img

Hoiokula sandwicensis

(Brack.) S.E.Fawc. & A.R.Sm. (2021)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Thelypteridaceae Genus: Hoiokula

hō‘i‘o kula [hoio kula]

img

Bibliography

Name Published In: Fawcett, S., & Smith, A. R. A Generic Classification of the Thelypteridaceae p.49 (2021)

Other References

Palmer 2003:214 (K, O, Mo, L, M, H [as Pneumatopteris sandwicensis]); Palmer 2005:81 (KEY); Snow et al. 2011:15 (Syn. P. sandwicensis = Cyclosorus sandwicensis); Pteridophyte Phylogeny Group 2016:585 (Syn. C. sandwicensis = P. sandwicensis); Fawcett & Smith 2021:49 (COMBNOV, Syn. P. sandwicensis = H. sandwicensis)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Hoiokula sandwicensis W Maui, Lahaina District, idge N of Honokawai-Amalu valley, Haelaau Preserved_Specimen Cranwell, L.M. 2787 Maui BISH 1938-08-02
2 Hoiokula sandwicensis unknown Preserved_Specimen Rock, J.F.C. BISH
3 Hoiokula sandwicensis unknown Preserved_Specimen BISH
4 Hoiokula sandwicensis Kuhiwa Stream Preserved_Specimen Hobdy, R.W. 2909 Maui BISH 1987-07-28
5 Hoiokula sandwicensis West Maui Preserved_Specimen Mann, H. 476 Maui BISH 1864-01-01
6 Hoiokula sandwicensis Waiapaa Preserved_Specimen Munro, G.C. 74 Lanai BISH 1913-09-06
7 Hoiokula sandwicensis on Kaholuamanoa above Waimea Preserved_Specimen Heller, A.A. 2839 Kauai BISH 1895-09-10
8 Hoiokula sandwicensis Pauoa-Konahuanui trail Preserved_Specimen Garber, D.W. 302 Oahu BISH 1920-03-14
9 Hoiokula sandwicensis near Anuhea GC, Kilauea Preserved_Specimen Degener, O. 3817 Hawaii BISH 1930-02-02
10 Hoiokula sandwicensis Kahanui Preserved_Specimen Krajina, V.J. 620612071 Molokai BISH 1962-06-12
11 Hoiokula sandwicensis W ridge, Kipapa Gulch, Waipio, Koolau Mts Preserved_Specimen Hosaka, E.Y. 1132 Oahu BISH 1933-08-06
12 Hoiokula sandwicensis on Kaholuamanoa above Waimea Preserved_Specimen Heller, A.A. 2839 Kauai BISH 1895-09-10
13 Hoiokula sandwicensis on Kaholuamanoa above Waimea Preserved_Specimen Heller, A.A. 2839 Kauai BISH 1895-09-10
14 Hoiokula sandwicensis Kona or Kilauea Preserved_Specimen Lack, F. Hawaii BISH 1880-01-01
15 Hoiokula sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
16 Hoiokula sandwicensis NW of Olaa, Hiulani forest Preserved_Specimen Skottsberg, C.J.F. 419 Hawaii BISH 1922-09-08
17 Hoiokula sandwicensis Kilauea, in Twin Crater Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-01
18 Hoiokula sandwicensis Koolau Mts. between Punaluu and Kaipapau Preserved_Specimen Forbes, C.N. Oahu BISH 1909-05-03
19 Hoiokula sandwicensis Hawaii Preserved_Specimen U.S. Exploring Expedition Hawaii BISH 1840-01-01
20 Hoiokula sandwicensis Koolau Mts. between Punaluu and Kaipapau Preserved_Specimen Forbes, C.N. Oahu BISH 1909-05-03
21 Hoiokula sandwicensis Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
22 Hoiokula sandwicensis Punaluu Preserved_Specimen Rock, J.F.C. Oahu BISH 1908-08-01
23 Hoiokula sandwicensis Waianae Mts., Makaha Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
24 Hoiokula sandwicensis Koolau Mts. between Punaluu and Kaipapau Preserved_Specimen Forbes, C.N. Oahu BISH 1908-11-14
25 Hoiokula sandwicensis Waiakoali Preserved_Specimen Neal, M.C. Kauai BISH 1930-08-22
26 Hoiokula sandwicensis unknown Preserved_Specimen (Not on sheet) BISH
27 Hoiokula sandwicensis Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-22
28 Hoiokula sandwicensis Kauai Preserved_Specimen Forbes, C.N. Kauai BISH
29 Hoiokula sandwicensis Kokee stream Preserved_Specimen Wiebke, H. Kauai BISH 1926-06-30
30 Hoiokula sandwicensis Kaiholena Preserved_Specimen Munro, G.C. Lanai BISH 1915-10-17
31 Hoiokula sandwicensis Glenwood Preserved_Specimen Fullaway, D. Hawaii BISH 1920-01-01
32 Hoiokula sandwicensis South Hilo District, Waiakea. Buffer zone collections along Stainback Highway Preserved_Specimen Funk, E.J. Hawaii BISH 1998-12-15
33 Hoiokula sandwicensis Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-22
34 Hoiokula sandwicensis Hawaii Preserved_Specimen Ruhle Hawaii BISH 1956-01-01
