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Cyanea spathulata

(Hillebr.) A.Heller (1896)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Asterales Family: Campanulaceae Genus: Cyanea

hāhā [haha]

Description

Key Characters:

Growth Form: Branched shrubs 2–3 m tall.

Stems:

Roots:

Leaves: Leaves simple. Alternate. Blades narrowly oblanceolate to narrowly elliptic, blades 8–22 cm long, 1–3 cm wide. Apex obtuse to acute. Base attenuate, narrowing into a winged petiole or narrowly cuneate with non–winged petioles. Upper surfaces green, glabrous; lower surfaces pale green, glabrous or sparsely pubescent. Margins minutely callose–crenulate, often revolute. Petioles winged 1.5–3.5(–6) cm long, or non–winged petioles 5.5–12 cm long. Stipules absent.

Flowers: Flowers in inflorescences 8–10–flowered, the rachis condensed or obsolete, peduncles 15–50 mm long, pedicels 8–15 mm long. Flowers bisexual (perfect); hypanthium broadly ellipsoid to broadly obovoid, 2–3 mm long, 1.5–2.5 mm wide. Calyx lobes dentiform, 0.5–1 mm long, apex acute to acuminate. Corolla white or greenish white, sometimes tinged pale purplish, 15–30 mm long, 1.5–3 mm wide, glabrous, the tube suberect, the lobes spreading. Stamens 5, alternate with corolla lobes, connate; staminal column glabrous; anthers glabrous, the lower 2 with apical tufts of white hairs. Ovary inferior, 2-celled; ovule placentation axile; stigma 2-lobed, wet or dry, appressed and nonreceptive as the style grows through the anther tube, pushing out the pollen, after which the stigmas spread apart and become receptive.

Fruit: Berries purple; globose; 5–8 mm wide. Seeds numerous.

Ploidy:

Habitat: Occurring in diverse mesic forest to wet forest; western and south-central Kaua'i.

Elevation Range: 700–1,200 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Minnesota Bot. Stud. 1: 909 (1896)

