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Melicope radiata

(H.St.John) T.G.Hartley & B.C.Stone (1989)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Melicope

alani, alani kuahiwi

Description

Key Characters:

Growth Form: Shrubs or small trees 2–6 m tall, new growth glabrate or sparsely appressed grayish puberulent to short–pilose.

Stems:

Roots:

Leaves: Leaves simple. Opposite. Blades elliptic to ovate or sometimes suborbicular, 3–14 cm long, 3–7 cm wide. Apex acute or sometimes rounded. Base broadly cuneate, subtruncate, or subcordate. Surfaces glabrous or midrib on lower surfaces sparsely puberulent; with an anise–citrus odor when crushed; upper surfaces pale green; lower surfaces slightly paler and duller with usually reddish veins, thin, coriaceous, brittle. Margins entire. Primary lateral veins usually 7–22 pairs, inconspicuous, connected by a weakly arched vein 1–3 mm from margin, venation pinnate, lateral veins connected by an arched submarginal vein, higher order venation forming a reticulate pattern. Petioles (6–)12–36 mm long, glabrate. Stipules absent.

Flowers: Flowers 1–3 in glabrate, axillary cymes, peduncles 10–12 mm long, pedicels 8–30 mm long, bracteoles minute, deltate, 1 mm or less long, sparsely puberulent, bracts small, opposite. Calyx of 4 sepals; sepals deltate–ovate, ca. 3 mm long, glabrate to very sparsely puberulent, imbricate. Corolla of 4 petals; petals narrowly deltate–lanceolate, 7–9 mm long, glabrous, apex acute, valvate. Stamens 8, in 2 whorls, distinct, reduced but always present in functionally pistillate flowers. Ovary superior, glabrous; style slender, ca. 4 mm long, nectary disk 8-lobed, ovary 4-celled and 4-lobed, carpels connate, only slightly distinct at apex to almost entirely distinct; ovules 2 per cell, hemitropous, 1 ascending; style 1, terminating in 4 short stigmatic branches.

Fruit: Capsules green to red–tinged; 20–36 mm wide; carpels connate ¼–⅓ their length; 10–18 mm long; becoming reflexed after dehiscence; exocarp and endocarp glabrous. the carpels dehiscing along the upper suture; exocarp. Seeds 1–2 per carpel; ca. 6–7 mm long. glossy black when ripe; crustaceous; ovoid; sometimes angled from compression; embryo straight in fleshy endosperm; radicle short; cotyledons compressed; ovate; putamen brown; warty.

Ploidy:

Habitat: Occurring in dry; mesic; and wet forest and subalpine shrubland.

Elevation Range: 300–1,650 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Taxon 38: 121 (1989)