35 Hoiokula sandwicensis Pauoa-Konahuanui trail Preserved_Specimen Garber, D.W. 301 Oahu BISH 1920-03-14
36 Hoiokula sandwicensis Mts. near Koele Preserved_Specimen Forbes, C.N. 64 Lanai BISH 1913-06-01
37 Hoiokula sandwicensis W ridge, Kipapa Gulch, Waipio, Koolau Mts Preserved_Specimen Hosaka, E.Y. 1153 Oahu BISH 1933-08-06
38 Hoiokula sandwicensis East Maui, Manawainui Planeze, below Ohia Camp and fence in gulch Preserved_Specimen Welton, P. 2173015 Maui BISH 2003-08-28
39 Hoiokula sandwicensis on Kaholuamanoa above Waimea Preserved_Specimen Heller, A.A. 2839 Kauai BISH 1895-09-10
40 Hoiokula sandwicensis S ridge, S. Opaeula Gulch, Koolau Mts., Paalaa Preserved_Specimen St.John, H. 12086 Oahu BISH 1932-09-25
41 Hoiokula sandwicensis Lanai Preserved_Specimen Forbes, C.N. 372 Lanai BISH 1917-09-01
42 Hoiokula sandwicensis near Anuhea GC, Kilauea Preserved_Specimen Degener, O. 3817 Hawaii BISH 1930-02-02
43 Hoiokula sandwicensis Honokakau drainage basin Preserved_Specimen Forbes, C.N. 400 Maui BISH 1917-09-25
44 Hoiokula sandwicensis Kuliouou, head of valley, Koolau Mts Preserved_Specimen Egler, F.E. 37108 Oahu BISH 1937-07-13
45 Hoiokula sandwicensis W ridge, Kipapa Gulch, Waipio, Koolau Mts Preserved_Specimen Hosaka, E.Y. 735 Oahu BISH 1932-09-18
46 Hoiokula sandwicensis Haalelepaakai Preserved_Specimen Munro, G.C. 173 Lanai BISH
47 Hoiokula sandwicensis Lanipo Trail Preserved_Specimen Topping, D.L. 2674 Oahu BISH 1923-07-01
48 Hoiokula sandwicensis East Maui, Koolau Gap, E side Preserved_Specimen Hobdy, R.W. 3392 Maui BISH 1991-07-18
49 Hoiokula sandwicensis Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. 11 Hawaii BISH 1929-06-29
50 Hoiokula sandwicensis Haalelepaakai Preserved_Specimen Munro, G.C. 15 Lanai BISH
51 Hoiokula sandwicensis Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. 11 Hawaii BISH 1929-06-29
52 Hoiokula sandwicensis head of Waikolu valley, pipe line trail Preserved_Specimen Cranwell, L.M. 2589 Molokai BISH 1938-08-13
53 Hoiokula sandwicensis East Maui, Manawainui Planeze, below Ohia Camp and fence in gulch Preserved_Specimen Welton, P. 2173015 Maui BISH 2003-08-28
54 Hoiokula sandwicensis Kamoku, Kaiholena Gulch Preserved_Specimen Hobdy, R.W. 2195 Lanai BISH 1984-10-22
55 Hoiokula sandwicensis Oahu Preserved_Specimen Judd, J.R. Oahu BISH 1888-01-01
56 Hoiokula sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH 1863-01-01
57 Hoiokula sandwicensis Forest of Waimea Preserved_Specimen Lyon, H.L. Hawaii BISH
58 Hoiokula sandwicensis Waiakeakua Preserved_Specimen Munro, G.C. 96 Lanai BISH 1927-06-14
59 Hoiokula sandwicensis Kalalau Lookout, Na Pali-Kona Forest Reserve Preserved_Specimen Kato, M. 18 Kauai BISH 1997-10-18
60 Hoiokula sandwicensis Koolau Mts. between Punaluu and Kaipapau Preserved_Specimen Forbes, C.N. Oahu BISH 1909-05-03
61 Hoiokula sandwicensis near Anuhea GC, Kilauea Preserved_Specimen Degener, O. 3817 Hawaii BISH 1930-02-02
62 Hoiokula sandwicensis East Maui, Koolau Forest Reserve Preserved_Specimen Gagné, B.H. 353 Maui BISH 1973-07-22
63 Hoiokula sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
64 Hoiokula sandwicensis Between Kalahuapueo and Hanalilolilo Preserved_Specimen Krajina, V.J. 620611017 Molokai BISH 1962-06-11
65 Hoiokula sandwicensis National Park tree fern forest Preserved_Specimen Degener, O. 31159 Hawaii BISH 1922-12-28
66 Hoiokula sandwicensis unknown Preserved_Specimen 64 BISH
67 Hoiokula sandwicensis 4 mi NW of Kulani Prison Camp, S. Hilo ohia woods Preserved_Specimen St.John, H. 25328 Hawaii BISH 1954-12-19
68 Hoiokula sandwicensis Kawiwi-Kaala ridge, Waianae Mts., Makaha-Waianae kai dense, moist woods Preserved_Specimen tufted St.John, H. 17574 Oahu BISH 1935-03-31
69 Hoiokula sandwicensis Kamakou, W fork of stream E of Kawela Ridge, leading to Puu o Kaeha gulch, ridge; wet forest with Metrosideros, Ilex, Xylosma, Psychotria, Syzygium Preserved_Specimen ground fern with thick, erect rhizome Lorence, D.H. 5642 Molokai BISH 1987-10-22
70 Hoiokula sandwicensis East Maui, Hana District, Waihoi Valley, N side along stream bank Preserved_Specimen Harrison, B. 2597 Maui BISH 1972-07-23
