, T. 432 Kauai BISH 6/2/1983
257 Melicope spathulata A.Gray back of Kaholuamanu woods PRESERVED_SPECIMEN Rock, J.F.C. 5969 Kauai BISH 9/4/1909
258 Melicope spathulata A.Gray S. Opaeula Ridge PRESERVED_SPECIMEN Suehiro, A. 137 Oahu BISH 9/25/1932
259 Melicope spathulata A.Gray Waianae Mts, Peacock Flats Trail, .3 mi before Cabin, 2100 ft elev, nr trail In mesic forest w/Coprosma, Hibiscus, Antidesma, kukui, Psychotria, Ohia, Guava, Schinus plants PRESERVED_SPECIMEN Perlman, S. 5798 Oahu BISH 4/30/1987
260 Melicope spathulata A.Gray Olaa Flume PRESERVED_SPECIMEN Forbes, C.N. 652.H Hawaii BISH 6/2/1915
261 Melicope spathulata A.Gray Kaholuamana PRESERVED_SPECIMEN Forbes, C.N. 328 K Kauai BISH 9/1/1909
262 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 10226 Kauai BISH 10/1/1911
263 Melicope spathulata A.Gray Kokee State Park, along Pihea Train from Puu o Kila to jct. with Alakai Swamp Trail PRESERVED_SPECIMEN Wagner, W.L. 5036 Kauai BISH 8/30/1983
264 Melicope spathulata A.Gray Honolulu District. Aiea trail, near summit, between Kaivipoo hill and large power relay Metrosidero-Dicranopteris linearis lowland wet forest. With Syzgium sandwicense, Machaerina angustifolia, Ilex anomala, Melicope oahuensis, Zanthoxylum oahuense, Antidesma platyphylla, Pritcardia martii, Clidemia hirta, Psidium cattleianum PRESERVED_SPECIMEN Perlman, S. 23644 Oahu BISH 8/20/2013
265 Melicope spathulata A.Gray along Punaluu (Castle) Trail, Koolau Mts In wet Metrosideros forest, with Sadleria cyatheoides, Pritchardia martii, Cyrtandra, Stenogyne, Scaevola mollis, Dicranopteris emarginata, Euphorbia rockii, Coprosma longifolia, Pittosporum dolosum, Hibiscus arnottianus. PRESERVED_SPECIMEN Carlquist, S. 1745 Oahu BISH 7/6/1965
266 Melicope spathulata A.Gray Mt. Olympus PRESERVED_SPECIMEN Rock, J.F.C. s.n. Oahu BISH 11/30/1912
267 Melicope spathulata A.Gray Forest along N. Border of Ewa Forest Reserve Forest PRESERVED_SPECIMEN Christophersen, E. 1608 Oahu BISH 2/20/1931
268 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 1968 Kauai BISH 3/3/1909
269 Melicope spathulata A.Gray Wainiha, upper NE fork Closed Metrosideros forest, 8-12 m canopy. Understory dominated by Antidesma w/ mixed wet associates, Syzygium, Broussaisia, Perrottetia, Cyrtandra spp, Psychotria spp, Dubautia spp, Labordia spp, Coprosma waimeae, Cheirodendron spp, Tetraplasandra PRESERVED_SPECIMEN Wood, K.R. 13362 Kauai BISH 9/10/2008
270 Melicope spathulata A.Gray Kohala Mountains PRESERVED_SPECIMEN Lamoureux, C.H. 2748 Hawaii BISH 6/18/1964
271 Melicope spathulata A.Gray PRESERVED_SPECIMEN h. mann, w. t. brigham 94 GH
272 Melicope spathulata A.Gray Olaa Forest Reserve, behind Volcano dump PRESERVED_SPECIMEN Wagner, W.L. 5273 Hawaii BISH 2/28/1984