Other References

Wagner et al. 1990:465 (K [as C. spathulata subsp. spathulata]); Lammers 1998:33 (revert to C. spathulata, as C. spathulata subsp. longipetiolata now = C. kahiliensis); Lammers 2004:100 (KEY)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Cyanea spathulata (Hillebr.) A.Heller forest of kaholuamano Forest PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Kauai BISH 10/1/1911
2 Cyanea spathulata (Hillebr.) A.Heller kalalau pali PRESERVED_SPECIMEN forbes, c.n. collector number: 1026.k. Kauai BISH 7/3/1917
3 Cyanea spathulata (Hillebr.) A.Heller waimea district. kohua ridge, north facing slopes above mohihi stream scrub PRESERVED_SPECIMEN wood, k.r. query, m., kirkpatrick, m. collector number: 15080 Maui BISH 7/11/2012
4 Cyanea spathulata (Hillebr.) A.Heller kaholuamanoa behind waimea PRESERVED_SPECIMEN forbes, c.n. collector number: 387.k. Kauai BISH 9/1/1909
5 Cyanea spathulata (Hillebr.) A.Heller Kaholuamano PRESERVED_SPECIMEN Rock, J.F.C. 9008 Kauai BISH 10/1/1911
6 Cyanea spathulata (Hillebr.) A.Heller waimea district. kokee; end of camp 10 road, on trail to circle bog, but drop down north into tributary of waiakoali, along steam on bank Growing near shore with Leucaena leucocephala and other exotics. PRESERVED_SPECIMEN tangalin, n. wood, k.r. collector number: 2437 Kauai BISH 12/8/2010
7 Cyanea spathulata (Hillebr.) A.Heller Waimea PRESERVED_SPECIMEN Knudsen, V. s.n. Kauai BISH 2/12/1905
8 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Waiakoali Along stream in predominately native, mesic to wet forest. W/ Diparia sp., Metrosideros polymorpha, Broussaisia arguta, Cheirodendron spp., Labordia degeneri, Dicranopteris linearis, Diplopterygium pinnatum, Melicope clusiifolia, Hedychium gardnerianum,. Cibot PRESERVED_SPECIMEN Walsh, S. SKW537 Kauai BISH 9/1/2017
9 Cyanea spathulata (Hillebr.) A.Heller wainiha, upper ne fork Dry native shrub with mixed grass community. Extensive soil erosion PRESERVED_SPECIMEN wood, k.r. collector number: 13276 Maui BISH 9/12/2008
10 Cyanea spathulata (Hillebr.) A.Heller headwater of n fork waialua river, just e of mts waialeale and kawaikini Deep valley surrounded by steep waterfall-laced cliffs, low primary vegetation of small, stunted trees 1-2 m tall, numerous pteridophytes and Gunnera over saturated ground PRESERVED_SPECIMEN lorence, d.h. plews, j.h.r.; wood, k.r; doyle, m. collector number: 5367 Kauai BISH 7/23/1987
11 Cyanea spathulata (Hillebr.) A.Heller kaholuaumanu dry limestone woods PRESERVED_SPECIMEN rock, j.f.c. collector number: 9008 Kauai BISH 10/1/1911
12 Cyanea spathulata (Hillebr.) A.Heller Waimea PRESERVED_SPECIMEN Knudsen 11 Kauai BISH
13 Cyanea spathulata (Hillebr.) A.Heller Forest of Kaholuamano Forest PRESERVED_SPECIMEN Rock, J.F.C. s.n. Kauai BISH 10/1/1911
14 Cyanea spathulata (Hillebr.) A.Heller kahaluamano PRESERVED_SPECIMEN rock, j.f.c. Kauai BISH 3/1/1909
15 Cyanea spathulata (Hillebr.) A.Heller kaholuamanoa, above waimea PRESERVED_SPECIMEN heller, a.a. collector number: 2768 Kauai BISH 1895-09-02
16 Cyanea spathulata (Hillebr.) A.Heller Found with Syzygium sandwicensis, Kadua affinis, Pittosporum gayanum, P. glabrum, Ilex anomala, Melicope anisata, M. clusiifolia, Myrsine dentata, M. Punctata, Psychotria hexandra, P. mariniana, P. greenwel. PRESERVED_SPECIMEN Wood, K.R. 15221 Kauai BISH 9/14/2012
17 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Kohua Ridge, north facing slopes above Mohihi Stream Mestrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Cheirodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphoriba atrococca, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena au PRESERVED_SPECIMEN Wood, K.R. 15080 Kauai BISH 7/11/2012
