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Sphaeropteris cooperi

(Hook. ex F.Muell.) R.M.Tryon (1970)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Cyatheales Family: Cyatheaceae Genus: Sphaeropteris

Australian tree fern, scaly tree fern

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Potentially naturalizing
Lana'i Naturalized
Kaho'olawe Potentially naturalizing
Maui Naturalized
Hawai'i Unknown

Dispersal Agents

Water/Ocean

Bibliography

Name Published In: Contr. Gray Herb. 200: 24 (1970)

Other References

Palmer 2003:243 (K, O, M, H); Staples & Herbst 2005:32 (DESCR [as Cyathea cooperi]); Oppenheimer 2008:26 (L); Palmer & Thomas in press (Mo); Paul Higashino pers. comm. 2025 (Ka? [observed but no spm. from under rain catchment])

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Sphaeropteris cooperi South drainage of upper Honopu; enter via Honopu trail; drop into gulch to north of trail 4180 ft.; Elev. 3700- 3900 feet. Metrosideros-Dicranopteris Montane Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4990 Kauai PTBG 2/25/1996
2 Sphaeropteris cooperi w maui, hoholua, maunalei arboretum Closed-canopy, alien dominated forest. Assoc. vegetation: Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, Psidium. PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 187 Maui BISH 3/7/1997
3 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN Oppenheimer, H.L. Hansen, G. Collector Number: H60403 Maui BISH 6/10/2004
4 Sphaeropteris cooperi Honolulu: Kahala; 3653 Kahala Avenue; residence of C. F. Johnston In full sun. PRESERVED_SPECIMEN Lau, J. 2136 Oahu BISH 3/17/1986
5 Sphaeropteris cooperi Honolulu; Bishop Museum In shade house at back of campus. Grown in 18' wooden box. PRESERVED_SPECIMEN Evans, M.A. 1 Oahu BISH 6/20/1990
6 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN Oppenheimer, H.L. H90631 Maui BISH 9/28/2006
7 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN Oppenheimer, H.L. H60403 Maui BISH 6/10/2004
8 Sphaeropteris cooperi east maui, hana district, kahikinui, wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h90631 Maui BISH 9/28/2006
9 Sphaeropteris cooperi east maui, hana district, hana district,kipahulu valley PRESERVED_SPECIMEN gagné, b.h. collector number: 3056 MPM 4/24/1991
10 Sphaeropteris cooperi East Maui, Hana District, Hana District,Kipahulu Valley Growing as an aggresive weed. PRESERVED_SPECIMEN Gagne, B.H. 3056 Maui BISH 4/24/1991
11 Sphaeropteris cooperi Honolulu; Makiki; grounds of Contemporary Museum, 2411 Makiki Heights Drive In garden. PRESERVED_SPECIMEN Staples, G. 848 Oahu BISH 7/13/1992
12 Sphaeropteris cooperi East Maui, Hana District, Hana District,Kipahulu Valley Growing as an aggressive weed. PRESERVED_SPECIMEN Gagne, B.H. 3056 Maui BISH 4/24/1991
13 Sphaeropteris cooperi Koloa Dist.; Lihue-Koloa Forest Reserve, Wahiawa drainage; along main fork & upper tributaries of Wahiawa stream to crest of ridge just SW of Kapalaoa Diverse wet forest dominated by Metrosideros spp., Antidesma, Psychotria, Perrottetia, Hedyotis tryblium, Cyrtandra spp., Labordia spp., Broussaisia, Dubautia, Melicope spp., Freycinetia, Syzygium, Tetraplasandra & w/mixed open patches of Dicranopteris/Diplopterygium especially on the sides & crest of ridge; invaded by Psidium cattleianum, Rubus rosifolius. PRESERVED_SPECIMEN Flynn, T. 4593 Kauai BISH 4/12/1991
14 Sphaeropteris cooperi honolulu; bishop museum In shade house at back of campus. Grown in 18' wooden box. PRESERVED_SPECIMEN evans, m. collector number: 01 Oahu BISH 6/20/1990
15 Sphaeropteris cooperi Waianae Range: along the summit of the Waianae Mountains from Mauna Kapu to Pali Kea and beyond Cloud forest on the windward side. PRESERVED_SPECIMEN Wilson, K.A. 2519 Oahu BISH 2/6/1996
16 Sphaeropteris cooperi W Maui, Hoholua, Maunalei Arboretum Closed-canopy, alien dominated forest. Assoc. vegetation: Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, Psidium. PRESERVED_SPECIMEN Meidell, J.S. 187 Maui BISH 3/7/1997
17 Sphaeropteris cooperi oahu forest national wildlife refuge, kipapa stream, upslope ca 300 yd from stream. Dicranopteris-dominated slope. PRESERVED_SPECIMEN hanley, j. collector number: s.n. Oahu BISH 9/8/2011