273 Melicope spathulata A.Gray Piko Trail, on ridge between Kukuiala Gulch and 2nd gulch from W. end of Makuleia, Mokuleia, Mokuleia Forest Reserve Moist woods PRESERVED_SPECIMEN St. John, H. 14094 Oahu BISH 2/4/1934
274 Melicope spathulata A.Gray Kohua Ridge forest PRESERVED_SPECIMEN Degener, O. 23919 Kauai BISH 9/7/1955
275 Melicope spathulata A.Gray Kawailoa Very wet rainforest PRESERVED_SPECIMEN Bryan Jr., E.H. 867 Oahu BISH 11/20/1934
276 Melicope spathulata A.Gray Kohala high plateau in dense forest PRESERVED_SPECIMEN Rock, J.F.C. s.n. Hawaii BISH 3/20/1957
277 Melicope spathulata A.Gray Pupukea-Kahuku trail, Kaumala-Waimea Rain forest PRESERVED_SPECIMEN Baker, R.J. s.n. Oahu BISH 2/19/1933
278 Melicope spathulata A.Gray Along road below cabin, at Portuguese Gate. Along crest of west branch of Honokane Cibotium-Metrosideros-Cheirodendron forest with Clermontia, Pelea, Cyrtandra, Gouldia, Straussia PRESERVED_SPECIMEN Gillett, G.W. 1894 Hawaii BISH 6/25/1966
279 Melicope spathulata A.Gray Waimea, Alakai, between Lehua makanoe and Kilohana PRESERVED_SPECIMEN Cranwell, L.M. 2938 Kauai BISH 8/16/1938
280 Melicope spathulata A.Gray Koolau Mts., Moanalua Valley, summit crest ridge In Metrosideros-Dicranopteris linearis lowland wet forest with Antidesma, Clermontia, Cyrtandra, Zanthoxylum, Myrsine, Melicope, Pritchardia, Psychotria, Tetraplasandra, Cheirodendron, Bobea. PRESERVED_SPECIMEN Perlman, S. 5942 Oahu BISH 6/7/1987
281 Melicope spathulata A.Gray Crest of Koolau Mts. between Puu Keahiakahoe and Puu Kahuauli scattered Clidemia. PRESERVED_SPECIMEN Nagata, K.M. 2596 Oahu BISH 2/9/1983
282 Melicope spathulata A.Gray Awaawapuhi trail. Honopu Rain forest PRESERVED_SPECIMEN Woolford, H.A. 423 Kauai BISH 12/29/1956
283 Melicope spathulata A.Gray Kipahulu, W part of valley PRESERVED_SPECIMEN Lamoureux, C.H. 4043 Maui BISH 8/10/1967
284 Melicope spathulata A.Gray Koolau Summit Trail between Laie Summit and Kahuku cabin PRESERVED_SPECIMEN Stemmermann, R.L. 1200 Oahu BISH 10/30/1976
285 Melicope spathulata A.Gray Vicinity forest ranger station at Kokee, transition between wet and dry forest near 3500 ft altitutde: Pipeline trail PRESERVED_SPECIMEN Stern, W.L. 1307 Kauai BISH 8/1/1961
286 Melicope spathulata A.Gray Punaluu Mts PRESERVED_SPECIMEN Rock, J.F.C. 841 Oahu BISH 12/24/1908
287 Melicope spathulata A.Gray East Maui, below Kula pipe line PRESERVED_SPECIMEN Munro, G.C. 796 Maui BISH 2/22/1928
288 Melicope spathulata A.Gray Waipio-Waiawa Ridge PRESERVED_SPECIMEN MacDaniels, L.H. 16 Oahu BISH 10/6/1926
289 Melicope spathulata A.Gray East Maui, Hana District, Waihoi, base of N wall stream, under Uluhe, Perrottetia PRESERVED_SPECIMEN Harrison, B. 89 Maui BISH 7/12/1972