18 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi Valley Mist drainage on N side of valley, below Honopu trail Diverse mesic forest. Metrosideros-Acacia koa overstory with Dodonaea viscosa, Claoxylon, Coprosma, Cyanea hirtella, Hedychium gardnerianum. PRESERVED_SPECIMEN Trauernicht, C. Tangalin, N. Collector Number: 641 Kauai BISH 12/4/2008
19 Cyanea spathulata (Hillebr.) A.Heller Kahaluamano PRESERVED_SPECIMEN Rock, J.F.C. s.n. Kauai BISH 3/1/1909
20 Cyanea spathulata (Hillebr.) A.Heller waimea district. kohua ridge, north facing slopes above mohihi stream Metrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Cheirodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena au PRESERVED_SPECIMEN wood, k.r. query, m., kirkpatrick, m. collector number: 15074 Kauai BISH 7/11/2012
21 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi Valley. Below Honopu trail Moist drainage on N side of valley. Diverse mesic forest. Metrosideros-Acacia koa overstory with Dodonaea viscosa, Claoxylon, Coprosma, Cyanea hirtella, Hedychium gardnerianum. PRESERVED_SPECIMEN Trauernicht, C. 642 Kauai BISH 12/4/2008
22 Cyanea spathulata (Hillebr.) A.Heller Waioli; hanging valley w/ main falls. Metrosideros- Cheirodendron Lowland Wet Forest. PRESERVED_SPECIMEN K. R. Wood 2144 Kauai PTBG 11/19/1992
23 Cyanea spathulata (Hillebr.) A.Heller Found with Syzygium sandwicensis, Kadua affinis, Pittosporum gayanum, P. glabrum, Ilex anomala, Melicope anisata, M. clusiifolia, Myrsine dentata, M. Punctata, Psychotria hexandra, P. mariniana, P. greenwel. PRESERVED_SPECIMEN Wood, K.R. Wood, M. Collector Number: 15221.00 Kauai BISH 9/14/2012
24 Cyanea spathulata (Hillebr.) A.Heller e rim of kalalaua valley Among staghorn PRESERVED_SPECIMEN degener, o. collector number: 21468 Kauai BISH 12/27/1951
25 Cyanea spathulata (Hillebr.) A.Heller Hawaii - Kauai, Waimea PRESERVED_SPECIMEN Faurie, U. J. (Coll. Delessert) G
26 Cyanea spathulata (Hillebr.) A.Heller lihue-koloa forest reserve. PRESERVED_SPECIMEN eric b. knox 2384b IND 7/1/1991
27 Cyanea spathulata (Hillebr.) A.Heller PRESERVED_SPECIMEN o. degener 7867 MPM 6/30/1926
28 Cyanea spathulata (Hillebr.) A.Heller lihue-koloa forest reserve. PRESERVED_SPECIMEN eric b. knox 2384a NY 7/1/1991
29 Cyanea spathulata (Hillebr.) A.Heller Makaleha Mtns., north-facing cliffs of Kekoiki Peak. Wet Windward cliffs. PRESERVED_SPECIMEN Natalia Tangalin 2372 Kauai PTBG 7/8/2010
30 Cyanea spathulata (Hillebr.) A.Heller KAUAI: Kokee State Park, along Hwy 550 near mile 18.3. Along western side of ridge jutting into Kalalau Valley. PRESERVED_SPECIMEN Tim Flynn 1226 Kauai PTBG 9/11/1985
31 Cyanea spathulata (Hillebr.) A.Heller Waiakoali, Na Pali Kona Forest Reserve. Metrosideros-Cheirodendron montane wet slopes and drainage. PRESERVED_SPECIMEN K. R. Wood 15221 Kauai PTBG 9/14/2012
32 Cyanea spathulata (Hillebr.) A.Heller Kokee, End of Camp 10 Road, along Mohihi Stream Metrosideros-Cheirodendron-Dicranopteris Wet Montane Forest PRESERVED_SPECIMEN Natalia Tangalin 3347 Kauai PTBG 9/19/2011
33 Cyanea spathulata (Hillebr.) A.Heller Mohihi, slopes and drainage north of Mohihi-Waialae trail. Metrosideros mixed montane wet forest with some steep slopes dominated by matting ferns of Dicranopteris, Diplopterygium, or Sticherus PRESERVED_SPECIMEN K. R. Wood 18228 Kauai PTBG 6/19/2019
34 Cyanea spathulata (Hillebr.) A.Heller Makaleha Mts.; North facing steep slopes below Kekoiki. Metrosideros- Dicranopteris Lowland Wet Forest. PRESERVED_SPECIMEN K. R. Wood 2849 Kauai PTBG 11/10/1993
35 Cyanea spathulata (Hillebr.) A.Heller Kohua Ridge, North facing slopes above Mohihi Stream. Metrosideros-Acacia montane mesic forest. PRESERVED_SPECIMEN K. R. Wood 15080 Kauai PTBG 7/11/2012