18 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley In shady wet valley in Acacia--Metrosideros forest which has been altered by introduced tree species, such as Aleurites moluccana and Ficus sp. With such shrubby species as Cyrtandra and Cordyline terminalis. PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Oahu BISH 6/10/2004
19 Sphaeropteris cooperi Waiopa Gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN Oppenheimer, H.L. Perlman, S. Collector Number: H90705 Lanai BISH 9/5/2007
20 Sphaeropteris cooperi honolulu; makiki; grounds of contemporary museum, 2411 makiki heights drive In garden. PRESERVED_SPECIMEN staples, g.w. medbury, j.s. collector number: 848 Oahu BISH 7/13/1992
21 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Maui BISH 6/10/2004
22 Sphaeropteris cooperi waianae range: along the summit of the waianae mountains from mauna kapu to pali kea and beyond PRESERVED_SPECIMEN wilson, k.a. palmer, d.d.; obata, j.; gustafason, b. collector number: 2519 Kauai BISH 2/6/1996
23 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H90631 Maui BISH 9/28/2006
24 Sphaeropteris cooperi east maui, hana district, hana district,kipahulu valley Growing as an aggresive weed. PRESERVED_SPECIMEN gagné, b.h. collector number: 3056 Maui BISH 4/24/1991
25 Sphaeropteris cooperi hanalilolilo sink hole With Acacia and Gouldia. PRESERVED_SPECIMEN hughes, g.d. collector number: s.n. Oahu BISH 2/10/1992
26 Sphaeropteris cooperi Hanalilolilo Sink Hole Mesic forest. PRESERVED_SPECIMEN Hughes, G.D. s.n. Molokai BISH 2/10/1992
27 Sphaeropteris cooperi w maui, hoholua, maunalei arboretum PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 187 Maui BISH 3/7/1997
28 Sphaeropteris cooperi koloa dist.; lihue-koloa forest reserve, wahiawa drainage; along main fork & upper tributaries of wahiawa stream to crest of ridge just sw of kapalaoa PRESERVED_SPECIMEN flynn, t. lorence, d.h.; perlman, s.; wood, k. collector number: 4593 Maui BISH 4/12/1991
29 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley Under bridge in ephemeral creek. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0330 Maui HAPI 11/22/2007
30 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley HUMAN_OBSERVATION Big Island Invasive Species Committee (BIISC) Maui HAPI 11/13/2008
31 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters HUMAN_OBSERVATION Big Island Invasive Species Committee (BIISC) Maui HAPI 11/13/2008
32 Sphaeropteris cooperi Oahu Forest National Wildlife Refuge, Kipapa Stream, upslope ca 300 yd from stream. Dicranopteris-dominated slope. PRESERVED_SPECIMEN Hanley, J. s.n. Oahu BISH 9/8/2011
33 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Lanai BISH 6/10/2004
34 Sphaeropteris cooperi kalihi valley, above waterfall Two plants along trail. Non-native dominated lowland mesic forest. PRESERVED_SPECIMEN us army kawelo, k. collector number: usarmy 185 Oahu BISH 5/9/2010
35 Sphaeropteris cooperi waiopa gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN oppenheimer, h.l. perlman, s. collector number: h90705 Lanai BISH 9/5/2007
36 Sphaeropteris cooperi waianae range: along the summit of the waianae mountains from mauna kapu to pali kea and beyond Locally common in sparse bushy vegetation, in seriously eroded slope of fine clay deeply weathered red volcanic soil PRESERVED_SPECIMEN wilson, k.a. palmer, d.d.; obata, j.; gustafason, b. collector number: 2519 Kauai BISH 2/6/1996
37 Sphaeropteris cooperi honolulu: kahala; 3653 kahala avenue; residence of c. f. johnston In full sun. PRESERVED_SPECIMEN lau, j. cory, c. collector number: 2136 Oahu BISH 3/17/1986
38 Sphaeropteris cooperi Waiopa Gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN Oppenheimer, H.L. H90705 Lanai BISH 9/5/2007
39 Sphaeropteris cooperi honolulu, bishop museum, shade house, back side of campus PRESERVED_SPECIMEN evans, m. collector number: 1 Oahu BISH 6/20/1990
40 Sphaeropteris cooperi Maui, road to Haleakalae Under bridge in ephemeral creek. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0330 Consortium of Pacific Northwest Herbaria 11/22/2007
41 Sphaeropteris cooperi Kalihi Valley, above waterfall Two plants along trail. Non-native dominated lowland mesic forest. PRESERVED_SPECIMEN Kawelo, K. USARMY 185 Oahu BISH 5/9/2010
42 Sphaeropteris cooperi Lawai Valley. The National Tropical Botanical Garden. Bamboo Bridge at W end of Hau Tunnel. Plants destroyed because species has become naturalized throughout island. Cultivated plants. PRESERVED_SPECIMEN Tim Flynn 5787 Kauai PTBG 4/21/1995