71 Hoiokula sandwicensis East Maui, Waihoi Valley, S fork of Waiohonu Stream under ledge at base of waterfall; with Peperomia, Selaginella Preserved_Specimen Harrison, B. 124 Maui BISH 1972-07-29
72 Hoiokula sandwicensis Kona, Keauhou 1 along road, ohia forest Preserved_Specimen Christ, J.H. Hawaii BISH 1958-05-15
73 Hoiokula sandwicensis N fork of Wailua River, Blue Hole, E of Mts. Waialeale and Kawaikini with 1-2 m trees, pteridophytes, Gunnera; over saturated, rocky ground Preserved_Specimen ground fern, rhizome thick, suberect Lorence, D.H. 5338 Kauai BISH 1987-07-23
74 Hoiokula sandwicensis Mt. Konahuanui, N of Honolulu light shade on ridge Preserved_Specimen Kato, M. 1057 Oahu BISH 1998-08-30
75 Hoiokula sandwicensis Chain of Craters Rd. ~7.4 mi toward Kalapana from intersection with Crater Rim Rd., up Napau Crater Trail ~1.5 mi.; ~7 m 122 deg. to R of trail Metrosideros, Cibotium forest Preserved_Specimen ~2' ferm Newell, C.L. 847 Hawaii BISH 1967-01-24
76 Hoiokula sandwicensis Kulani Prison Rd. ~10.3 mi from intersection with Kona-Hilo Hwy.; ~20 m SE of road Metrosideros, Cibotium forest Preserved_Specimen ~3' fern Newell, C.L. 727 Hawaii BISH 1966-09-01
77 Hoiokula sandwicensis Kaala, summit damp soil along path in swamp Preserved_Specimen Terrestrial. Grether, D.F. 3923 Oahu BISH 1945-10-31
78 Hoiokula sandwicensis Kau, Paauau, Keaiwa Gulch shaded moist fern forest Preserved_Specimen 3-5 ft. Russ, G.W. 26 Hawaii BISH 1927-07-12
79 Hoiokula sandwicensis Peahinaia ridge wet mesic forest Preserved_Specimen Stemmermann, R.L. 1253 Oahu BISH 1976-11-13
80 Hoiokula sandwicensis Kipapa Stream, Waipio, Koolau Mts wet gorge Preserved_Specimen St.John, H. 10018 Oahu BISH 1922-11-10
81 Hoiokula sandwicensis Wainiha Valley, side gulch on W side 3/4 mi. upstream from intake dense vegetation at base of waterfall Preserved_Specimen tuft; rhizome erect, 3 diam. St.John, H. 13963 Kauai BISH 1934-01-01
82 Hoiokula sandwicensis upper Honopu, from first Kalalau lookout, down dividing ridge with Metrosideros, Acacia, Dicranopteris Preserved_Specimen rhizome sub-erect; stipe arching; lamina dull green with brown costa; rachis yellow brown with white hairs; scales dark brown, crozier scales black brown Wood, K.R. 4965 Kauai BISH 1996-02-21
83 Hoiokula sandwicensis Kulani Trail, Olaa FR, Puna Cibotium, Metrosideros woods Preserved_Specimen scabs blackish, pinnae brittle St.John, H. 24998 Hawaii BISH 1953-01-28
84 Hoiokula sandwicensis Kipapa Gulch, Waipio Metrosideros- Antidesma forest. Preserved_Specimen Grant, M.L. 7254 Oahu BISH 1934-08-07
85 Hoiokula sandwicensis N outer slope of Haleakala, 2 mi N of Paliku Cabin, below Puu Alaea floor of rain forest; with Metrosideros, Cheirodendron, Pelea, Myrsine, Pteris, Sadleria, Athyrium, Ctenitis Preserved_Specimen rosette fern Henrickson, J.S. 3688 Maui BISH 1969-06-22
86 Hoiokula sandwicensis Haleakala, head of Kuhiwa Stream, S of Nahiku moist shady gulch, woods Preserved_Specimen St.John, H. 21055 Maui BISH 1945-08-20
87 Hoiokula sandwicensis Kipapa-Waiahole crest, Koolau Mts., Waipio Metrosideros, Cheirodendron forest Preserved_Specimen Grant, M.L. 7214 Oahu BISH 1934-08-07
88 Hoiokula sandwicensis East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, Moomoonui Stream Base of shady, moist waterfall face with Selaginella arbuscula, Vandenboschia, Ageratina riparia, Sphenomeris chinensis, Cyrtandra, Peperomia. Preserved_Specimen Imada, C. 200264 Maui BISH 2002-11-05
89 Hoiokula sandwicensis Wiamea District. Honopu, west of National Guard Station Acacia koa-Metrosideros polymorpha - with matting ferns of Dicranopteris linearis, Diplopterygium pinnatum, Sticherus owhyensis, mesic to transitional wet forest drainages with Bobea brevipes, Ilex anomala, Wikstroemia furcata, Claoxylon sandwicensis, Polyscias waimeae, Melicope anisata, M. barbigera, Kadua affinis, Psychotria greenwelliae, P. hexandra, Planchonella sandw Preserved_Specimen Terrestrial fern, along gulch, rhizome on streambank, drooping to sub-erect, 15 cm. long, fertile, uncommon. Wood, K.R. 18394 Kauai BISH 2020-04-30