36 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi valley, north aspect on banks. In Acacia koa- Metrosideros mesic forest. PRESERVED_SPECIMEN K. R. Wood 5353 Kauai PTBG 7/5/1996
37 Cyanea spathulata (Hillebr.) A.Heller Na Pali-Kona Forest Reserve; along Nawaimaka Stream. Elev. 3180-3200 ft. Through disturbed forest of Acacia koa and Metrosideros polymorpha. PRESERVED_SPECIMEN Tim Flynn 5451 Kauai PTBG 9/9/1993
38 Cyanea spathulata (Hillebr.) A.Heller Wai'oli; back of valley. Metrosideros-Dicranopteris lowland wet forest. PRESERVED_SPECIMEN K. R. Wood 2042 Kauai PTBG 8/6/1992
39 Cyanea spathulata (Hillebr.) A.Heller Wai'oli; back of valley. Metrosideros-Dicranopteris lowland wet forest. PRESERVED_SPECIMEN K. R. Wood 2051 Kauai PTBG 8/6/1992
40 Cyanea spathulata (Hillebr.) A.Heller Kokee; end of Camp 10 Road, on trail to Circle Bog, but drop down North into tributary of Waiakoali, along steam on bank. Metrosideros-Dicranopteris-Cheirodendron Wet Forest PRESERVED_SPECIMEN Natalia Tangalin 2437 Kauai PTBG 12/8/2010
41 Cyanea spathulata (Hillebr.) A.Heller border of kawaihau and halalei dist., makaleha mts., n facing slopes below kekoiki PRESERVED_SPECIMEN k. r. wood 2849 Kauai US 11/10/1993
42 Cyanea spathulata (Hillebr.) A.Heller Na Pali-Kona Forest Reserve. Pihea Trail through Kawaikoi Valley. Near largest tributary from south side. Near downslope side of trail. PRESERVED_SPECIMEN John H. R. Plews s.n. Kauai PTBG 7/18/1987
43 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi Valley. Moist drainage on N side of valley, below Honopu trail. Diverse mesic forest. PRESERVED_SPECIMEN Clay Trauernicht 641 Kauai PTBG 12/4/2008
44 Cyanea spathulata (Hillebr.) A.Heller Honopu valley, near stream bottum, on steep slopes. In Metrosideros -Dicranopteris linearis wet forest. PRESERVED_SPECIMEN Steve Perlman 13736 Kauai PTBG 8/17/1993
45 Cyanea spathulata (Hillebr.) A.Heller Wainiha, upper northeastern fork. Closed Metrosideros forest, 8-12 m canopy. PRESERVED_SPECIMEN K. R. Wood 13276 Kauai PTBG 9/12/2008
46 Cyanea spathulata (Hillebr.) A.Heller KAUAI: Kawai`iki, off Kaluakaulu ridge, upper forest and drainage to the south of Koai’e and north of Waialae, Metrosideros polymorpha mixed mesic forest w/ Dubautia laevigata, Hedyotis terminalis, Cheirodendron trigynum, Dianella sandwicensis, Poa sandwicensis, Bidens cosmoides, Peperomia membranacea PRESERVED_SPECIMEN K. R. Wood 9003 Kauai PTBG 6/27/2001
47 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi Valley. Moist drainage on N side of valley, below Honopu trail. Diverse mesic forest. PRESERVED_SPECIMEN Clay Trauernicht 642 Kauai PTBG 12/4/2008
48 Cyanea spathulata (Hillebr.) A.Heller Wai'oli; back of valley. Metrosideros-Dicranopteris lowland wet forest. PRESERVED_SPECIMEN K. R. Wood 2044 Kauai PTBG 8/6/1992
49 Cyanea spathulata (Hillebr.) A.Heller Kalalau below and west of Puu o Kila. Metrosideros-Cheirodendron montane wet forest. PRESERVED_SPECIMEN K. R. Wood 8130 Kauai PTBG 1/11/2000
50 Cyanea spathulata (Hillebr.) A.Heller Wainiha, upper northeastern fork. Closed Metrosideros forest, 8-12 m canopy. PRESERVED_SPECIMEN K. R. Wood 13254 Kauai PTBG 9/4/2008
51 Cyanea spathulata (Hillebr.) A.Heller Koaie canyon drainages, above "Twin Falls", north of Kawaiiki ridge, south drainage of twin falls, N fa 22 deg. 05.471'N & 159 deg. 36.027'W In Acacia koa- Metrosideros mesic forest. PRESERVED_SPECIMEN Steve Perlman 17695 Kauai PTBG 9/25/2001
52 Cyanea spathulata (Hillebr.) A.Heller Honopu, NW of Kainamanu, Acacia koa - Metrosideros polymorpha - Dicranopteris linearis montane mesic PRESERVED_SPECIMEN K. R. Wood 15163 Kauai PTBG 8/8/2012