43 Sphaeropteris cooperi Kokee, .3 mile down Kumuwela Ridge Road. West of road off of ridge top in small, weedy gully bottom. PRESERVED_SPECIMEN Tim Flynn 1587 Kauai PTBG 3/4/1986
44 Sphaeropteris cooperi Hanalei Valley. Elev. 2220-2340 ft. Wet forest. PRESERVED_SPECIMEN Tim Flynn 5316 Kauai PTBG 3/22/1993
45 Sphaeropteris cooperi W. Maui, Honokahua, South of Valley. PRESERVED_SPECIMEN Hank Oppenheimer H60403 Maui PTBG 6/10/2004
46 Sphaeropteris cooperi Kokee State Park, Mohihi Road near emergency shower. Along the edge of roadside. PRESERVED_SPECIMEN Tim Flynn 3794 Kauai PTBG 3/1/1990
47 Sphaeropteris cooperi Lihue-Koloa Forest Reserve, Wahiawa drainage; along main fork and upper tributaries of Wahiawa stream to crest of ridge just SW of Kapalaoa. Elev. 2200-3200 ft. Diverse wet forest. PRESERVED_SPECIMEN Tim Flynn 4593 Kauai PTBG 4/13/1991
48 Sphaeropteris cooperi Limahuli Upper Preserve. Metrosideros wet forest. PRESERVED_SPECIMEN Emory Griffin-Noyes EGN 510 Kauai PTBG 2/14/2012
49 Sphaeropteris cooperi hosmer grove area, haleakala national park. outside of area planted by national park research team with trees from continental n. america. Wet forest PRESERVED_SPECIMEN leila m. shultz; john s. shultz 6395 Kauai US 12/23/1982
50 Sphaeropteris cooperi W. Maui, Maunalei Arboretum, Honolua. ASL. Closed canopy, alien dominated forest of Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, and Psidium guajava. PRESERVED_SPECIMEN J. Scott Meidell & Hank Oppenheimer 188 Maui PTBG
51 Sphaeropteris cooperi Halelea Forest Reserve. Along the powerline trail NW of the summit camp area. Stunted forest dominated by Metrosideros, Antidesma, and Dicranopteris. PRESERVED_SPECIMEN Tim Flynn 3546 Kauai PTBG 10/3/1989
52 Sphaeropteris cooperi E. Maui, east of Honomanu. In Metrosideros- Dicranopteris linearis wet forest. PRESERVED_SPECIMEN K. R. Wood 8772 Maui PTBG 10/17/2000
53 Sphaeropteris cooperi Limahuli Upper Preserve Metrosideros wet forest PRESERVED_SPECIMEN Emory Griffin-Noyes EGN 512 Kauai PTBG 2/23/2012
54 Sphaeropteris cooperi Near boundary with Waimea District. Upper part of Limahuli Valley from ridge separating Limahuli & Hanakapei Valleys (710 m alt.) down to Limahuli Stream (520 m alt.) above big waterfall. Nearly undisturbed rain forest with Metrosideros and Syzygium dominant. PRESERVED_SPECIMEN David H. Lorence 5721 Kauai PTBG 12/10/1987
55 Sphaeropteris cooperi Makaha Valley; upper valley via Milolii Rd.; at 3500 ft. contour north face of upper drainage; west of road. Elev. 3400-3600 ft. Acacia-Metrosideros Montane Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4983 Kauai PTBG 2/22/1996
56 Sphaeropteris cooperi Lawai Valley. The National Tropical Botanical Garden. Bamboo Bridge at W end of Hau Tunnel. Cultivated plants. PRESERVED_SPECIMEN Tim Flynn 5787 Kauai PTBG 4/21/1995
57 Sphaeropteris cooperi Waiopa Gulch. In gulch bottom. In lowland wet to mesic forest. PRESERVED_SPECIMEN Hank Oppenheimer H90705 Lanai PTBG 9/5/2007
58 Sphaeropteris cooperi koke`e, vicinity of wichman cabin on mohihi rd. PRESERVED_SPECIMEN Kauai US 1/17/2005
59 Sphaeropteris cooperi South drainage of upper Honopu; enter via Honopu trail; drop into gulch to north of trail 4180 ft.; Elev. 3700- 3900 feet. Metrosideros-Dicranopteris Montane Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4990 Kauai PTBG 2/25/1996
60 Sphaeropteris cooperi w maui, hoholua, maunalei arboretum Closed-canopy, alien dominated forest. Assoc. vegetation: Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, Psidium. PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 187 Maui BISH 3/7/1997
61 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN Oppenheimer, H.L. Hansen, G. Collector Number: H60403 Maui BISH 6/10/2004
62 Sphaeropteris cooperi Honolulu: Kahala; 3653 Kahala Avenue; residence of C. F. Johnston In full sun. PRESERVED_SPECIMEN Lau, J. 2136 Oahu BISH 3/17/1986
63 Sphaeropteris cooperi Honolulu; Bishop Museum In shade house at back of campus. Grown in 18' wooden box. PRESERVED_SPECIMEN Evans, M.A. 1 Oahu BISH 6/20/1990
64 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN Oppenheimer, H.L. H90631 Maui BISH 9/28/2006
65 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN Oppenheimer, H.L. H60403 Maui BISH 6/10/2004