Other References

Wagner et al. 1990:1202 (H [as Pelea radiata]); Hartley & Stone 1989:121 (COMBNOV:Syn. P. radiata)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kalapana extension - above wahaula visitor center Partial shade. With Pisonia PRESERVED_SPECIMEN reeser, d.w. perman, t. Hawaii BISH
2 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kona District, Honaunau Forest Reserve, along jeep road in wet Metrosideros forest PRESERVED_SPECIMEN Herbst, D.R. 5427 Hawaii BISH 7/17/1975
3 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone N Kona Distr., Waipunalei Ahu., Tr 13(63); E slope of Mauna Kea, Hilo FR, along W fork of Kaawalii Stream Wet koa-`ohi`a forest with other native trees and a treefern, native shrub understory. PRESERVED_SPECIMEN Warshauer, F.R. FRW 1691 Hawaii BISH 1/24/1978
4 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone keauhou ranch, kau district, near rainshed on eastern boundary Growing in Pennisetum clandestinum (kikuyu) pasture with scattered Metrosideros, Cheirodendron and Perrottetia. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 648 Hawaii BISH 10/23/1980
5 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka NAR Metrosideros lowland mesic forest with 70% canopy cover @ 20-25 m tall with understory of Myrsine lessertiana, Dodonaea viscosa, Alyxia oliviformis, Pipturus albidus, Antidesma platyphyllum & A. pulvinatum, Myoporum, Tetraplasandra hawaiiense, Dryopteris fusco-atra, Asplenium polyphyllum, Lepisorus thunbergiana, Asplenium nidus PRESERVED_SPECIMEN Wood, K.R. 12345 Hawaii BISH 4/13/2007
6 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Keauhou Ranch, Kau District Disturbed open Metrosideros-Acacia koa forest. Growing in full sun in kikuyu grass. PRESERVED_SPECIMEN Davis, J. 390 Hawaii BISH 10/1/1980
7 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kopapola/punaulu/kau. PRESERVED_SPECIMEN charles n. forbes forbes 399.h MO 8/7/1911
8 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone manuka n.a.r., transect 4, above hwy 11, across kaulanamauna crossing south kona-kau dist boundary, near kahahawai. PRESERVED_SPECIMEN s. p. perlman, l. abbott & s. gon 10725 MU 4/10/1989
9 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kona side. PRESERVED_SPECIMEN Eggler, W. A. 256 Hawaii US 1965-00-00
10 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Ocean View Estates, along Hukilau Drive between Pineapple Parkway and Plumeria Lane. Aa lava flow with low, relatively dry Metrosideros shrubland. PRESERVED_SPECIMEN David H. Lorence 5868 Hawaii PTBG 3/12/1988
11 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Ocean View Estates; Hukilau Drive. Ohia successional forest on weathered lava. PRESERVED_SPECIMEN W. L. Wagner 5966 Hawaii PTBG 3/12/1988
12 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone HAWAII: *Manuka NAR, Metrosideros lowland mesic forest with 70% canopy cover @ 20-25 m tall with understory of Myrsine lessertiana, Dodonaea viscosa, Alyxia oliviformis, Pipturus albidus, Antidesma platyphyllum & A. pulvinatum, Myoporum, Tetraplasandra hawaiiense, Dryopteris fusco-atra, Asplenium polyphyllum, Lepisorus thunbergiana, Asplenium nidus PRESERVED_SPECIMEN K. R. Wood 12345 Hawaii PTBG 4/13/2007
13 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kahaualea NAR, trail to Puu O`o lava flow, * Metrosideros - Dicranopteris - Ciboteum lowland wet forest with Hedyotis terminalis, Cheirodendron, Cyrtandra platyphylla, C. paludosa, Freycinetia, Broussaisia, Metrosideros - Dicranopteris - Ciboteum lowland wet fores PRESERVED_SPECIMEN K. R. Wood 12335 Hawaii PTBG 4/12/2007
14 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka NAR, ca .5 miles south of South Kona-Kau District boundary & Kaulanamauna, makai from end of old logging road. Metrosideros polymorpha lowland mesic forest. PRESERVED_SPECIMEN Steve Perlman 10684 Hawaii PTBG 4/20/1989