53 Cyanea spathulata (Hillebr.) A.Heller Nuololo; Kuia Natural Area Reserve. Metrosideros-Dicranopteris Montane Mesic Forest in drainage below/ PRESERVED_SPECIMEN K. R. Wood 2715 Kauai PTBG 8/17/1993
54 Cyanea spathulata (Hillebr.) A.Heller Kaunuohua ridge, near Puu o Kila lookout, in small stream. Metrosideros- Dicranopteris linearis wet forest. PRESERVED_SPECIMEN Steve Perlman 23722 Kauai PTBG 9/27/2013
55 Cyanea spathulata (Hillebr.) A.Heller Waioli; hanging valley w/ main falls. Metrosideros- Cheirodendron Lowland Wet Forest. PRESERVED_SPECIMEN K. R. Wood 2132 Kauai PTBG 11/19/1992
56 Cyanea spathulata (Hillebr.) A.Heller Wainiha; drainage south of Mahina- kehau. Metrosideros-Dicranopteris Lowland Wet Forest. PRESERVED_SPECIMEN K. R. Wood 2340 Kauai PTBG 1/29/1993
57 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi valley. PRESERVED_SPECIMEN K. R. Wood 5864 Kauai PTBG 12/7/1996
58 Cyanea spathulata (Hillebr.) A.Heller Waiakoali Along stream in predominately native, mesic to wet forest. PRESERVED_SPECIMEN Seana Walsh SKW537 Kauai PTBG 9/1/2017
59 Cyanea spathulata (Hillebr.) A.Heller Kohua Ridge, North facing slopes above Mohihi Stream. Metrosideros-Acacia montane mesic forest. PRESERVED_SPECIMEN K. R. Wood 15078 Kauai PTBG 7/11/2012
60 Cyanea spathulata (Hillebr.) A.Heller Wailua, Blue Hole, N. fork of the Wailua River, back wall, forested ridge at of back bowl and base of cliffs. Metrosideros Lowland Wet Forest patches and Shrubland. PRESERVED_SPECIMEN Natalia Tangalin 3001 Kauai PTBG 2/14/2012
61 Cyanea spathulata (Hillebr.) A.Heller Nuololo, northern branch. Metrosideros-Acacia montane mesic forest PRESERVED_SPECIMEN K. R. Wood 18344 Kauai PTBG 12/3/2019
62 Cyanea spathulata (Hillebr.) A.Heller Kohua Ridge, North facing slopes above Mohihi Stream. Metrosideros-Acacia montane mesic forest. PRESERVED_SPECIMEN K. R. Wood 15074 Kauai PTBG 7/11/2012
63 Cyanea spathulata (Hillebr.) A.Heller Kuia NAR, Kuia Valley, accessed from Nuololo Trail on N facing slope. Metrosideros-Acacia Mesic Forest PRESERVED_SPECIMEN Natalia Tangalin 2072 Kauai PTBG 7/1/2009
64 Cyanea spathulata (Hillebr.) A.Heller Kokee, Waiakoali Stream, ca. 50 yds past second footbridge. PRESERVED_SPECIMEN Tim Flynn 311 Kauai PTBG 9/2/1982
65 Cyanea spathulata (Hillebr.) A.Heller Kokee, End of Camp 10 Road, along Mohihi Stream Metrosideros-Cheirodendron-Dicranopteris Wet Montane Forest PRESERVED_SPECIMEN Natalia Tangalin 3351 Kauai PTBG 9/19/2011
66 Cyanea spathulata (Hillebr.) A.Heller Kokee State Park. Lower Pihea Trail between Alakai Swamp Trail and Kawaikoi Stream. PRESERVED_SPECIMEN John H. R. Plews s.n. Kauai PTBG 1/1/1985
67 Cyanea spathulata (Hillebr.) A.Heller on kaholuamanoa, above waimea PRESERVED_SPECIMEN a. a. heller 2768 Kauai US 1895-09-02
68 Cyanea spathulata (Hillebr.) A.Heller kaholuamano, above waimea PRESERVED_SPECIMEN a. a. heller 2768 Kauai US 1895-08-30
69 Cyanea spathulata (Hillebr.) A.Heller koaie canyon drainages, above "twin falls", north of kawaiiki ridge, south drainage of twin falls, n facing PRESERVED_SPECIMEN s. p. perlman & k. wood 17695 Kauai US 9/25/2001
70 Cyanea spathulata (Hillebr.) A.Heller kaholuamano, above waimea PRESERVED_SPECIMEN a. a. heller 2768 Kauai US 1895-09-02
71 Cyanea spathulata (Hillebr.) A.Heller along kokee stream PRESERVED_SPECIMEN o. degener 7867 Kauai US 6/30/1926
72 Cyanea spathulata (Hillebr.) A.Heller Honopu; upper north fork headwaters; follow dividing ridge of Kalalau & Honopu; 3800 ft contour south to gulch; follow gulch WSW. Metrosideros-Acacia-Dicranopteris Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4964 Kauai PTBG 2/21/1996