66 Sphaeropteris cooperi east maui, hana district, kahikinui, wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h90631 Maui BISH 9/28/2006
67 Sphaeropteris cooperi east maui, hana district, hana district,kipahulu valley PRESERVED_SPECIMEN gagné, b.h. collector number: 3056 MPM 4/24/1991
68 Sphaeropteris cooperi East Maui, Hana District, Hana District,Kipahulu Valley Growing as an aggresive weed. PRESERVED_SPECIMEN Gagne, B.H. 3056 Maui BISH 4/24/1991
69 Sphaeropteris cooperi Honolulu; Makiki; grounds of Contemporary Museum, 2411 Makiki Heights Drive In garden. PRESERVED_SPECIMEN Staples, G. 848 Oahu BISH 7/13/1992
70 Sphaeropteris cooperi East Maui, Hana District, Hana District,Kipahulu Valley Growing as an aggressive weed. PRESERVED_SPECIMEN Gagne, B.H. 3056 Maui BISH 4/24/1991
71 Sphaeropteris cooperi Koloa Dist.; Lihue-Koloa Forest Reserve, Wahiawa drainage; along main fork & upper tributaries of Wahiawa stream to crest of ridge just SW of Kapalaoa Diverse wet forest dominated by Metrosideros spp., Antidesma, Psychotria, Perrottetia, Hedyotis tryblium, Cyrtandra spp., Labordia spp., Broussaisia, Dubautia, Melicope spp., Freycinetia, Syzygium, Tetraplasandra & w/mixed open patches of Dicranopteris/Diplopterygium especially on the sides & crest of ridge; invaded by Psidium cattleianum, Rubus rosifolius. PRESERVED_SPECIMEN Flynn, T. 4593 Kauai BISH 4/12/1991
72 Sphaeropteris cooperi honolulu; bishop museum In shade house at back of campus. Grown in 18' wooden box. PRESERVED_SPECIMEN evans, m. collector number: 01 Oahu BISH 6/20/1990
73 Sphaeropteris cooperi Waianae Range: along the summit of the Waianae Mountains from Mauna Kapu to Pali Kea and beyond Cloud forest on the windward side. PRESERVED_SPECIMEN Wilson, K.A. 2519 Oahu BISH 2/6/1996
74 Sphaeropteris cooperi W Maui, Hoholua, Maunalei Arboretum Closed-canopy, alien dominated forest. Assoc. vegetation: Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, Psidium. PRESERVED_SPECIMEN Meidell, J.S. 187 Maui BISH 3/7/1997
75 Sphaeropteris cooperi oahu forest national wildlife refuge, kipapa stream, upslope ca 300 yd from stream. Dicranopteris-dominated slope. PRESERVED_SPECIMEN hanley, j. collector number: s.n. Oahu BISH 9/8/2011
76 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley In shady wet valley in Acacia--Metrosideros forest which has been altered by introduced tree species, such as Aleurites moluccana and Ficus sp. With such shrubby species as Cyrtandra and Cordyline terminalis. PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Oahu BISH 6/10/2004
77 Sphaeropteris cooperi Waiopa Gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN Oppenheimer, H.L. Perlman, S. Collector Number: H90705 Lanai BISH 9/5/2007
78 Sphaeropteris cooperi honolulu; makiki; grounds of contemporary museum, 2411 makiki heights drive In garden. PRESERVED_SPECIMEN staples, g.w. medbury, j.s. collector number: 848 Oahu BISH 7/13/1992
79 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Maui BISH 6/10/2004
80 Sphaeropteris cooperi waianae range: along the summit of the waianae mountains from mauna kapu to pali kea and beyond PRESERVED_SPECIMEN wilson, k.a. palmer, d.d.; obata, j.; gustafason, b. collector number: 2519 Kauai BISH 2/6/1996
81 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H90631 Maui BISH 9/28/2006
82 Sphaeropteris cooperi east maui, hana district, hana district,kipahulu valley Growing as an aggresive weed. PRESERVED_SPECIMEN gagné, b.h. collector number: 3056 Maui BISH 4/24/1991
83 Sphaeropteris cooperi hanalilolilo sink hole With Acacia and Gouldia. PRESERVED_SPECIMEN hughes, g.d. collector number: s.n. Oahu BISH 2/10/1992
84 Sphaeropteris cooperi Hanalilolilo Sink Hole Mesic forest. PRESERVED_SPECIMEN Hughes, G.D. s.n. Molokai BISH 2/10/1992
85 Sphaeropteris cooperi w maui, hoholua, maunalei arboretum PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 187 Maui BISH 3/7/1997
86 Sphaeropteris cooperi koloa dist.; lihue-koloa forest reserve, wahiawa drainage; along main fork & upper tributaries of wahiawa stream to crest of ridge just sw of kapalaoa PRESERVED_SPECIMEN flynn, t. lorence, d.h.; perlman, s.; wood, k. collector number: 4593 Maui BISH 4/12/1991
87 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley Under bridge in ephemeral creek. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0330 Maui HAPI 11/22/2007