90 Hoiokula sandwicensis Wiamea District. Honopu, west of National Guard Station Acacia koa-Metrosideros polymorpha - with matting ferns of Dicranopteris linearis, Diplopterygium pinnatum, Sticherus owhyensis, mesic to transitional wet forest drainages with Bobea brevipes, Ilex anomala, Wikstroemia furcata, Claoxylon sandwicensis, Polyscias waimeae, Melicope anisata, M. barbigera, Kadua affinis, Psychotria greenwelliae, P. hexandra, Planchonella sandw Preserved_Specimen Terrestrial fern, along gulch, rhizome on streambank, drooping to sub-erect, 15 cm. long, fertile, uncommon. Wood, K.R. 18394 Kauai BISH 2020-04-30
91 Hoiokula sandwicensis West Maui, Waihee tall shrubbery near end of trail Preserved_Specimen hairs light tan, fronds pale green Takeuchi, W.N. 7521 Maui BISH 1992-07-29
92 Hoiokula sandwicensis Chain of Craters Rd. ~7.4 mi toward Kalapana from intersection with Crater Rim Rd., up Napau Crater Trail ~1.5 mi.; ~7 m 122 deg. to R of trail Metrosideros, Cibotium forest Preserved_Specimen ~1' fern Newell, C.L. 846 Hawaii BISH 1967-01-24
93 Hoiokula sandwicensis Waimea District. Honopu, west of National Guard Station. Acacia koa-Metrosideros polymorpha - w/ matting ferns of Dicranopteris linearis, Diplopterygium pinnatum, Sticherus owhyhensis, mesic to transitional wet forest and drainages w/ Bobea brevipes, Ilex anomala, Wikstroemia furcata, Claoxylon sandwicensis, Polyscias waimeae, Melicope anisata, M. barbigera, Kadua affinis, Psychotria greenwelliae, P. hexandra, Planchonella sandw Preserved_Specimen Terrestrial fern, along gulch, rhizome on streambank crooping to sub-eret, 15 cm long, fertile, uncommon. Wood, K.R. 18394 Kauai BISH 2020-04-30
94 Hoiokula sandwicensis Punaluu mauka. Rainforest Preserved_Specimen Degener, O. 35019 Hawaii BISH 1970-08-15
95 Hoiokula sandwicensis Makaha Valley, Waianae Mts rain forest, on the ground Preserved_Specimen Wilson, K.A. 190 Oahu BISH 1952-11-30
96 Hoiokula sandwicensis Saddle Rd., 26 mi from Hilo, Waiakea mossy woods Preserved_Specimen St.John, H. 22399 Hawaii BISH 1946-12-26
97 Hoiokula sandwicensis East Molokai, Wailau Valley, near coast wet forest Preserved_Specimen Fosberg, F.R. 9607 Molokai BISH 1933-07-03
98 Hoiokula sandwicensis N fork of Wailua River, Blue Hole, E of Mts. Waialeale and Kawaikini with 1-2 m trees, pteridophytes, Gunnera; over saturated, rocky ground Preserved_Specimen ground fern with thick, prostrate creeping rhizome, fronds clustered Lorence, D.H. 5318 Kauai BISH 1987-07-23
99 Hoiokula sandwicensis Kilauea tree fern forest Preserved_Specimen Degener, O. 31157 Hawaii BISH 1922-12-15
100 Hoiokula sandwicensis Kilauea Settlement, Olaa, Puna moist Metrosideros, Cibotium woods Preserved_Specimen St.John, H. 18473 Hawaii BISH 1937-12-22
101 Hoiokula sandwicensis Near upper end of Manana Stream Lush wet gulch bottom under thick Metrosideros canopy with Broussaisia, Antidesma, and Psychotria, lots of Clidemia and a variety of terrestrial and epiphytic ferns: Cibotium, Callistopteris, Sphenomeris, Elaphoglossum, Mecodium, Adenophorus, Vandenboschia, etc. Preserved_Specimen Clumping fern with fronds 3.5 ft. tall; pinnae rough to the touch. Common. Imada, C. 200444 Oahu BISH 2004-05-19
102 Hoiokula sandwicensis Kulani Prison Rd. ~10.3 mi from intersection with Kona-Hilo Hwy.; ~20 m SE of road Metrosideros, Cibotium forest Preserved_Specimen ~1' ferm Newell, C.L. 726 Hawaii BISH 1966-09-01
103 Hoiokula sandwicensis West Maui, Waihee, end of trail tall shrubbery Preserved_Specimen hairs light tan, fronds pale green Takeuchi, W.N. 7521 Maui BISH 1992-07-29
104 Hoiokula sandwicensis Kahuku, 8 mi NE of Nene Cabin Metrosideros forest Preserved_Specimen St.John, H. 26814 Hawaii BISH 1971-06-25
105 Hoiokula sandwicensis National Park, SW corner of Wilderness Area, near Wright Rd jungle Preserved_Specimen Degener, O. 31155 Hawaii BISH 1967-01-19
106 Hoiokula sandwicensis East Maui, Koolau Gap, E side pendant on wet cliffs Preserved_Specimen starved Hobdy, R.W. 3393 Maui BISH 1991-07-18