290 Melicope spathulata A.Gray North facing slope of Haleanau gulch, Schofield Barracks Military Reservation West Range, Waianae Mt. Range Vegetation: Athyrium, Charpentiera, Cyrtandra, Metrosideros, Pisonia and Rubus. PRESERVED_SPECIMEN Higashino, P.K. 5589 Oahu BISH 2/27/1977
291 Melicope spathulata A.Gray Koolau, ridge above Kahana Bay PRESERVED_SPECIMEN Skottsberg, C.J.F. 1853 Oahu BISH 9/17/1926
292 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 5339 Kauai BISH 9/1/1909
293 Melicope spathulata A.Gray East Maui, Makawao District, Makawao FR, Waiohiwi stream headwaters Vegetation: Acacia, Metrosideros, Cheirodendron, Myrsine, Melicope, Broussaisia, Cyrtandra, Smilax, Alyxia, Asplenium spp, Athyrium, Diplazium, Marattia, Peperomia, Hedychium, Rubus. PRESERVED_SPECIMEN Oppenheimer, H.L. H20606 Maui BISH 2/7/2006
294 Melicope spathulata A.Gray Hualalai, summit? PRESERVED_SPECIMEN Rock, J.F.C. 3800? Hawaii BISH 6/9/1909
295 Melicope spathulata A.Gray S. Hilo Dist., Paukaa Ahu., Tr 21(106) E slope of Mauna Kea, on ridge betwn Honolii & Pohakupaa Streams Wet koa-`ohi`a forest with other native trees and a treefern, native shrub understory. PRESERVED_SPECIMEN Warshauer, F.R. FRW 1403 Hawaii BISH 9/13/1977
296 Melicope spathulata A.Gray Kawaihau District, ridge, w. side of Makaleha Mts Shaded, wet N. facing slope. PRESERVED_SPECIMEN Gagne, W.C. 582 Kauai BISH 9/4/1970
297 Melicope spathulata A.Gray Pupukea-Kahuku Trail, Koolau Range, Malaekahana-Waimea ridge Wet woods PRESERVED_SPECIMEN St. John, H. 11574 Oahu BISH 3/6/1932
298 Melicope spathulata A.Gray Hanalei to Lihue, power line trail PRESERVED_SPECIMEN Forbes, C.N. s.n. Kauai BISH 8/6/1909
299 Melicope spathulata A.Gray Kokee - Kumuwela, in mesic forest PRESERVED_SPECIMEN Stone, B.C. 3358 Kauai BISH 4/13/1960
300 Melicope spathulata A.Gray Olinda, along pipe-line trail In dark forest PRESERVED_SPECIMEN Degener, O. 8615 Maui BISH 7/29/1927
301 Melicope spathulata A.Gray Honopu: Awaawapuhi Valley In mesic forest PRESERVED_SPECIMEN Stone, B.C. 1609 Kauai BISH 12/29/1956
302 Melicope spathulata A.Gray Wahiawa-Kohana trail PRESERVED_SPECIMEN Forbes, C.N. 2202.O Oahu BISH 8/17/1915
303 Melicope spathulata A.Gray W bank of Honomanu trail PRESERVED_SPECIMEN Forbes, C.N. 2587 M Maui BISH 6/23/1920
304 Melicope spathulata A.Gray Near Glenwood PRESERVED_SPECIMEN Rock, J.F.C. s.n. Hawaii BISH 7/1/1918
305 Melicope spathulata A.Gray Kokee - Kumuwela, in mesic forest Kokee-Kamuwela, in mesic forest. PRESERVED_SPECIMEN Stone, B.C. 3358 Kauai BISH 4/13/1960
306 Melicope spathulata A.Gray Peahinaia ridge PRESERVED_SPECIMEN Stemmermann, R.L. 1271 Oahu BISH 11/14/1976
307 Melicope spathulata A.Gray Waikane-Schofield Trail, Koolau Range, Waianae Uka Thicket on divide PRESERVED_SPECIMEN St. John, H. 17626 Oahu BISH 4/28/1935