15 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kaiholena Unit, TNC Ka`u Preserve, Old Plantation Springs area. Closed to open Metrosideros forest. PRESERVED_SPECIMEN K. R. Wood 12396 Hawaii PTBG 5/15/2007
16 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Honomalino; 2300 ft elev. transect for TNC survey (700m). Elev. 700-750 m. Metrosideros lowland mesic forrest. PRESERVED_SPECIMEN K. R. Wood 1529 Hawaii PTBG 12/18/1991
17 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Upper Waiakea Forest Reserve, near Stainback Highway. Wet forest. PRESERVED_SPECIMEN Marc Appelhans MA698 Hawaii PTBG 10/11/2016
18 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Hanomolino; north boundary access rd., Mac farms. Metrosideros mesic forest PRESERVED_SPECIMEN K. R. Wood 7772 Hawaii PTBG 4/8/1999
19 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka N.A.R., transect 5, east from Kaulanamauna near old logging road. Mix of pioneer veg on lava & Metrosideros montane mesic forest PRESERVED_SPECIMEN Steve Perlman 10718 Hawaii PTBG 4/13/1989
20 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kahaualea natural area reserve. PRESERVED_SPECIMEN h. l. oppenheimer h40022 Maui US 4/21/2000
21 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone NW end of Honaunau Forest Reserve, S. Kona Dist. PRESERVED_SPECIMEN St. John, H.; Baldwin, P.; Longley, P. 18530 Hawaii US 12/26/1937
22 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna forest reserve 4 miles se of belt rd. PRESERVED_SPECIMEN o. degener & i. degener 31479 Hawaii US 2/20/1968
23 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone nw end of honaunau forest reserve, s. kona dist. PRESERVED_SPECIMEN h. st. john, p. baldwin & p. longley 18530 Kauai US 12/26/1937
24 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna forest reserve, 4 miles se of belt rood PRESERVED_SPECIMEN o. degener & i. degener 31479a Hawaii US 2/20/1960
25 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna; upper Kealakomo, above pali, S of Volcano-Kalapana trail, old lava flow. PRESERVED_SPECIMEN Stone, B. C. 2994 Hawaii US 6/29/1959
26 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone manuka n.a.r., transect 5, e from kaulanamauna near old logging road. PRESERVED_SPECIMEN s. p. perlman, l. abbott & s. gon 10718 Hawaii US 4/13/1989
27 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kau district, ocean view estates, hukilau dr. PRESERVED_SPECIMEN w. l. wagner, c. imada & w. n. takeuchi 5966 Hawaii US 3/12/1988
28 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna; upper kealakomo, above pali, s of volcano-kalapana trail, old lava flow. PRESERVED_SPECIMEN b. c. stone 2994 Hawaii US 6/29/1959
29 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna Forest Reserve 4 miles SE of Belt rd. PRESERVED_SPECIMEN Degener, O.; Degener, I. 31479 Hawaii US 2/20/1968
30 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kapua PRESERVED_SPECIMEN macdaniels, l.h. collector number: 273 PH 11/16/1926
31 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Upper Waiakea Forest Reserve, near Stainback Highway. Wet forest. PRESERVED_SPECIMEN Marc Appelhans MA696 Hawaii PTBG 10/11/2016
32 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kahaualea, NAR survey transect 10, sta. 1.0-5.0, above Royal Gardens. Ohia-hapu mesophytic forest. PRESERVED_SPECIMEN Steve Perlman 10314 Hawaii PTBG 12/6/1988
33 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone HAWAII: *Kaiholena Unit, northwestern corner of TNC Ka`u Preserve, Nanuamaia to Pali Muku, * closed to open Metrosideros forest PRESERVED_SPECIMEN K. R. Wood 12404 Hawaii PTBG 5/16/2007
34 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kahaualea NAR, trail to Puu O`o lava flow, * Metrosideros - Dicranopteris - Ciboteum lowland wet forest with Hedyotis terminalis, Cheirodendron, Cyrtandra platyphylla, C. paludosa, Freycinetia, Broussaisia, Metrosideros - Dicranopteris - Ciboteum lowland wet forest PRESERVED_SPECIMEN K. R. Wood 12336 Hawaii PTBG 4/12/2007