88 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley HUMAN_OBSERVATION Big Island Invasive Species Committee (BIISC) Maui HAPI 11/13/2008
89 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters HUMAN_OBSERVATION Big Island Invasive Species Committee (BIISC) Maui HAPI 11/13/2008
90 Sphaeropteris cooperi Oahu Forest National Wildlife Refuge, Kipapa Stream, upslope ca 300 yd from stream. Dicranopteris-dominated slope. PRESERVED_SPECIMEN Hanley, J. s.n. Oahu BISH 9/8/2011
91 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Lanai BISH 6/10/2004
92 Sphaeropteris cooperi kalihi valley, above waterfall Two plants along trail. Non-native dominated lowland mesic forest. PRESERVED_SPECIMEN us army kawelo, k. collector number: usarmy 185 Oahu BISH 5/9/2010
93 Sphaeropteris cooperi waiopa gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN oppenheimer, h.l. perlman, s. collector number: h90705 Lanai BISH 9/5/2007
94 Sphaeropteris cooperi waianae range: along the summit of the waianae mountains from mauna kapu to pali kea and beyond Locally common in sparse bushy vegetation, in seriously eroded slope of fine clay deeply weathered red volcanic soil PRESERVED_SPECIMEN wilson, k.a. palmer, d.d.; obata, j.; gustafason, b. collector number: 2519 Kauai BISH 2/6/1996
95 Sphaeropteris cooperi honolulu: kahala; 3653 kahala avenue; residence of c. f. johnston In full sun. PRESERVED_SPECIMEN lau, j. cory, c. collector number: 2136 Oahu BISH 3/17/1986
96 Sphaeropteris cooperi Waiopa Gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN Oppenheimer, H.L. H90705 Lanai BISH 9/5/2007
97 Sphaeropteris cooperi honolulu, bishop museum, shade house, back side of campus PRESERVED_SPECIMEN evans, m. collector number: 1 Oahu BISH 6/20/1990
98 Sphaeropteris cooperi Maui, road to Haleakalae Under bridge in ephemeral creek. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0330 Consortium of Pacific Northwest Herbaria 11/22/2007
99 Sphaeropteris cooperi Kalihi Valley, above waterfall Two plants along trail. Non-native dominated lowland mesic forest. PRESERVED_SPECIMEN Kawelo, K. USARMY 185 Oahu BISH 5/9/2010
100 Sphaeropteris cooperi Lawai Valley. The National Tropical Botanical Garden. Bamboo Bridge at W end of Hau Tunnel. Plants destroyed because species has become naturalized throughout island. Cultivated plants. PRESERVED_SPECIMEN Tim Flynn 5787 Kauai PTBG 4/21/1995
101 Sphaeropteris cooperi Kokee, .3 mile down Kumuwela Ridge Road. West of road off of ridge top in small, weedy gully bottom. PRESERVED_SPECIMEN Tim Flynn 1587 Kauai PTBG 3/4/1986
102 Sphaeropteris cooperi Hanalei Valley. Elev. 2220-2340 ft. Wet forest. PRESERVED_SPECIMEN Tim Flynn 5316 Kauai PTBG 3/22/1993
103 Sphaeropteris cooperi W. Maui, Honokahua, South of Valley. PRESERVED_SPECIMEN Hank Oppenheimer H60403 Maui PTBG 6/10/2004
104 Sphaeropteris cooperi Kokee State Park, Mohihi Road near emergency shower. Along the edge of roadside. PRESERVED_SPECIMEN Tim Flynn 3794 Kauai PTBG 3/1/1990
105 Sphaeropteris cooperi Lihue-Koloa Forest Reserve, Wahiawa drainage; along main fork and upper tributaries of Wahiawa stream to crest of ridge just SW of Kapalaoa. Elev. 2200-3200 ft. Diverse wet forest. PRESERVED_SPECIMEN Tim Flynn 4593 Kauai PTBG 4/13/1991
106 Sphaeropteris cooperi Limahuli Upper Preserve. Metrosideros wet forest. PRESERVED_SPECIMEN Emory Griffin-Noyes EGN 510 Kauai PTBG 2/14/2012
107 Sphaeropteris cooperi hosmer grove area, haleakala national park. outside of area planted by national park research team with trees from continental n. america. Wet forest PRESERVED_SPECIMEN leila m. shultz; john s. shultz 6395 Kauai US 12/23/1982
108 Sphaeropteris cooperi W. Maui, Maunalei Arboretum, Honolua. ASL. Closed canopy, alien dominated forest of Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, and Psidium guajava. PRESERVED_SPECIMEN J. Scott Meidell & Hank Oppenheimer 188 Maui PTBG
109 Sphaeropteris cooperi Halelea Forest Reserve. Along the powerline trail NW of the summit camp area. Stunted forest dominated by Metrosideros, Antidesma, and Dicranopteris. PRESERVED_SPECIMEN Tim Flynn 3546 Kauai PTBG 10/3/1989
110 Sphaeropteris cooperi E. Maui, east of Honomanu. In Metrosideros- Dicranopteris linearis wet forest. PRESERVED_SPECIMEN K. R. Wood 8772 Maui PTBG 10/17/2000