308 Melicope spathulata A.Gray Waioli, hanging valley, N face of Namolokama, prominent ridge to E of main falls windswept; Metrosideros, Cheirodendron, Sticherus, Diplopterygium, Dicranopteris, Psychotria mariniana, P. hexandra, Dubautia laxa, D. knudsenii, Bidens campylotheca, Freycinetia arborea, Sadleria pallida, Hedyotis tryblium PRESERVED_SPECIMEN Wood, K.R. 12186 Kauai BISH 10/18/2006
309 Melicope spathulata A.Gray Waimea District, Kohua Ridge North facing slopes above Awini Stream Metrosideros-Acacia montane mesic forest with Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena aurea, Polyscias Kavaiensis, Planch PRESERVED_SPECIMEN Wood, K.R. 14975 Kauai BISH 5/16/2012
310 Melicope spathulata A.Gray Within a kipuka near the Puu Makaala access road, Upper Waiakea Forest Feserve, S. Hilo District Open to closed Metrosideros-Cibotium forest PRESERVED_SPECIMEN Clarke, G. 636 Hawaii BISH 10/27/1981
311 Melicope spathulata A.Gray Waimea District. Koaie, headwater gulches and stream, well above Koaie river, north of Mohihi-Waialae trail. Metrosideros transitional mesic to wet forest and riparian vegetation w/ Cheirodendron, Perrottetia sandwicensis, Kadua affinis, Ilex anomala, Syzygium sandwicensis, Dodonaea viscosa, Coprosma waimeae, Cyrtandra kauaiensis, Melicope clusiifolia, Labordia degeneri, Cyane PRESERVED_SPECIMEN Wood, K.R. 15963 Kauai BISH 6/24/2014
312 Melicope spathulata A.Gray Koolau Mts.. N. Kaliuwaa PRESERVED_SPECIMEN Rock, J.F.C. 65 Oahu BISH 8/1/1908
313 Melicope spathulata A.Gray S. Opaeula Ridge PRESERVED_SPECIMEN Suehiro, A. s.n. Oahu BISH 9/25/1932
314 Melicope spathulata A.Gray Kokee, W rim, Kalalau Valley dry; Hedyotis acuminata, Pelea anisata, Acacia koa, Metrosideros, Dodonaea, Rubiaceae PRESERVED_SPECIMEN Herbst, D.R. 1198 Kauai BISH 8/10/1968
315 Melicope spathulata A.Gray Alakai Swamp trail, Kawaikoi Stream, from Lehua makanoe PRESERVED_SPECIMEN Takeuchi, W.N. Alakai 19 Kauai BISH 9/25/1982
316 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 5645 Kauai BISH 9/4/1909
317 Melicope spathulata A.Gray Alakai Swamp, nr Pihea Shallow gully PRESERVED_SPECIMEN Takeuchi, W.N. 2464 Kauai BISH 9/21/1985
318 Melicope spathulata A.Gray Kauai shady PRESERVED_SPECIMEN Hobdy, R.W. 16 Kauai BISH 5/21/1905
319 Melicope spathulata A.Gray Back of Wahiawa PRESERVED_SPECIMEN Swezey, O.H. s.n. Oahu BISH 10/31/1920
320 Melicope spathulata A.Gray Kokee - Kumuwela, PRESERVED_SPECIMEN Stone, B.C. 3360 Kauai BISH 4/13/1960
321 Melicope spathulata A.Gray Makawao, Haiku Uka, NW Haleakala, between Haipuaena and Puohokamoa stream; with ohia, native trees and shrubs; treefern PRESERVED_SPECIMEN Higashino, P.K. 9355 Maui BISH 6/10/1980
322 Melicope spathulata A.Gray Waimea District, Kohua Ridge North facing slopes above Awini Stream Metrosideros-Acacia montane mesic forest with Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena aurea, Polyscias Kavaiensis, Planch PRESERVED_SPECIMEN Wood, K.R. Query, M. Collector Number: 14975 Kauai BISH 5/16/2012