35 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka NAR, nature trail up to Kipuka Kilioe. Metrosideros lowland mesic forest ca. 25-30 m tall. PRESERVED_SPECIMEN K. R. Wood 13207 Hawaii PTBG 8/7/2008
36 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka NAR, transect 4, above Hwy 11, across Kaulanamauna crossing South Kona-Kau Dist. Boundary, near Kahahawai. Mix of pioneer veg on lava & Metrosideros montane mesic forest PRESERVED_SPECIMEN Steve Perlman 10725 Hawaii PTBG 4/10/1989
37 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna District, Kahaualea NAR, trail to Puu O`o lava flow. Lowland wet forest. PRESERVED_SPECIMEN Wood, K. R.; Agorastos, N. 12335 Hawaii US 4/12/2007
38 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone W Maui; Wailuku District; Iao Valley; Nakalaloa Stream; Back of Valley Lowland mesic forest PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Perlman, J. 1942 Maui US 5/28/1992
39 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kau district, ocean view estates, hukilau dr. and pineapple parkway PRESERVED_SPECIMEN w. l. wagner, c. imada & w. n. takeuchi 5963 Hawaii US 3/12/1988
40 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna district, kahaualea nar, trail to puu o`o lava flow. Lowland wet forest. PRESERVED_SPECIMEN k. r. wood & n. agorastos 12335 Hawaii US 4/12/2007
41 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna Forest Reserve, 4 miles SE of Belt Rood PRESERVED_SPECIMEN Degener, O.; Degener, I. 31479a Hawaii US 2/20/1960
42 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka N.A.R., transect 5, E from Kaulanamauna near old logging road. PRESERVED_SPECIMEN Perlman, S. P.; Abbott, L.; Gon, S. 10718 Hawaii US 4/13/1989
43 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kona side. PRESERVED_SPECIMEN w. a. eggler 256 Hawaii US 8/5/1965
44 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kahaualea Natural Area Reserve. PRESERVED_SPECIMEN H. L. Oppenheimer H40022 Hawaii US 4/21/2000
45 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kahaualea Natural Area Reserve. PRESERVED_SPECIMEN Oppenheimer, H. L. H40022 Hawaii US 4/21/2000
46 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Manuka N.A.R., transect 4, above Hwy 11, across Kaulanamauna crossing South Kona-Kau Dist Boundary, near Kahahawai. PRESERVED_SPECIMEN Perlman, S. P.; Abbott, L.; Gon, S. 10725 Hawaii US 4/10/1989
47 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kau District, Ocean View Estates, Hukilau Dr. and Pineapple Parkway PRESERVED_SPECIMEN Wagner, W. L.; Imada, C. T.; Takeuchi, W. N. 5963 Hawaii US 3/12/1988
48 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone w maui; wailuku district; iao valley; nakalaloa stream; back of valley Lowland mesic forest PRESERVED_SPECIMEN k. r. wood, s. p. perlman & j. perlman 1942 Maui US 5/28/1992
49 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kona district, honaunau forest reserve, along north jeep road. PRESERVED_SPECIMEN d. r. herbst & g. spence 5427 Oahu US 7/17/1975
50 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna; upper Kealakomo. N of the pali, S of the Volcano-Kalapana trail. PRESERVED_SPECIMEN Stone, B. C.; Stone, J. G. 3089-b Hawaii US 8/10/1959
51 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kau District, Ocean View Estates, Hukilau Dr. PRESERVED_SPECIMEN Wagner, W. L.; Imada, C. T.; Takeuchi, W. N. 5966 Hawaii US 3/12/1988
52 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna; upper kealakomo. n of the pali, s of the volcano-kalapana trail. PRESERVED_SPECIMEN b. c. stone & j. g. stone 3089-b Hawaii US 8/10/1959
53 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone N Hilo Distr., Honohina Ahu., Tr 17(114); E slope of Mauna Kea, on ridge E of Opea-Peleau Homesteads wet `ohi`a forest with other native trees and an exotic and native shrub understory. growing along stream bank. PRESERVED_SPECIMEN Warshauer, F.R. FRW 1576 Hawaii BISH 11/29/1977