111 Sphaeropteris cooperi Limahuli Upper Preserve Metrosideros wet forest PRESERVED_SPECIMEN Emory Griffin-Noyes EGN 512 Kauai PTBG 2/23/2012
112 Sphaeropteris cooperi Near boundary with Waimea District. Upper part of Limahuli Valley from ridge separating Limahuli & Hanakapei Valleys (710 m alt.) down to Limahuli Stream (520 m alt.) above big waterfall. Nearly undisturbed rain forest with Metrosideros and Syzygium dominant. PRESERVED_SPECIMEN David H. Lorence 5721 Kauai PTBG 12/10/1987
113 Sphaeropteris cooperi Makaha Valley; upper valley via Milolii Rd.; at 3500 ft. contour north face of upper drainage; west of road. Elev. 3400-3600 ft. Acacia-Metrosideros Montane Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4983 Kauai PTBG 2/22/1996
114 Sphaeropteris cooperi Lawai Valley. The National Tropical Botanical Garden. Bamboo Bridge at W end of Hau Tunnel. Cultivated plants. PRESERVED_SPECIMEN Tim Flynn 5787 Kauai PTBG 4/21/1995
115 Sphaeropteris cooperi Waiopa Gulch. In gulch bottom. In lowland wet to mesic forest. PRESERVED_SPECIMEN Hank Oppenheimer H90705 Lanai PTBG 9/5/2007
116 Sphaeropteris cooperi koke`e, vicinity of wichman cabin on mohihi rd. PRESERVED_SPECIMEN Kauai US 1/17/2005
117 Sphaeropteris cooperi South drainage of upper Honopu; enter via Honopu trail; drop into gulch to north of trail 4180 ft.; Elev. 3700- 3900 feet. Metrosideros-Dicranopteris Montane Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4990 Kauai PTBG 2/25/1996
118 Sphaeropteris cooperi w maui, hoholua, maunalei arboretum Closed-canopy, alien dominated forest. Assoc. vegetation: Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, Psidium. PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 187 Maui BISH 3/7/1997
119 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN Oppenheimer, H.L. Hansen, G. Collector Number: H60403 Maui BISH 6/10/2004
120 Sphaeropteris cooperi Honolulu: Kahala; 3653 Kahala Avenue; residence of C. F. Johnston In full sun. PRESERVED_SPECIMEN Lau, J. 2136 Oahu BISH 3/17/1986
121 Sphaeropteris cooperi Honolulu; Bishop Museum In shade house at back of campus. Grown in 18' wooden box. PRESERVED_SPECIMEN Evans, M.A. 1 Oahu BISH 6/20/1990
122 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN Oppenheimer, H.L. H90631 Maui BISH 9/28/2006
123 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN Oppenheimer, H.L. H60403 Maui BISH 6/10/2004
124 Sphaeropteris cooperi east maui, hana district, kahikinui, wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h90631 Maui BISH 9/28/2006
125 Sphaeropteris cooperi east maui, hana district, hana district,kipahulu valley PRESERVED_SPECIMEN gagné, b.h. collector number: 3056 MPM 4/24/1991
126 Sphaeropteris cooperi East Maui, Hana District, Hana District,Kipahulu Valley Growing as an aggresive weed. PRESERVED_SPECIMEN Gagne, B.H. 3056 Maui BISH 4/24/1991
127 Sphaeropteris cooperi Honolulu; Makiki; grounds of Contemporary Museum, 2411 Makiki Heights Drive In garden. PRESERVED_SPECIMEN Staples, G. 848 Oahu BISH 7/13/1992
128 Sphaeropteris cooperi East Maui, Hana District, Hana District,Kipahulu Valley Growing as an aggressive weed. PRESERVED_SPECIMEN Gagne, B.H. 3056 Maui BISH 4/24/1991
129 Sphaeropteris cooperi Koloa Dist.; Lihue-Koloa Forest Reserve, Wahiawa drainage; along main fork & upper tributaries of Wahiawa stream to crest of ridge just SW of Kapalaoa Diverse wet forest dominated by Metrosideros spp., Antidesma, Psychotria, Perrottetia, Hedyotis tryblium, Cyrtandra spp., Labordia spp., Broussaisia, Dubautia, Melicope spp., Freycinetia, Syzygium, Tetraplasandra & w/mixed open patches of Dicranopteris/Diplopterygium especially on the sides & crest of ridge; invaded by Psidium cattleianum, Rubus rosifolius. PRESERVED_SPECIMEN Flynn, T. 4593 Kauai BISH 4/12/1991
130 Sphaeropteris cooperi honolulu; bishop museum In shade house at back of campus. Grown in 18' wooden box. PRESERVED_SPECIMEN evans, m. collector number: 01 Oahu BISH 6/20/1990
131 Sphaeropteris cooperi Waianae Range: along the summit of the Waianae Mountains from Mauna Kapu to Pali Kea and beyond Cloud forest on the windward side. PRESERVED_SPECIMEN Wilson, K.A. 2519 Oahu BISH 2/6/1996
132 Sphaeropteris cooperi W Maui, Hoholua, Maunalei Arboretum Closed-canopy, alien dominated forest. Assoc. vegetation: Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, Psidium. PRESERVED_SPECIMEN Meidell, J.S. 187 Maui BISH 3/7/1997