323 Melicope spathulata A.Gray Kipapa Gulch, So. ridge. Koolau Mts on wet wooded ridge PRESERVED_SPECIMEN Hosaka, E.Y. 970 Oahu BISH 4/16/1933
324 Melicope spathulata A.Gray Mt. Kaala PRESERVED_SPECIMEN Suehiro, A. 137 Oahu BISH 1/8/1933
325 Melicope spathulata A.Gray Kalalau Valley, east rim Rainforest PRESERVED_SPECIMEN Degener, O. 27150 Kauai BISH 11/16/1960
326 Melicope spathulata A.Gray Olinda forest PRESERVED_SPECIMEN Rock, J.F.C. 8534 Maui BISH 9/1/1910
327 Melicope spathulata A.Gray Alakai Swamp nr Kawaikoi Stream Under dominant Cheirodendron, Coprosma and Rubus common in S layer PRESERVED_SPECIMEN Takeuchi, W.N. 2469 Kauai BISH 9/21/1985
328 Melicope spathulata A.Gray Hanalei District; Kumuwela, gulch just S of Maluapopoki stream Metrosideros Acacia mesic forest withSantalum pyrularium, Leptecophylla tameamea, Coprosma waimeae, Dodonaea viscosa, Wikstroemia furcata, Melicope ovata, M. barbigera, Cyanea leptostegia, Claoxylon sandiwcense, Planchonella sandwic PRESERVED_SPECIMEN Wood, K.R. 1158 m Kauai BISH 9/19/2012
329 Melicope spathulata A.Gray East Maui, Hana District, central Kipahulu Valley wet koa ohia forest; with native trees, shrubs, treefern PRESERVED_SPECIMEN Higashino, P.K. 9414 Maui BISH 7/22/1980
330 Melicope spathulata A.Gray Milolii Ridge, Waimea wet forest PRESERVED_SPECIMEN St. John, H. 13774 Kauai BISH 12/27/1933
331 Melicope spathulata A.Gray Waikamoi, lower pipe line trail PRESERVED_SPECIMEN Hobdy, R.W. 2607 Maui BISH 6/28/1986
332 Melicope spathulata A.Gray Punaluu Mts.,, Along trail to Punaluu Camp and beyond PRESERVED_SPECIMEN Rock, J.F.C. 8834 Oahu BISH 8/1/1911
333 Melicope spathulata A.Gray Waimea, Alakai Swamp wet ohia, native trees and shrubs, treeferns; bank of gulch PRESERVED_SPECIMEN Higashino, P.K. 9685 Kauai BISH 5/16/1981
334 Melicope spathulata A.Gray Halemanu PRESERVED_SPECIMEN Rock, J.F.C. 1997 Kauai BISH 2/14/1909
335 Melicope spathulata A.Gray Tree along the Kawaikoi Trail just before going down the steeper switchback portion PRESERVED_SPECIMEN Corn, C. s.n. Kauai BISH 8/29/1982
336 Melicope spathulata A.Gray Kipapa Gulch. Waipio Metrosideros-Antidesma Forest PRESERVED_SPECIMEN Grant, M.L. 7252 Oahu BISH 8/7/1934
337 Melicope spathulata A.Gray Koolau Mts., Punaluu, Castle trail Forest tree on wet ridge PRESERVED_SPECIMEN Stone, B.C. 1142 Oahu BISH 9/24/1955
338 Melicope spathulata A.Gray Makawao Forest Reserve, Olinda pipe line rain forest PRESERVED_SPECIMEN St. John, H. 25625 Maui BISH 12/25/1955
339 Melicope spathulata A.Gray W Waimea drainage basin PRESERVED_SPECIMEN Forbes, C.N. 782 K Kauai BISH 7/3/1917
340 Melicope spathulata A.Gray main divide, Waikane-Schoefield trail, Koolau Range PRESERVED_SPECIMEN Hume, E.P. 537 Oahu BISH 4/17/1932