54 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone subdivision makai of ocean view estates, kau In branches of guava trees. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6663 Oahu BISH 6/6/1980
55 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna; upper Kealakomo In small patch of forest above pali, south of Volcano-Kalapana trail, old lava flow; with Metrosideros, Tetraplasandra, Myrsine. PRESERVED_SPECIMEN Stone, B.C. 2994 Hawaii BISH 6/29/1959
56 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kapua Rough lava PRESERVED_SPECIMEN MacDaniels, L.H. 273 Hawaii BISH 11/16/1926
57 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna; upper kealakomo, north of the pali, s of the volcano-kalapana trail PRESERVED_SPECIMEN stone, b.c. stone, j.g. collector number: 3089b Hawaii BISH 8/10/1959
58 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kohala mts., waimea PRESERVED_SPECIMEN forbes, c.n. collector number: 492.h Hawaii BISH 9/1/1911
59 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kona District, Honaunau Forest Reserve, along north jeep road. Small, sparingly branched tree. PRESERVED_SPECIMEN Herbst, D. R.; Spence, G. 5427 Hawaii US 7/17/1975
60 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kau, kapapala (puuaulu) PRESERVED_SPECIMEN forbes, c.n. collector number: 399.h Hawaii BISH 8/7/1911
61 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna forest reserve four miles southeast of belt road wet, shrubby jungle PRESERVED_SPECIMEN degener, o. degener, i. collector number: 31479 Hawaii BISH 2/20/1968
62 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna; upper Kealakomo, north of the pali, S of the Volcano-Kalapana trail dry forest on aa lava (Metrosideros, Myrsine, Tetraplasandra, Santalum, Canthium, Diospyros). The location is arid; rains are occasional and violent; the lava is extremely porous. PRESERVED_SPECIMEN Stone, B.C. 3089a Hawaii BISH 8/10/1959
63 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kohala PRESERVED_SPECIMEN Rock, J.F.C. 8374 Hawaii BISH 6/1/1910
64 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Hawaii: *Puna District, Kahaualea NAR, trail to Puu O'o lava flow *Metrosideros - Dicranopteris - Ciboteium lowland wet forest with Hedyotis terminalis, Cheirodendron, Cyrtandra platyphylla, C. paludosa, Frecinetia, Broussaisia PRESERVED_SPECIMEN Wood, K.R. 12335 Hawaii BISH 4/12/2007
65 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone NW side of Kamalo Valley Gulch. Up ridge to summit west of Kamakou Metrosideros-Styphelia-Dodonaea w/ Vaccinium, Scaevola, Wikstroemia, Osteomeles. PRESERVED_SPECIMEN Wood, K.R. 1377 Molokai BISH 11/12/1991
66 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puuwaawaa Open mesic forest (same lava tube hole as previous urera). Associated species: Urera, banana poka, ferns, Rubus hawaiensis. PRESERVED_SPECIMEN montgomery, s.l. ruffin, w. collector number: 23 Hawaii BISH 7/3/1971
67 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kalapana trail infrequent in ohia forest. PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 792 Hawaii BISH 8/17/1943
68 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone n kona distr., waipunalei ahu., tr 13(63); e slope of mauna kea, hilo fr, along w fork of kaawalii stream Wet koa-`ohi`a forest with other native trees and a treefern, native shrub understory. PRESERVED_SPECIMEN warshauer, f.r. collector number: frw 1691 Hawaii BISH 1/24/1978
69 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kau District, Ocean View Estates; Hukilau Drive `ohi`a successional forest on weathered lava PRESERVED_SPECIMEN Wagner, W.L. 5965 Hawaii BISH 3/12/1988
70 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kau district, ocean view estates, hukilau drive and pineapple parkway PRESERVED_SPECIMEN wagner, w.l. imada, c.; takeuchi, w. collector number: 5963 Oahu BISH 3/12/1988