73 Cyanea spathulata (Hillebr.) A.Heller waimea dist, na pali-kona for res, along nawaimaka stream PRESERVED_SPECIMEN t. w. flynn & g. kawakai 5451 Kauai US 9/9/1993
74 Cyanea spathulata (Hillebr.) A.Heller waimea district, kokee, end of camp 10 road, on trail to circle bog, but drop down north into tributary of waiakoali, along stream on bank. Wet forest. PRESERVED_SPECIMEN n. tangalin & k. r. wood 2437 Kauai US 12/8/2010
75 Cyanea spathulata (Hillebr.) A.Heller waimea dist., nawaimaka stream Montane mesic forest. PRESERVED_SPECIMEN k. r. wood, t. w. flynn & s. p. perlman 2444 Kauai US 3/19/1993
76 Cyanea spathulata (Hillebr.) A.Heller kaholuamanu, behind waimea PRESERVED_SPECIMEN c. n. forbes 849.k Kauai US
77 Cyanea spathulata (Hillebr.) A.Heller kaholuamano PRESERVED_SPECIMEN a. s. hitchcock 15369 Kauai US 10/20/1916
78 Cyanea spathulata (Hillebr.) A.Heller Waimea District, Kokee, end of Camp 10 Road, on trail to Circle Bog, but drop down North into tributary of Waiakoali, along stream on bank. Wet forest. PRESERVED_SPECIMEN N. Tangalin & K. R. Wood 2437 Kauai US 12/8/2010
79 Cyanea spathulata (Hillebr.) A.Heller Koaie canyon drainages, above "Twin Falls", north of Kawaiiki ridge, south drainage of twin falls, N facing PRESERVED_SPECIMEN S. P. Perlman & K. Wood 17695 Kauai US 9/25/2001
80 Cyanea spathulata (Hillebr.) A.Heller waimea district, waiakoaki, na pali kona forest reserve. Montane wet slopes. PRESERVED_SPECIMEN k. r. wood & m. kirkpatrick 15221 Kauai US 9/14/2012
81 Cyanea spathulata (Hillebr.) A.Heller lihue-koloa forest reserve. PRESERVED_SPECIMEN eric b. knox 2384c VT 7/1/1991
82 Cyanea spathulata (Hillebr.) A.Heller kaholuamanoa PRESERVED_SPECIMEN forbes, c.n. collector number: 343.k. Kauai BISH 9/1/1909
83 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi Valley. Below Honopu trail Moist drainage on N side of valley. Diverse mesic forest. Metrosideros-Acacia koa overstory with Dodonaea viscosa, Claoxylon, Coprosma, Cyanea hirtella, Hedychium gardnerianum. PRESERVED_SPECIMEN Trauernicht, C. Tangalin, N. Collector Number: 642 Kauai BISH 12/4/2008
84 Cyanea spathulata (Hillebr.) A.Heller Waimea canyon drainage, Nawaimaka valley, along bank of stream, at 22 deg. 04.269'N & 159 deg. 36.045'W on south bank of stream, scattered shrubs along streams. In Acacia koa- Metrosideros mesic forest. PRESERVED_SPECIMEN Steve Perlman 17628 Kauai PTBG 8/7/2001
85 Cyanea spathulata (Hillebr.) A.Heller Kaholuamanoa PRESERVED_SPECIMEN Forbes, C.N. s.n. Kauai BISH 9/1/1909
86 Cyanea spathulata (Hillebr.) A.Heller Waimea District, Waiakoaki, Na Pali Kona Forest Reserve. Montane wet slopes. PRESERVED_SPECIMEN K. R. Wood & M. Kirkpatrick 15221 Kauai US 9/14/2012
87 Cyanea spathulata (Hillebr.) A.Heller Makaleha Mtns., N facing claiffs of Kekoiki Peak. Wet Windward cliffs. W/ Metrosideros, Dicranopteris, Diplopterygium, Selaginella arbuscula, Broussaisia arguta, Peperomia, Anoectochilus sandvicensis, Pipturus ruber, Perrottetia sandwicensis, Cyrtandra paliku, Cyanea fissa. PRESERVED_SPECIMEN Tangalin, N. 2372 Kauai BISH 7/8/2010
88 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Kokee, End of Camp 10 Road, along Mohihi Stream Metrosideros-Cheirodendron-Dicranopteris Wet Montane Forst Psychotria, Kadu, Bobea, Myrsine, Nestegis, Cyanea spathulata, C. fiss (gayana), Kadua ffinis, Saddleria, Cibotium, Syzgium, Dubautia latifolia PRESERVED_SPECIMEN Tangalin, N. 3347 Kauai BISH 9/19/2011
89 Cyanea spathulata (Hillebr.) A.Heller Kalalau Pali PRESERVED_SPECIMEN Forbes, C.N. 1026.K. Kauai BISH 7/3/1917
90 Cyanea spathulata (Hillebr.) A.Heller opaewelu stream PRESERVED_SPECIMEN rock, j.f.c. collector number: 5664 Oahu US 9/7/1909
91 Cyanea spathulata (Hillebr.) A.Heller lihue-koloa forest reserve. PRESERVED_SPECIMEN eric b. knox 2384 Hawaii US 7/1/1991