341 Melicope spathulata A.Gray Nualolo trail, Kokee, valley floor moist, shade PRESERVED_SPECIMEN Herbst, D.R. 2066 Kauai BISH 8/16/1971
342 Melicope spathulata A.Gray Kahana Trail PRESERVED_SPECIMEN Lyon, H.L. s.n. Oahu BISH
343 Melicope spathulata A.Gray Kumuwela Ridge, Waimea moist forest PRESERVED_SPECIMEN St. John, H. 13803 Kauai BISH 12/28/1933
344 Melicope spathulata A.Gray At ca. 2,800 feet elevation, on contour portion of trail beyond the stream, Punaluu (Castle) Trail, Koolau Mts In wet Metrosideros polymorpha forest, with Euphorbia rockii, Scaevola mollis, Cheirodendron, Broussaisia, Labordia, various ferns. PRESERVED_SPECIMEN Carlquist, S. 2197 Oahu BISH 8/22/1966
345 Melicope spathulata A.Gray Between W and E Wailuaiki ridge PRESERVED_SPECIMEN Forbes, C.N. 2664 M Maui BISH 7/2/1920
346 Melicope spathulata A.Gray Waimano, Waimano Trail, Koolau Range Thicket on divide in cloud zone PRESERVED_SPECIMEN St. John, H. 20959 Oahu BISH 5/13/1945
347 Melicope spathulata A.Gray Kaluanui, near stream, small valley head PRESERVED_SPECIMEN Hume, E.P. 106 Oahu BISH 3/8/1931
348 Melicope spathulata A.Gray Maui PRESERVED_SPECIMEN Ozaki, E.T. 1390 Maui BISH 1/9/1955
349 Melicope spathulata A.Gray Wailua mauka PRESERVED_SPECIMEN s.n. Oahu BISH
350 Melicope spathulata A.Gray Poamoho Trail Open, shrubby rain-forest PRESERVED_SPECIMEN Degener, O. 27377 Oahu BISH 4/30/1961
351 Melicope spathulata A.Gray W rim, Kalalau Valley dark woods PRESERVED_SPECIMEN Degener, O. 27155 Kauai BISH 11/29/1960
352 Melicope spathulata A.Gray S. Kona Distr., Kukuiopae Ahu., Tr 67(56); W slopes of Mauna Loa; 2.5-3 km. N. of Kaapuna 1950 lava flow Wet `ohi`a forest with other native trees and a treefern and native shrub understory. 1 neighboring plant, 2 plants 200 m. away; growing on aa. PRESERVED_SPECIMEN Warshauer, F.R. FRW 2024 Hawaii BISH 7/26/1978
353 Melicope spathulata A.Gray Waimea District. Kohua Ridge, north facing slopes above Mohihi Stream Metrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Chierodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracena au PRESERVED_SPECIMEN Wood, K.R. 15091 Kauai BISH 7/12/2012
354 Melicope spathulata A.Gray First N for, of Wahiawa Stream, NW of Wahiawa Bog dense wet forest with moderate to heavy Psidium growth, except upper part of valley dominated by Metrosideros with understory of Psychotria, Hedyotis terminalis, Cyanea (3 species), Dicranopteris (in open areas), 3 spp. of Cyrtandra, Labordia hirtella (in upper part). PRESERVED_SPECIMEN Wagner, W.L. 6060 Kauai BISH 4/10/1988
355 Melicope spathulata A.Gray Wailuku Forest Reserve PRESERVED_SPECIMEN Adee, K.T. s.n. Hawaii BISH 6/7/1979