107 Hoiokula sandwicensis Kona, Keauhou, above Kahaluu FR, ~4.7 mi above lower gate along jeep road and open areas in Metrosideros forest Preserved_Specimen Herbst, D.R. 5410 Hawaii BISH 1975-07-15
108 Hoiokula sandwicensis Haleakala, head of Kuhiwa Stream, S of Nahiku moist shady gulch, woods Preserved_Specimen St.John, H. 21055 Maui BISH 1945-08-20
109 Hoiokula sandwicensis Olinda pipe line, Makawao Forest Reserve wet forest Preserved_Specimen Palmer, M.R. 101 Maui BISH 1955-12-25
110 Hoiokula sandwicensis Kulani Prison Rd. ~10.3 mi from intersection with Kona-Hilo Hwy.; ~20 m SE of road Metrosideros, Cibotium forest Preserved_Specimen ~3' fern Newell, C.L. 728 Hawaii BISH 1966-09-01
111 Hoiokula sandwicensis Mapulehu Valley rocky gorge Preserved_Specimen St.John, H. 12750 Molokai BISH 1932-12-28
112 Hoiokula sandwicensis E. Maui, east Kopili'ula Acacia-Metrosideros, 50% open 7m canopy, understory of uluhe mat ferns dissected by riparian veg with 'ohia & Cheirodendron, understory of Ilex, Clermontia arborescens, Scaevola chamissoniana, Myrisne lessertiana, Melicope spp, Hedyotis spp, Coprosma spp, Perrottetia, and Broussaisia, shrubs of Vaccinium, Styphelia, Cyrtandra, and Rubus hawaiensis, ferns and bryophytes are common Preserved_Specimen lithophytic rhizome sub-erect 15 cm long x 15 cm wide, fronds arching out, scales at base of stipes broad dark brown, appressed against stipe, pinnae with 6-7 pairs of veins, not all uniting, very common Wood, K.R. 11290 Maui BISH 2005-05-03
113 Hoiokula sandwicensis Laie shaded mossy slopes with Phyllostegia Preserved_Specimen Takeuchi, W.N. 2617 Oahu BISH 1986-01-01
114 Hoiokula sandwicensis Kona, Keauhou 1 ohia rain forest Preserved_Specimen Christ, J.H. Hawaii BISH 1958-05-15
115 Hoiokula sandwicensis W Maui, E rim of Ukumehame Gulch moist Metrosideros forest Preserved_Specimen St.John, H. 25676 Maui BISH 1955-12-28
116 Hoiokula sandwicensis S ridge, S. Opaeula Gulch, Koolau Mts., Paalaa wet gulch Preserved_Specimen St.John, H. 12066 Oahu BISH 1932-09-25
117 Hoiokula sandwicensis N fork of Wailua River, Blue Hole, E of Mts. Waialeale and Kawaikini with 1-2 m trees, pteridophytes, Gunnera; over saturated, rocky ground Preserved_Specimen ground fern, rhizome thick, suberect Lorence, D.H. 5366 Kauai BISH 1987-07-23
118 Hoiokula sandwicensis Waikolu Valley, below Puu Kaeo wet forest, valley wall Preserved_Specimen St.John, H. 23472 Molokai BISH 1948-12-22
119 Hoiokula sandwicensis Waianae Mts., Kaala stream bank on steep E slope Preserved_Specimen St.John, H. 10046 Oahu BISH 1929-11-24
120 Hoiokula sandwicensis Kahikinui, Manawaimui gulch pendant on cliff faces Preserved_Specimen gracile Hobdy, R.W. 3024 Maui BISH 1989-02-28
121 Hoiokula sandwicensis W Maui, E rim of Ukumehame Gulch moist Metrosideros forest Preserved_Specimen St.John, H. 25676 Maui BISH 1955-12-28
122 Hoiokula sandwicensis Peahinaia ridge wet mesic forest Preserved_Specimen Stemmermann, R.L. 1238 Oahu BISH 1976-11-13
123 Hoiokula sandwicensis Puu Kaala, Waianae Mts, Waianaeuka Wet forest Preserved_Specimen F.R. Fosberg 10366 Oahu BISH 1933-05-11
124 Hoiokula sandwicensis Puu Hualalai, Puuwaawaa moist woods Preserved_Specimen St.John, H. 11374 Hawaii BISH 1931-12-29
125 Hoiokula sandwicensis Chain of Craters Rd. ~7.4 mi toward Kalapana from intersection with Crater Rim Rd., up Napau Crater Trail ~1.5 mi.; ~7 m 122 deg. to R of trail Metrosideros, Cibotium forest Preserved_Specimen ~2' ferm Newell, C.L. 849 Hawaii BISH 1967-01-24
126 Hoiokula sandwicensis O'ahu Forest National Wildlife Refuge, Kipapa Trail, 2 miles from refuge boundary, S side of trail no bank in shade of overhanging mat of Dicraopteris linearis Metrosideros-Acacia-Dicranopteris wet lowland forest. Preserved_Specimen Large fern; blades ~1 m in length, coriaceous, once-pinnate with shallow lobes; fertile,s ori near costae. Uncommon. Reynolds, L.S. 217 Oahu BISH 2006-07-20
127 Hoiokula sandwicensis Hulopoe Gulch, head of, at Munro Trail dense jungle Preserved_Specimen rhizome prostrate Degener, O. 31156 Lanai BISH 1963-07-26