92 Cyanea spathulata (Hillebr.) A.Heller Wainiha, upper NE fork Closed Metrowideros forest, 8-12 m canopy. Understory dominated by Antidesma w/mixed wet associates, Syzygium, Broussaisia, Perrottetia, Cyrtandra spp, Psychotria spp, Dubautia spp, Labordia spp, Coprosma waimeae, Cheirodendron spp, Tetraplasandra PRESERVED_SPECIMEN Wood, K.R. 13276 Kauai BISH 9/12/2008
93 Cyanea spathulata (Hillebr.) A.Heller waimea district. pihea trail about 1/2 mile from the junction of the crossover to the kawaikoi trail PRESERVED_SPECIMEN gustafson, r. herbst, d.r. collector number: 3046 Oahu BISH 7/22/1984
94 Cyanea spathulata (Hillebr.) A.Heller awaawapuhi valley. below honopu trail Moist drainage on N side of valley. Diverse mesic forest. Metrosideros-Acacia koa overstory with Dodonaea viscosa, Claoxylon, Coprosma, Cyanea hirtella, Hedychium gardnerianum. PRESERVED_SPECIMEN trauernicht, c. tangalin, n. collector number: 642 Kauai BISH 12/4/2008
95 Cyanea spathulata (Hillebr.) A.Heller PRESERVED_SPECIMEN wood, k.r. wood, m. collector number: 15221.00 Oahu BISH 9/14/2012
96 Cyanea spathulata (Hillebr.) A.Heller Waimea Drainage Basin, W side PRESERVED_SPECIMEN Forbes, C.N. 996.K. Kauai BISH 7/3/1917
97 Cyanea spathulata (Hillebr.) A.Heller Waimea PRESERVED_SPECIMEN Knudsen, V. s.n. Kauai BISH
98 Cyanea spathulata (Hillebr.) A.Heller E rim of Kalalaua Valley Among staghorn PRESERVED_SPECIMEN Degener, O. 21468 Kauai BISH 12/27/1951
99 Cyanea spathulata (Hillebr.) A.Heller Opaewelu Stream PRESERVED_SPECIMEN Rock, J.F.C. 5664 Kauai BISH 9/7/1909
100 Cyanea spathulata (Hillebr.) A.Heller awaawapuhi valley Mist drainage on N side of valley, below Honopu trail Diverse mesic forest. Metrosideros-Acacia koa overstory with Dodonaea viscosa, Claoxylon, Coprosma, Cyanea hirtella, Hedychium gardnerianum. PRESERVED_SPECIMEN trauernicht, c. tangalin, n. collector number: 641 Kauai BISH 12/4/2008
101 Cyanea spathulata (Hillebr.) A.Heller trail to kauluwehi swamp PRESERVED_SPECIMEN rock, j.f.c. Kauai BISH 10/1/1916
102 Cyanea spathulata (Hillebr.) A.Heller Along Nawaimaka Stream, Na Pali-Kona Forest Reserve Disturbed forest of Acacia koa and Metrosideros polymorpha PRESERVED_SPECIMEN Flynn, T. 5451 Kauai BISH 9/9/1993
103 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Kohua Ridge, north facing slopes above Mohihi Stream Metrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Cheirodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena au PRESERVED_SPECIMEN Wood, K.R. 15074 Kauai BISH 7/11/2012
104 Cyanea spathulata (Hillebr.) A.Heller waimea PRESERVED_SPECIMEN knudsen, v. Kauai BISH 1870-08-08
105 Cyanea spathulata (Hillebr.) A.Heller waimea dist., na pali-kona forest reserve, pihea trail through kawaikoi valley Common on roadside. PRESERVED_SPECIMEN j. plews s.n. Hawaii BISH 7/18/1987
106 Cyanea spathulata (Hillebr.) A.Heller waimea PRESERVED_SPECIMEN knudsen collector number: 11 Kauai BISH
107 Cyanea spathulata (Hillebr.) A.Heller Awaawapuhi Valley Mist drainage on N side of valley, below Honopu trail Diverse mesic forest. Metrosideros-Acacia koa overstory with Dodonaea viscosa, Claoxylon, Coprosma, Cyanea hirtella, Hedychium gardnerianum. PRESERVED_SPECIMEN Trauernicht, C. 641 Kauai BISH 12/4/2008
108 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Kokee; end of Camp 10 Road, on trail to Circle Bog, but drop down North into tributary of Waiakoali, along steam on bank Metrosideros-Dicranopteris-Cheirodendron Wet Forest Asplenium schizPsychotria, Coprosma, Saddleria, Broussaisia arguta, Elaphoglossum crassicaule, Vaccinium, Cyrtandra, Pittosporum, Platydesma, Peperomia kokeana, Seaginella, Deparia, Dryopteirs, Callis PRESERVED_SPECIMEN Tangalin, N. 2437 Kauai BISH 12/8/2010