71 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puuwaawaa Open mesic forest (same lava tube hole as previous urera). Associated species: Urera, banana poka, ferns, Rubus hawaiensis. PRESERVED_SPECIMEN Montgomery, S.L. 23 Hawaii BISH 7/3/1971
72 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone manuka nar;, nature trail up to kipuka kilioe Metrosideros lowland mesic forest ca 25 - 30 m tall. W/ understory of Nestegis, Pisonia, Charpentiera, Psychotria hawaiiensis, Claoxylon, Myrsine lessertiana, Dodonaea viscosa, Alyxia oliviformis, Pipturus albidus, Antidesma platyphyllum & A. pulvinatum. PRESERVED_SPECIMEN wood, k.r. agorastos, n. collector number: 13207 Hawaii BISH 8/7/2008
73 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kalapana trail infrequent in ohia forest. PRESERVED_SPECIMEN Fagerlund, G.O. 792 Hawaii BISH 8/17/1943
74 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Mauna Loa Truck trail Growing in partial shade PRESERVED_SPECIMEN Fagerlund, G.O. 683 Hawaii BISH 7/27/1943
75 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kilauea, Bird Park PRESERVED_SPECIMEN Neal, M.C. s.n. Hawaii BISH 8/3/1927
76 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone N Hilo Distr., Laupahoehoe Ahu., Tr 14(38); E slope of Mauna Kea, along Kilau Stream wet koa-`ohi`a forest with other native trees and a treefern, native shrub understory. PRESERVED_SPECIMEN Warshauer, F.R. FRW 1309b Hawaii BISH 8/13/1977
77 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kalapana trail in ohia forest PRESERVED_SPECIMEN Fagerlund, G.O. 796 Hawaii BISH 8/17/1943
78 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kapuka Key & Puauhi, ranch side PRESERVED_SPECIMEN Forbes, C.N. 399.H Hawaii BISH 8/7/1911
79 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kapapala near kipuka puaulu lava crevice PRESERVED_SPECIMEN takeuchi, w.n. collector number: 3657 Hawaii BISH 8/2/1987
80 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kilauea-Mauna Loa Road, Keauhou, Puna Dist Open Acacia koa forest PRESERVED_SPECIMEN St. John, H. 18411 Hawaii BISH 12/19/1937
81 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna Distr., Kamoamoa Ahu., Tr 37(53); 2 km. S of benchmark 2893, Volcano Hwy, S of Napau Crater Wet `ohi`a forest with other native trees and a treefern and native shrub understory. PRESERVED_SPECIMEN Stemmermann, R.L. RLS 3810 Hawaii BISH 6/1/1979
82 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone n hilo distr., laupahoehoe ahu., tr 14(38); e slope of mauna kea, along kilau stream dry, intermittent pond on flat above Hanauma Bay PRESERVED_SPECIMEN warshauer, f.r. collector number: frw 1309 Oahu BISH 8/13/1977
83 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone SW. rift of Mauna Loa above Ocean View subdivision, approaching Puu Ohohia in kipuka under Metrosideros-Acacia koa cover, with Styphelia, Dodonaea. PRESERVED_SPECIMEN Imada, C.T. 198 Hawaii BISH 2/16/1986
84 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kalapana Extension - Above Wahaula Visitor Center Partial shade. With Pisonia PRESERVED_SPECIMEN Reeser, D.W. s.n. Hawaii BISH
85 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone keauhou ranch, kau district Disturbed open Metrosideros-Acacia koa forest. Growing in full sun in kikuyu grass. PRESERVED_SPECIMEN davis, j. collector number: 390 Hawaii BISH 10/1/1980
86 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kau district, ocean view estates; hukilau drive `ohi`a successional forest on weathered lava PRESERVED_SPECIMEN wagner, w.l. imada, c.; takeuchi, w. collector number: 5965 Hawaii BISH 3/12/1988
87 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone s. kona distr., kealakekua ahu., tr 60(115); w slopes of mauna loa; 2-4 km. e of keopuka Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Shaded. Growing on aa. PRESERVED_SPECIMEN jacobi, j.d. collector number: jdj 1140 Hawaii BISH 8/4/1978
88 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Subdivision makai of Ocean View Estates, Kau in mixed ohia forest on aa PRESERVED_SPECIMEN Stemmermann, R.L. 6663 Hawaii BISH 6/6/1980