128 Hoiokula sandwicensis E. Maui, east Kopili'ula Acacia-Metrosideros, 50% open 7m canopy, understory of uluhe mat ferns dissected by riparian veg with 'ohia & Cheirodendron, undersory of Ilex, Clermontia arborescens, Scaevola chamissoniana, Myrsine lessertiana, Melicope spp, Hedyotis spp, Coprosma spp, Perrottetia, and Broussaisia, shrubs of Vaccinium, Styphelia, Cyrtandra and Rubus hawaiensis, ferns and bryophytes are common Preserved_Specimen lithophytic rhizome sub-erect 15 cm long x 15 cm wide, fronds arching out, scales at base of stipes broad dark brown, appressed against stipe, pinnae with 6-7 paris of veins, not all uniting, very common Wood, K.R. 11290 Maui BISH 2005-05-03
129 Hoiokula sandwicensis Lanai City, above high central ridge; forest; wet Preserved_Specimen fern in ground Stone, B.C. 873 Lanai BISH 1955-07-04
130 Hoiokula sandwicensis top of Kaala wet forest Preserved_Specimen Christophersen, E. 1744 Oahu BISH 1931-05-10
131 Hoiokula sandwicensis N fork of Wailua River, Blue Hole, E of Mts. Waialeale and Kawaikini with 1-2 m trees, pteridophytes, Gunnera; over saturated, rocky ground Preserved_Specimen ground fern, rhizome thick, suberect Lorence, D.H. 5366 Kauai BISH 1987-07-23
132 Hoiokula sandwicensis Haleakala Crater, foot of pali E of Waikekehia wet, shaded gulch Preserved_Specimen St.John, H. 17756 Maui BISH 1936-12-25
133 Hoiokula sandwicensis National Park, Napau Crater lehua, fern forest Preserved_Specimen Stone, B.C. Hawaii BISH 1959-08-07
134 Hoiokula sandwicensis East Maui, Haleakala, Naholuku at Paliku, N of Kuiki along steep gully to saddle ridge, in mesic Metrosideros-scrub forest Preserved_Specimen fronds 1 m Higashino, P.K. 944 Maui BISH 1975-06-21
135 Hoiokula sandwicensis Kulani Prison Rd. ~7.4 mi from intersection with Kona-Hilo Hwy.; ~50 m into forest at L of road Metrosideros, Cibotium, Pritchardia forest Preserved_Specimen ~3' fern Newell, C.L. 342 Hawaii BISH 1966-08-09
136 Hoiokula sandwicensis 9.5 mi below Kulani Prison camp, on road from Hilo palm grove in moist Metrosideros woods on old lava Preserved_Specimen fronds closely spaced on horizontal rhizome Fosberg, F.R. 41565 Hawaii BISH 1961-03-18
137 Hoiokula sandwicensis Chain of Craters Rd. ~7.4 mi toward Kalapana from intersection with Crater Rim Rd., up Napau Crater Trail ~1.5 mi.; ~7 m 122 deg. to R of trail Metrosideros, Cibotium forest Preserved_Specimen ~6 fern Newell, C.L. 845 Hawaii BISH 1967-01-24
138 Hoiokula sandwicensis Nuuanu, Southern Koolau Mountains. Ahupuaa of Waikiki, Moku of Kona. Along Switchback trail to Konahuanui near Kaumuhonu. Growing on side of trail in wet forest dominated by Metrosideros tremuloides, M. polymorpha, Citharexylum caudatum, Psidium cattleianum, Kadua affinis, Bobea elatior, Antidesma platyphyllum, Sadleria pallida, Vandenboschia davallioides, Leucobryum seemannii, Plagiochila, Pyrrhybryum, and others. Preserved_Specimen Terrestrial fern, 1-pinnate, with minute hairs on rachis and lamina, otherwise mostly glabrous. Dark brown lanceolate scales present at base. Thomas, M.K. 317 Oahu BISH 2022-04-03
139 Hoiokula sandwicensis Hawaii County. Puu Makaala Natural Area Reserve Site, NEON Plot PUUM_014; WGS84. MEP05/ACKO overstory. CHTR2/ILAN/MYLE2/CIGL, mid story. Ground covery mainly DRWA. With Metrosideros polymorpha, Acacia koa, Cheirodendron triggnum, Ilex anomala, Myrsine lessertiana, Cibotium glaucum, Dryopteris wallichiana. Preserved_Specimen Fruiting Donoso 4 Hawaii BISH 2019-09-07
140 Hoiokula sandwicensis Hawaii County. Puu Makaala Natural Area Reserve Site, NEON Plot PUUM_014; WGS84. MEP05/ACKO overstory. CHTR2/ILAN/MYLE2/CIGL, mid story. Ground covery mainly DRWA. With Metrosideros polymorpha, Acacia koa, Cheirodendron triggnum, Ilex anomala, Myrsine lessertiana, Cibotium glaucum, Dryopteris wallichiana. Preserved_Specimen Fruiting Donoso 4 Hawaii BISH 2019-09-07
141 Hoiokula sandwicensis West Maui, Hanaula wet understory Preserved_Specimen Hobdy, R.W. 297 Maui BISH 1978-07-25