109 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Pihea Trail about 1/2 mile from the junction of the crossover to the Kawaikoi trail Moderately wet mesic forest assoc. with Pelea, Smilax, Cheirodendron, Psychotria, Pittosporum, Cyrtandra, etc. PRESERVED_SPECIMEN Gustafson, R. 3046 Kauai BISH 7/22/1984
110 Cyanea spathulata (Hillebr.) A.Heller Kaholuamanoa behind Waimea PRESERVED_SPECIMEN Forbes, C.N. 387.K. Kauai BISH 9/1/1909
111 Cyanea spathulata (Hillebr.) A.Heller makaleha mtns., n facing claiffs of kekoiki peak. Wet Windward cliffs. W/ Metrosideros, Dicranopteris, Diplopterygium, Selaginella arbuscula, Broussaisia arguta, Peperomia, Anoectochilus sandvicensis, Pipturus ruber, Perrottetia sandwicensis, Cyrtandra paliku, Cyanea fissa. PRESERVED_SPECIMEN tangalin, n. wood, k.r.; kishida, w. collector number: 2372 Kauai BISH 7/8/2010
112 Cyanea spathulata (Hillebr.) A.Heller Waiakealoha PRESERVED_SPECIMEN Rock, J.F.C. 2456 Kauai BISH 3/1/1909
113 Cyanea spathulata (Hillebr.) A.Heller Kaholuaumanu PRESERVED_SPECIMEN Rock, J.F.C. 9008 Kauai BISH 10/1/1911
114 Cyanea spathulata (Hillebr.) A.Heller kalalau pali PRESERVED_SPECIMEN forbes, c.n. collector number: 1026(a).k. Kauai BISH 7/3/1917
115 Cyanea spathulata (Hillebr.) A.Heller along nawaimaka stream, na pali-kona forest reserve Disturbed forest of Acacia koa and Metrosideros polymorpha PRESERVED_SPECIMEN flynn, t. kawakami, g. collector number: 5451 Kauai BISH 9/9/1993
116 Cyanea spathulata (Hillebr.) A.Heller Kaholuamanoa PRESERVED_SPECIMEN Forbes, C.N. 343.K. Kauai BISH 9/1/1909
117 Cyanea spathulata (Hillebr.) A.Heller waimea district. kohua ridge, north facing slopes above mohihi stream Metrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Cheirodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Dodonaea viscosa, Myrisne lanaiensis, Nestegis sandwicensis, Dracaena au PRESERVED_SPECIMEN wood, k.r. query, m., kirkpatrick, m. collector number: 15078 Kauai BISH 7/11/2012
118 Cyanea spathulata (Hillebr.) A.Heller Trail to Kauluwehi swamp PRESERVED_SPECIMEN Rock, J.F.C. s.n. Kauai BISH 10/1/1916
119 Cyanea spathulata (Hillebr.) A.Heller waiakealoha with Vaccinium, Deschampsia PRESERVED_SPECIMEN rock, j.f.c. collector number: 2456 Maui BISH 3/1/1909
120 Cyanea spathulata (Hillebr.) A.Heller kaholuamanoa PRESERVED_SPECIMEN forbes, c.n. Kauai BISH 9/1/1909
121 Cyanea spathulata (Hillebr.) A.Heller Headwater of N fork Waialua River, just E of Mts Waialeale and Kawaikini Deep valley surrounded by steep waterfall-laced cliffs, low primary vegetation of small, stunted trees 1-2 m tall, numerous pteridophytes and Gunnera over saturated ground PRESERVED_SPECIMEN Lorence, D.H. 5367 Kauai BISH 7/23/1987
122 Cyanea spathulata (Hillebr.) A.Heller Kalalau Pali PRESERVED_SPECIMEN Forbes, C.N. 1026(a).K. Kauai BISH 7/3/1917
123 Cyanea spathulata (Hillebr.) A.Heller Waimea District. Kohua Ridge, north facing slopes above Mohihi Stream Metrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Cheirodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Dodonaea viscosa, Myrisne lanaiensis, Nestegis sandwicensis, Dracaena au PRESERVED_SPECIMEN Wood, K.R. 15078 Kauai BISH 7/11/2012
124 Cyanea spathulata (Hillebr.) A.Heller Kaholuamanoa, above Waimea PRESERVED_SPECIMEN Heller, A.A. 2768 Kauai BISH 1895-00-00
125 Cyanea spathulata (Hillebr.) A.Heller kaholuamano PRESERVED_SPECIMEN rock, j.f.c. collector number: 9008 Kauai BISH 10/1/1911
126 Cyanea spathulata (Hillebr.) A.Heller waimea drainage basin, w side PRESERVED_SPECIMEN forbes, c.n. collector number: 996.k. Kauai BISH 7/3/1917
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