356 Melicope spathulata A.Gray East Maui, Makawao, Kalialinui, between Waiohiwi and Kahakapo gulches ridge, with koa, ohia, native fern and shrub, treefern PRESERVED_SPECIMEN Warshauer, F.R. s.n. Maui BISH 6/28/1980
357 Melicope spathulata A.Gray Honopu: Awaawapuhi Valley PRESERVED_SPECIMEN Lane, I.E. 56-614 Kauai BISH 12/29/1956
358 Melicope spathulata A.Gray Waimea District. Nuololo, north facing slopes above northern branch Metrosideros-Acacia montane mesic forest with Alphitonia, Pouteria, Xylosma hawaiiensis, Claoxylon, Mysine laniensis, Wikstroemia furcata, Dodonaea, Kadua affinis, Melicope ovata, M. anisata, M. barbigera, Dracaena aurea, Plyscias kavaien. PRESERVED_SPECIMEN Wood, K.R. 15104 Kauai BISH 7/1/2012
359 Melicope spathulata A.Gray below Kokee and Kilohana PRESERVED_SPECIMEN Skottsberg, C.J.F. 981 Kauai BISH 10/27/1922
360 Melicope spathulata A.Gray Waiawa top of Koolau range PRESERVED_SPECIMEN Russ, G.W. s.n. Oahu BISH 3/1/1930
361 Melicope spathulata A.Gray Laie Trail open area along trail PRESERVED_SPECIMEN Herbst, D.R. 1106 Oahu BISH 5/5/1968
362 Melicope spathulata A.Gray upper Awaawapuhi Valley diverse mesic forest; Melicope haupuensis, Dodonaea viscosa, Psychotria greenwelliae, Metrosideros polymorpha, Cryptocarya mannii, Nothocestrum peltatum, Rubus argutus PRESERVED_SPECIMEN Trauernicht, C. 605 Kauai BISH 11/11/2008
363 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 1978 Kauai BISH 3/3/1909
364 Melicope spathulata A.Gray Honomanu Trail PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH 5/1/1911
365 Melicope spathulata A.Gray near Puu Kainapuaa, Koolau mts shrub in Clidemia PRESERVED_SPECIMEN Takeuchi, W.N. s.n. Oahu BISH 12/1/1985
366 Melicope spathulata A.Gray Wahiawa Mts PRESERVED_SPECIMEN Forbes, C.N. 229 Kauai BISH 8/1/1909
367 Melicope spathulata A.Gray Koolau Mts., Bryan's Mtn House Trail at Grant Boundry PRESERVED_SPECIMEN Krajina, V.J. 620520055 Oahu BISH 5/20/1962
368 Melicope spathulata A.Gray Kipapa-Waiawa Ridge. Waipio Dist Metrosideros-Acacia Forest PRESERVED_SPECIMEN Grant, M.L. 7248 Oahu BISH 8/7/1934
369 Melicope spathulata A.Gray Awaawapuhi trail, Honopu district PRESERVED_SPECIMEN Lane, I.E. s.n. Kauai BISH 12/1/1956
370 Melicope spathulata A.Gray NW Alakai swamp, Napali-Kona FR woods PRESERVED_SPECIMEN St. John, H. 10737 Kauai BISH 12/27/1930
371 Melicope spathulata A.Gray back of Kaholuamanu woods PRESERVED_SPECIMEN Rock, J.F.C. 6020 Kauai BISH 9/4/1909
372 Melicope spathulata A.Gray NW Haleakala, ridge W of Opana Gulch, Koolau Forest Reserve wet koa ohia forest with native trees and shrubs; treeferns PRESERVED_SPECIMEN Warshauer, F.R. FRW 2736 Maui BISH 7/28/1980
373 Melicope spathulata A.Gray Honomu, Papaikou not common PRESERVED_SPECIMEN Bryan, L.W. s.n. Hawaii BISH 3/22/1937
374 Melicope spathulata A.Gray Kaholuamanu PRESERVED_SPECIMEN Rock, J.F.C. 5344 Kauai BISH 9/4/1909