142 Hoiokula sandwicensis W Maui, Hanaula, just E of forest reserve boundary gate Montane Ohia Mixed Wet Forest Preserved_Specimen Welton, P. 1054 Maui BISH 1991-07-13
143 Hoiokula sandwicensis Poamoho. Ahupuaa of Paalaa. Northern Koolau Mountains. Growing on vertical ledge within Dicranopteri linearis in wet forest. Area dominated by Metrosideros polymorpha, Acacia koa, Syzygium sandwicense, Psychotria mariniana, Dicranopteris linearis, Diplopterygium pinnatum, Vaccinium, and others. Preserved_Specimen Thomas, M.K. 679 Oahu BISH 2024-06-06
144 Hoiokula sandwicensis Along Munro Trail, NW of Lanaihale Dense wet forest. Preserved_Specimen Degener, O. 35017 Lanai BISH 1963-08-26
145 Hoiokula sandwicensis N fork of Wailua River, Blue Hole, E of Mts. Waialeale and Kawaikini with 1-2 m trees, pteridophytes, Gunnera; over saturated, rocky ground Preserved_Specimen ground fern with thick, prostrate creeping rhizome, fronds clustered Lorence, D.H. 5318 Kauai BISH 1987-07-23
146 Hoiokula sandwicensis Kipapa Gulch, Waipio, Koolau Mts wooded slope Preserved_Specimen Hosaka, E.Y. 1210 Oahu BISH 1933-08-08
147 Hoiokula sandwicensis Top of Kaala wet forest Preserved_Specimen Christophersen, E. 1744 Oahu BISH 1931-05-10
148 Hoiokula sandwicensis Kalaupapa; Waikolu, S, stream head of valley Preserved_Specimen medium terrestrial fern Hughes, G.D. 1099 Molokai BISH 2004-11-23
149 Hoiokula sandwicensis East Maui, Koolau Ditch, W of Kailua Preserved_Specimen ~drawf Hobdy, R.W. 597 Maui BISH 1979-05-22
150 Hoiokula sandwicensis W Maui, Hanaula, just E of forest reserve boundary gate Montane Ohia Mixed Wet Forest Preserved_Specimen Welton, P. 1054 Maui BISH 1991-07-13
151 Hoiokula sandwicensis Kipapa Gulch, Waipio, Koolau Mts wet, wooded slope Preserved_Specimen Hosaka, E.Y. 1184 Oahu BISH 1933-08-07
152 Hoiokula sandwicensis Punaluu, Castle Trail Preserved_Specimen no indusia Ozaki, E.T. 1577 Oahu BISH 1956-11-26
153 Hoiokula sandwicensis Mapulehu Valley rocky gorge Preserved_Specimen St.John, H. 12750 Molokai BISH 1932-12-28
154 Hoiokula sandwicensis West Maui, Hanaula, Pohakea gulch bottom Preserved_Specimen Hobdy, R.W. 526 Maui BISH 1979-05-10
155 Hoiokula sandwicensis W Maui, Lahaina District, Lihau Montane Ohia Mixed Wet Forest at 4000 ft with Metrosideros, Clermontia arborescens, Cheirodendron, Cibotium, Hedyotis, Coprosma, Broussaisia, Myrsine, Ilex, Sadleria pallida, Labordia hedyosmifolia, Stenogyne, Peperomia spp. and native ferns Preserved_Specimen Welton, P. 1054 Maui BISH 1991-07-13
156 Hoiokula sandwicensis Kalanilehua, Olaa. Preserved_Specimen Frond apex. Giffard, W.M. 349 Hawaii BISH 1928-07-01
157 Hoiokula sandwicensis West Maui, Hanaula Springs trail Preserved_Specimen unusually lobed immature Hobdy, R.W. 1847 Maui BISH 1983-11-06
158 Hoiokula sandwicensis Kipahulu Valley, W part Preserved_Specimen terrestrial Lamoureux, C.H. 3969 Maui BISH 1967-08-09
159 Hoiokula sandwicensis Kaimahonu Ridge, Pauoa Flats Preserved_Specimen Russell, P. 143 Oahu BISH 1937-07-26
160 Hoiokula sandwicensis Maui Preserved_Specimen robust Hobdy, R.W. 3390 Maui BISH 1991-07-18
161 Hoiokula sandwicensis Kaiholena Gulch Preserved_Specimen Hobdy, R.W. 2195 Lanai BISH 1984-10-22
162 Hoiokula sandwicensis Kalanilehua, Olaa. Preserved_Specimen Next to apex. Giffard, W.M. 349 Hawaii BISH 1928-07-01
163 Hoiokula sandwicensis East Maui, Ukulele Preserved_Specimen frond 1.66-1.54 m x 34-38 cm; pinnae 17.5 cm long; stipe 98 cm long, with brownish black scales at base, few along rachis Forbes, C.N. 922 Maui BISH 1919-07-24
164 Hoiokula sandwicensis Head of Waikolu Valley, Hanalilolilo moist woods, along stream Preserved_Specimen St.John, H. 12362 Molokai BISH 1932-12-21
165 Hoiokula sandwicensis West Maui, Hanaula Springs trail Preserved_Specimen some sori along rachis and not on veins, upper surface nearly glabrous Hobdy, R.W. 2123 Maui BISH 1984-08-30