133 Sphaeropteris cooperi oahu forest national wildlife refuge, kipapa stream, upslope ca 300 yd from stream. Dicranopteris-dominated slope. PRESERVED_SPECIMEN hanley, j. collector number: s.n. Oahu BISH 9/8/2011
134 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley In shady wet valley in Acacia--Metrosideros forest which has been altered by introduced tree species, such as Aleurites moluccana and Ficus sp. With such shrubby species as Cyrtandra and Cordyline terminalis. PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Oahu BISH 6/10/2004
135 Sphaeropteris cooperi Waiopa Gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN Oppenheimer, H.L. Perlman, S. Collector Number: H90705 Lanai BISH 9/5/2007
136 Sphaeropteris cooperi honolulu; makiki; grounds of contemporary museum, 2411 makiki heights drive In garden. PRESERVED_SPECIMEN staples, g.w. medbury, j.s. collector number: 848 Oahu BISH 7/13/1992
137 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley In shade of planted Cryptomeria, with Dicranopteris ground cover. PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Maui BISH 6/10/2004
138 Sphaeropteris cooperi waianae range: along the summit of the waianae mountains from mauna kapu to pali kea and beyond PRESERVED_SPECIMEN wilson, k.a. palmer, d.d.; obata, j.; gustafason, b. collector number: 2519 Kauai BISH 2/6/1996
139 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters Single plant observed and destroyed in small gulch. Vegetation: Degraded Acacia/Metrosideros Montane Mesic Forest, w. Coprosma, Pteris, Polystichum, Sadleria, Lythrum, Dodonaea, Myrsine, Anthoxanthum, Holcus, Deschampsia, Axonopus, Physalis, Cyrtandra. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H90631 Maui BISH 9/28/2006
140 Sphaeropteris cooperi east maui, hana district, hana district,kipahulu valley Growing as an aggresive weed. PRESERVED_SPECIMEN gagné, b.h. collector number: 3056 Maui BISH 4/24/1991
141 Sphaeropteris cooperi hanalilolilo sink hole With Acacia and Gouldia. PRESERVED_SPECIMEN hughes, g.d. collector number: s.n. Oahu BISH 2/10/1992
142 Sphaeropteris cooperi Hanalilolilo Sink Hole Mesic forest. PRESERVED_SPECIMEN Hughes, G.D. s.n. Molokai BISH 2/10/1992
143 Sphaeropteris cooperi w maui, hoholua, maunalei arboretum PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 187 Maui BISH 3/7/1997
144 Sphaeropteris cooperi koloa dist.; lihue-koloa forest reserve, wahiawa drainage; along main fork & upper tributaries of wahiawa stream to crest of ridge just sw of kapalaoa PRESERVED_SPECIMEN flynn, t. lorence, d.h.; perlman, s.; wood, k. collector number: 4593 Maui BISH 4/12/1991
145 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley Under bridge in ephemeral creek. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0330 Maui HAPI 11/22/2007
146 Sphaeropteris cooperi W Maui, Lahaina District, Honokahua, S of Valley HUMAN_OBSERVATION Big Island Invasive Species Committee (BIISC) Maui HAPI 11/13/2008
147 Sphaeropteris cooperi East Maui, Hana District, Kahikinui, Wailaulau headwaters HUMAN_OBSERVATION Big Island Invasive Species Committee (BIISC) Maui HAPI 11/13/2008
148 Sphaeropteris cooperi Oahu Forest National Wildlife Refuge, Kipapa Stream, upslope ca 300 yd from stream. Dicranopteris-dominated slope. PRESERVED_SPECIMEN Hanley, J. s.n. Oahu BISH 9/8/2011
149 Sphaeropteris cooperi w maui, lahaina district, honokahua, s of valley PRESERVED_SPECIMEN oppenheimer, h.l. hansen, g. collector number: h60403 Lanai BISH 6/10/2004
150 Sphaeropteris cooperi kalihi valley, above waterfall Two plants along trail. Non-native dominated lowland mesic forest. PRESERVED_SPECIMEN us army kawelo, k. collector number: usarmy 185 Oahu BISH 5/9/2010
151 Sphaeropteris cooperi waiopa gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN oppenheimer, h.l. perlman, s. collector number: h90705 Lanai BISH 9/5/2007
152 Sphaeropteris cooperi waianae range: along the summit of the waianae mountains from mauna kapu to pali kea and beyond Locally common in sparse bushy vegetation, in seriously eroded slope of fine clay deeply weathered red volcanic soil PRESERVED_SPECIMEN wilson, k.a. palmer, d.d.; obata, j.; gustafason, b. collector number: 2519 Kauai BISH 2/6/1996