89 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone n hilo distr., honohina ahu., tr 17(114); e slope of mauna kea, on ridge e of opea-peleau homesteads wet `ohi`a forest with other native trees and an exotic and native shrub understory. growing along stream bank. PRESERVED_SPECIMEN warshauer, f.r. collector number: frw 1576 Hawaii BISH 11/29/1977
90 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna forest reserve 4 miles se of belt road Wet, shrubby jungle. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 31479 Hawaii BISH 12/31/1968
91 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kilauea-mauna loa road, keauhou, puna dist Open Acacia koa forest PRESERVED_SPECIMEN st.john, h. baldwin, p.; catto, r.j.; longley, p. collector number: 18411 Hawaii BISH 12/19/1937
92 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna; upper Kealakomo, north of the pali, S of the Volcano-Kalapana trail dry forest on aa lava (Metrosideros, Canthium, Diospyros, Myrsine). the location is arid PRESERVED_SPECIMEN Stone, B.C. 3089b Hawaii BISH 8/10/1959
93 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kapuka key & puauhi, ranch side PRESERVED_SPECIMEN forbes, c.n. collector number: 399.h Hawaii BISH 8/7/1911
94 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone southwest flank of mauna loa Vegetation: Closed Metrosideros; substrate mixed soil and aa rock outcrops; understory consisting of Athyrium, Cibotium, Cyrtomium, Nephrolepis, Psidium and Psychotria. PRESERVED_SPECIMEN higashino, p.k. meyer, p. collector number: 6047 Hawaii BISH 7/5/1977
95 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kapapala near Kipuka Puaulu lava crevice PRESERVED_SPECIMEN Takeuchi, W.N. 3657 Hawaii BISH 8/2/1987
96 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kau District, Ocean View Estates, Hukilau Drive and Pineapple Parkway on lava PRESERVED_SPECIMEN Wagner, W.L. 5963 Hawaii BISH 3/12/1988
97 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna; upper kealakomo In small patch of forest above pali, south of Volcano-Kalapana trail, old lava flow; with Metrosideros, Tetraplasandra, Myrsine. PRESERVED_SPECIMEN stone, b.c. collector number: 2994 Hawaii BISH 6/29/1959
98 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone so. of kalapana trail Metrosideros, Cibotium forest; with Cyrtandra, Perrottetia, Pipturus, Psychotria, Liparis PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 798 Hawaii BISH 8/17/1943
99 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kau District, Ocean View Estates; Hukilau Drive `ohi`a successional forest on weathered lava PRESERVED_SPECIMEN Wagner, W.L. 5966 Hawaii BISH 3/12/1988
100 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kona district, honaunau forest reserve, along jeep road in wet Metrosideros forest PRESERVED_SPECIMEN herbst, d.r. spence, g. collector number: 5427 Hawaii BISH 7/17/1975
101 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kohala District, above Waimea Reservoir, E. of Mt. Kaala PRESERVED_SPECIMEN Neal, M.C. 3 Hawaii BISH 6/6/1964
102 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone nw side of kamalo valley gulch. up ridge to summit west of kamakou Metrosideros-Styphelia-Dodonaea w/ Vaccinium, Scaevola, Wikstroemia, Osteomeles. PRESERVED_SPECIMEN wood, k.r. garnett, w.; perlman, s. collector number: 1377 Molokai BISH 11/12/1991
103 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone south kona distr., nw end of honaunau forest reserve Rain forest. PRESERVED_SPECIMEN st.john, h. baldwin, p.; longley, p. collector number: 18530 Hawaii BISH 12/26/1937
104 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kohala district, above waimea reservoir, e. of mt. kaala Metrosideros woods. PRESERVED_SPECIMEN neal, m.c. collector number: 3 Hawaii BISH 6/6/1964