166 Hoiokula sandwicensis Koolau summit ridge along the Niu/Waimanalo divide. Ahupuaa of Waikiki, Southern Koolau mountains. Growing in cloudy summit shrubland intermixed with Dicranopteris linearis, Macherina angustifolia, Vaccinium dentatum, Metrosideros polymoprha, Scaevola mollis, Elaphoglossum fauriei, and many bryophytes. Preserved_Specimen Thomas, M.K. 988 Oahu BISH 2025-02-06
167 Hoiokula sandwicensis Pauoa-Konahuanui trail Preserved_Specimen Garber, D.W. 302 Oahu BISH 1920-03-14
168 Hoiokula sandwicensis East Maui, Ukulele Preserved_Specimen frond 1.66-1.54 m x 34-38 cm; pinnae 17.5 cm long; stipe 98 cm long, with brownish black scales at base, few along rachis Forbes, C.N. 922 Maui BISH 1919-07-24
169 Hoiokula sandwicensis Kahanui 21.1N 157.0W Preserved_Specimen Krajina, V.J. 620612053 Molokai BISH 1962-06-12
170 Hoiokula sandwicensis near Anuhea GC, Kilauea Preserved_Specimen Degener, O. 3817 Hawaii BISH 1930-02-02
171 Hoiokula sandwicensis East Maui, Manawainui Planeze, below Ohia Camp and fence in gulch. Preserved_Specimen Welton, P. 2173016 Maui BISH 2003-08-28
172 Hoiokula sandwicensis Kilauea, Twin Crater Preserved_Specimen Smith, F.G. 41 Hawaii BISH 1929-03-18
173 Hoiokula sandwicensis Puu Huluhulu Crater Preserved_Specimen Kemp, R.M. 3 Hawaii BISH 1975-08-06
174 Hoiokula sandwicensis Hii Mts. Preserved_Specimen ~5' long fronds, pubsecent; rhizome creeping Forbes, C.N. 618 Kauai BISH 1916-10-20
175 Hoiokula sandwicensis Kalanilehua, Olaa. Preserved_Specimen Continuation to base of frond. Giffard, W.M. 349 Hawaii BISH 1928-07-01
176 Hoiokula sandwicensis N. of Puu Pala; between N. and S. Kohala 20.1N 155.7W Preserved_Specimen Krajina, V.J. 611231046 Hawaii BISH 1961-12-31
177 Hoiokula sandwicensis Wailau Trail Preserved_Specimen fronds 5-8'; spores black, stipe scaly tip Forbes, C.N. 341 Molokai BISH 1912-08-01
178 Hoiokula sandwicensis North fork Wailua River, Lihue Koloa Forest Reserve Preserved_Specimen Kato, M. 117 Kauai BISH 1997-10-28
179 Hoiokula sandwicensis Blue Hole stream drainage. Diverse, wet, open-canopied riparian vegetation. Collected on S side of N stream branch at base of steep. dripping rock face Preserved_Specimen Large fern with fronds growing in a rosette from a single base, to 140 cm long. Sparse reddish-black hairs at base of stipe. Trauernicht, C. 737 Kauai BISH 2009-02-12
180 Hoiokula sandwicensis East Maui, Koolau Gap, Ainahou Preserved_Specimen fronds and stipe hairier and more coriaceous than those at lower elevations Hobdy, R.W. 770 Maui BISH 1980-02-16
181 Hoiokula sandwicensis Olokui, Wailau, between upper forks of Wailele Stream ridge, moist forest Preserved_Specimen St.John, H. 23306 Molokai BISH 1948-02-06
182 Hoiokula sandwicensis Hii Mts. Preserved_Specimen ~5' long fronds, pubsecent; rhizome creeping Forbes, C.N. 618 Kauai BISH 1916-10-20
183 Hoiokula sandwicensis West Maui, Hanaula Springs trail Preserved_Specimen some sori along rachis and not on veins, upper surface nearly glabrous Hobdy, R.W. 2123 Maui BISH 1984-08-30
184 Hoiokula sandwicensis Hii Mts Preserved_Specimen ~5' long fronds, pubsecent; rhizome creeping Forbes, C.N. 618 Kauai BISH 1916-10-20
185 Hoiokula sandwicensis Wailau Trail Preserved_Specimen fronds 5-8'; spores black, stipe scaly tip Forbes, C.N. 341 Molokai BISH 1912-08-01
186 Hoiokula sandwicensis Kipahulu Valley, W part Preserved_Specimen terrestrial Lamoureux, C.H. 3969 Maui BISH 1967-08-09
187 Hoiokula sandwicensis Plum trail, Polipoli Preserved_Specimen Hobdy, R.W. 316 Maui BISH 1978-10-27
188 Hoiokula sandwicensis East Maui, Kula pipe line, between Waikamoi and Puohokamoa Streams Preserved_Specimen Krajina, V.J. 620417049 Maui BISH 1962-04-17
189 Hoiokula sandwicensis Lanai Preserved_Specimen Forbes, C.N. 372 Lanai BISH 1917-09-01
190 Hoiokula sandwicensis Kaiholena Gulch Preserved_Specimen Hobdy, R.W. 2195 Lanai BISH 1984-10-22
191 Hoiokula sandwicensis Koolau Mts, Punaluu Preserved_Specimen Rock, J.F.C. 221 Oahu BISH 1908-11-14
192 Hoiokula sandwicensis Kilauea; N.E. Ledge 19.4N 155.3W Preserved_Specimen Krajina, V.J. 611225003 Hawaii BISH 1961-12-25
193 Hoiokula sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
Back to Top