375 Melicope spathulata A.Gray Koolau Range, Kahuku-Kawailoa For.Res., Puu Kainapuaa wind-swept crest of ridge PRESERVED_SPECIMEN Suehiro, A. Not on sheet Oahu BISH 11/23/1941
376 Melicope spathulata A.Gray Kokee-Kumuwela plateau above Waimea, 3600-3900 ft PRESERVED_SPECIMEN Stone, B.C. 3395 Kauai BISH 4/14/1960
377 Melicope spathulata A.Gray Halemanu PRESERVED_SPECIMEN Rock, J.F.C. 2298 Kauai BISH 2/14/1909
378 Melicope spathulata A.Gray Kalalau Trail PRESERVED_SPECIMEN Forbes, C.N. 1046 K Kauai BISH 7/3/1917
379 Melicope spathulata A.Gray Alakai Swamp, nr junction Pihea trail and Alakai Swamp trail PRESERVED_SPECIMEN Takeuchi, W.N. 2465 Kauai BISH 9/21/1985
380 Melicope spathulata A.Gray land of Olaa, Olaa Forest Reserve, 2/3 mile WSW of Puu Makaala in wet, dense Metrosideros-fern forest. PRESERVED_SPECIMEN Herbst, D.R. 5507 Hawaii BISH 10/25/1975
381 Melicope spathulata A.Gray Kokee; Awaawapuhi train, Honopu Distr moist mesic-wet forest at 3500 ft., of Acacia Koa, Straussia, Fagara, Claoxylon, Pelea, etc. PRESERVED_SPECIMEN Stone, B.C. 3330 Kauai BISH 4/12/1960
382 Melicope spathulata A.Gray W Waimea drainage basin PRESERVED_SPECIMEN Forbes, C.N. 911 K Kauai BISH 7/3/1917
383 Melicope spathulata A.Gray L side of Kipahulu Valley ridge PRESERVED_SPECIMEN Forbes, C.N. 1642 M Maui BISH 11/15/1919
384 Melicope spathulata A.Gray Pauoa Oahu PRESERVED_SPECIMEN s.n. Oahu BISH
385 Melicope spathulata A.Gray Kawailoa In moist rainforest PRESERVED_SPECIMEN Bryan Jr., E.H. 814 Oahu BISH 10/2/1934
386 Melicope spathulata A.Gray Alakai Swamp, Kokee open forest PRESERVED_SPECIMEN MacDaniels, L.H. 781 Kauai BISH 2/18/1927
387 Melicope spathulata A.Gray Laie at 2075' elev stature = 10' in small gully with Antidesma platyphyllum, Psychotria mariniana, and Psidium cattleianum PRESERVED_SPECIMEN Takeuchi, W.N. 1980 Oahu BISH 2/3/1985
388 Melicope spathulata A.Gray Kumuwela Ridge, Waimea moist forest PRESERVED_SPECIMEN St. John, H. 13819 Kauai BISH 12/28/1933
389 Melicope spathulata A.Gray Maakua-Papali ridge, Kaipapau ; Forest Reserve Hauula Moist woods PRESERVED_SPECIMEN St. John, H. 13367 Oahu BISH 10/15/1933
390 Melicope spathulata A.Gray E bank of Honomanu gulch PRESERVED_SPECIMEN Forbes, C.N. 2568 M Maui BISH 6/21/1920
391 Melicope spathulata A.Gray Laie sporadic occurrence at elevations above the Gap; most plants sterile PRESERVED_SPECIMEN Takeuchi, W.N. 2202 Oahu BISH 9/19/1985
392 Melicope spathulata A.Gray Halemanu PRESERVED_SPECIMEN Rock, J.F.C. 2318 Kauai BISH 2/14/1909
393 Melicope spathulata A.Gray Along the Kawaikoi Trail just before the steeper switchback portion Other plants seen slong this portion of the steep trail. PRESERVED_SPECIMEN Corn, C. s.n. Kauai BISH 8/29/1982
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