153 Sphaeropteris cooperi honolulu: kahala; 3653 kahala avenue; residence of c. f. johnston In full sun. PRESERVED_SPECIMEN lau, j. cory, c. collector number: 2136 Oahu BISH 3/17/1986
154 Sphaeropteris cooperi Waiopa Gulch Occasional, naturalized, plants destroyed when found in gulch bottom in lowland wet to mesic forest. PRESERVED_SPECIMEN Oppenheimer, H.L. H90705 Lanai BISH 9/5/2007
155 Sphaeropteris cooperi honolulu, bishop museum, shade house, back side of campus PRESERVED_SPECIMEN evans, m. collector number: 1 Oahu BISH 6/20/1990
156 Sphaeropteris cooperi Maui, road to Haleakalae Under bridge in ephemeral creek. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0330 Consortium of Pacific Northwest Herbaria 11/22/2007
157 Sphaeropteris cooperi Kalihi Valley, above waterfall Two plants along trail. Non-native dominated lowland mesic forest. PRESERVED_SPECIMEN Kawelo, K. USARMY 185 Oahu BISH 5/9/2010
158 Sphaeropteris cooperi Lawai Valley. The National Tropical Botanical Garden. Bamboo Bridge at W end of Hau Tunnel. Plants destroyed because species has become naturalized throughout island. Cultivated plants. PRESERVED_SPECIMEN Tim Flynn 5787 Kauai PTBG 4/21/1995
159 Sphaeropteris cooperi Kokee, .3 mile down Kumuwela Ridge Road. West of road off of ridge top in small, weedy gully bottom. PRESERVED_SPECIMEN Tim Flynn 1587 Kauai PTBG 3/4/1986
160 Sphaeropteris cooperi Hanalei Valley. Elev. 2220-2340 ft. Wet forest. PRESERVED_SPECIMEN Tim Flynn 5316 Kauai PTBG 3/22/1993
161 Sphaeropteris cooperi W. Maui, Honokahua, South of Valley. PRESERVED_SPECIMEN Hank Oppenheimer H60403 Maui PTBG 6/10/2004
162 Sphaeropteris cooperi Kokee State Park, Mohihi Road near emergency shower. Along the edge of roadside. PRESERVED_SPECIMEN Tim Flynn 3794 Kauai PTBG 3/1/1990
163 Sphaeropteris cooperi Lihue-Koloa Forest Reserve, Wahiawa drainage; along main fork and upper tributaries of Wahiawa stream to crest of ridge just SW of Kapalaoa. Elev. 2200-3200 ft. Diverse wet forest. PRESERVED_SPECIMEN Tim Flynn 4593 Kauai PTBG 4/13/1991
164 Sphaeropteris cooperi Limahuli Upper Preserve. Metrosideros wet forest. PRESERVED_SPECIMEN Emory Griffin-Noyes EGN 510 Kauai PTBG 2/14/2012
165 Sphaeropteris cooperi hosmer grove area, haleakala national park. outside of area planted by national park research team with trees from continental n. america. Wet forest PRESERVED_SPECIMEN leila m. shultz; john s. shultz 6395 Kauai US 12/23/1982
166 Sphaeropteris cooperi W. Maui, Maunalei Arboretum, Honolua. ASL. Closed canopy, alien dominated forest of Cinnamomum burmannii, Ardisia elliptica, Diplazium sandwichianum, Thelypteris hudsoniana, and Psidium guajava. PRESERVED_SPECIMEN J. Scott Meidell & Hank Oppenheimer 188 Maui PTBG
167 Sphaeropteris cooperi Halelea Forest Reserve. Along the powerline trail NW of the summit camp area. Stunted forest dominated by Metrosideros, Antidesma, and Dicranopteris. PRESERVED_SPECIMEN Tim Flynn 3546 Kauai PTBG 10/3/1989
168 Sphaeropteris cooperi E. Maui, east of Honomanu. In Metrosideros- Dicranopteris linearis wet forest. PRESERVED_SPECIMEN K. R. Wood 8772 Maui PTBG 10/17/2000
169 Sphaeropteris cooperi Limahuli Upper Preserve Metrosideros wet forest PRESERVED_SPECIMEN Emory Griffin-Noyes EGN 512 Kauai PTBG 2/23/2012
170 Sphaeropteris cooperi Near boundary with Waimea District. Upper part of Limahuli Valley from ridge separating Limahuli & Hanakapei Valleys (710 m alt.) down to Limahuli Stream (520 m alt.) above big waterfall. Nearly undisturbed rain forest with Metrosideros and Syzygium dominant. PRESERVED_SPECIMEN David H. Lorence 5721 Kauai PTBG 12/10/1987
171 Sphaeropteris cooperi Makaha Valley; upper valley via Milolii Rd.; at 3500 ft. contour north face of upper drainage; west of road. Elev. 3400-3600 ft. Acacia-Metrosideros Montane Mesic Forest. PRESERVED_SPECIMEN K. R. Wood 4983 Kauai PTBG 2/22/1996
172 Sphaeropteris cooperi Lawai Valley. The National Tropical Botanical Garden. Bamboo Bridge at W end of Hau Tunnel. Cultivated plants. PRESERVED_SPECIMEN Tim Flynn 5787 Kauai PTBG 4/21/1995
173 Sphaeropteris cooperi Waiopa Gulch. In gulch bottom. In lowland wet to mesic forest. PRESERVED_SPECIMEN Hank Oppenheimer H90705 Lanai PTBG 9/5/2007
174 Sphaeropteris cooperi koke`e, vicinity of wichman cabin on mohihi rd. PRESERVED_SPECIMEN Kauai US 1/17/2005
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