105 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kau district, ocean view estates; hukilau drive `ohi`a successional forest on weathered lava PRESERVED_SPECIMEN wagner, w.l. imada, c.; takeuchi, w. collector number: 5966 Hawaii BISH 3/12/1988
106 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna Forest Reserve 4 miles SE of Belt Road Wet, shrubby jungle. PRESERVED_SPECIMEN Degener, O. 31479 Hawaii BISH 2/20/1968
107 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Keauhou Ranch, Kau District, near rainshed on eastern boundary Growing in Pennisetum clandestinum (kikuyu) pasture with scattered Metrosideros, Cheirodendron and Perrottetia. PRESERVED_SPECIMEN Cuddihy, L.W. 648 Hawaii BISH 10/23/1980
108 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone manuka nar Metrosideros lowland mesic forest with 70% canopy cover @ 20-25 m tall with understory of Myrsine lessertiana, Dodonaea viscosa, Alyxia oliviformis, Pipturus albidus, Antidesma platyphyllum & A. pulvinatum, Myoporum, Tetraplasandra hawaiiense, Dryopteris fusco-atra, Asplenium polyphyllum, Lepisorus thunbergiana, Asplenium nidus PRESERVED_SPECIMEN wood, k.r. agorastos, n. collector number: 12345 Hawaii BISH 4/13/2007
109 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Kohala Mts., Waimea PRESERVED_SPECIMEN Forbes, C.N. 492.H Hawaii BISH 9/1/1911
110 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone puna distr., kamoamoa ahu., tr 37(53); 2 km. s of benchmark 2893, volcano hwy, s of napau crater PRESERVED_SPECIMEN stemmermann, r.l. montgomery, s. collector number: rls 3810 Hawaii BISH 6/1/1979
111 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Southwest flank of Mauna Loa Vegetation: Closed Metrosideros; substrate mixed soil and aa rock outcrops; understory consisting of Athyrium, Cibotium, Cyrtomium, Nephrolepis, Psidium and Psychotria. PRESERVED_SPECIMEN Higashino, P.K. 6047 Hawaii BISH 7/5/1977
112 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kalapana trail in ohia forest PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 796 Hawaii BISH 8/17/1943
113 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Puna Forest Reserve four miles southeast of Belt Road wet, shrubby jungle PRESERVED_SPECIMEN Degener, O. 31479 Hawaii BISH 2/20/1968
114 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone Keauhou Ranch, Kau District Disturbed open Metrosideros-Acacia koa forest. Growing in full sun in kikuyu grass. PRESERVED_SPECIMEN Davis, J. Collector Number: 390 Hawaii BISH 10/1/1980
115 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone hawaii: *puna district, kahaualea nar, trail to puu o'o lava flow PRESERVED_SPECIMEN wood, k.r. agorastos, n. collector number: 12335 Oahu BISH 4/12/2007
116 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone mauna loa truck trail Intertidal PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 683 Hawaii BISH 7/27/1943
117 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone kohala PRESERVED_SPECIMEN rock, j.f.c. collector number: 8374 Oahu BISH 6/1/1910
118 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone sw. rift of mauna loa above ocean view subdivision, approaching puu ohohia in kipuka under Metrosideros-Acacia koa cover, with Styphelia, Dodonaea. PRESERVED_SPECIMEN imada, c.t. tokushige, w.; et al. collector number: 198 Hawaii BISH 2/16/1986
119 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone S. Kona Distr., Kealakekua Ahu., Tr 60(115); W slopes of Mauna Loa; 2-4 km. E of Keopuka Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Shaded. Growing on aa. PRESERVED_SPECIMEN Jacobi, J.D. JDJ 1140 Hawaii BISH 8/4/1978
120 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone South Kona Distr., NW end of Honaunau Forest Reserve Rain forest. PRESERVED_SPECIMEN St. John, H. 18530 Hawaii BISH 12/26/1937
121 Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone So. of Kalapana trail ohia-uluhe forest PRESERVED_SPECIMEN Fagerlund, G.O. 798 Hawaii BISH 8/17/1943
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