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Spermacoce remota

Lam. (1792)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Rubiaceae Genus: Spermacoce

buttonweed

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Description

Key Characters:

Growth Form: Annual or short-lived perennial herbs.

Stems: Stems often purple-tinged, erect or sprawling, 3-10 dm long, glabrous or sparsely retrorsely puberulent in lines on the angles.

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly elliptic, elliptic-lanceolate, or ovate, 1-7 cm long 0.4-3 cm wide. Apex acute to attenuate. Base rounded to cuneate. Blades often purple-tinged, glabrous except puberulent near margins, giving a slightly scabrid texture. Margins with 1 to numerous filiform setae. Petioles 2-4 mm long. Stipules 2-3 mm long, adnate to petioles, forming a sheath, with 5-7 slender, unequal setae 2-5 mm long, stipules of flower-bearing nodes with numerous setae.

Flowers: Flowers numerous in axillary and terminal, dense, sessile clusters, stipule-derived bracteoles with setae 2-3 mm long, subtended by 1-2 or more pairs of bract-like leaves, stipule-derived bracteoles with setae; hypanthium obconical, 2-2.5 mm long. Flowers usually small, bisexual (perfect), sessile, insect-pollinated, often heterostylous. Calyx lobes 2-4(-8) with open aestivation, deltate, ca. 0.6 mm long, ciliate, separated by a fringe. Corolla funnelform or salverform, white or lobes tinged pink, 2.4-2.6 mm long, valvate in bud, the tube usually very slender; nectary disk usually present. Stamens 4, alternate with the corolla lobes, inserted on corolla tube or throat, usually exserted; anthers dithecal, opening by longitudinal slits. Ovary inferior, occasionally partly so, or very rarely superior, 2-celled; ovules 1 per cell, attached to middle of septum; style many as carpels, terminal, slender, filiform, exserted; stigmas lobed or capitate, dry or occasionally wet.

Fruit: Fruit ellipsoid or obovoid-fusiform; 2-4 mm long; pubescent in upper part; dehiscent; the valves bifid; falling completely away. Seeds 2; dull dark brown; oblong-ellipsoid; 1.5-2.5 mm long; transversely sulcate; with well-developed oily endosperm; or endosperm occasionally scanty or absent.

Ploidy:

Habitat: Naturalized and often common in urban areas; along roadsides; and other low elevation disturbed sites.

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: Tabl. Encycl. 1: 273 (1792)

Other References

Wagner et al. 1990:1173 (K, O, M, H [as S. assurgens]); Lorence et al. 1995:51/Lorence & Flynn 2006:4/Lorence & Faccenda 2024 (KEY); Oppenheimer 2003:23 (Mo, L); Acevedo-Rodríguez & Strong 2012:847 (Syn. S. assurgens = S. remota)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Spermacoce remota near Olaa, Keaau Macadamia Orchard Preserved_Specimen Mangelsdorf, A.J. Hawaii BISH 1962-07-26
2 Spermacoce remota Stainback Highway Preserved_Specimen Degener, O. 31587 Hawaii BISH 1967-07-18
3 Spermacoce remota grounds of Pacific Tropical Botanical Garden Preserved_Specimen Wagner, W.L. 5139 Kauai BISH 1983-12-20
4 Spermacoce remota East Maui, Haleakala National Park in Pipiwai exclosure below bridges Preserved_Specimen Welton, P. 2457 Maui BISH 2020-10-27
5 Spermacoce remota Manoa, Cooper Rd., Honolulu Preserved_Specimen Egler, F.E. 37422 Oahu BISH 1937-06-03
6 Spermacoce remota roadside, Hilo Preserved_Specimen Degener, O. 32432 Hawaii BISH 1966-05-20
7 Spermacoce remota Kauman substation Preserved_Specimen Wagner, W.L. 4632 Hawaii BISH 1982-11-16
8 Spermacoce remota 2062A St. Louis Dr., Honolulu Preserved_Specimen Wagner, W.L. Oahu BISH 1988-04-27
9 Spermacoce remota Exp.. Sta. Grounds Preserved_Specimen Caum, E.L. Oahu BISH 1935-01-21
10 Spermacoce remota Honolulu Preserved_Specimen Woolford, H.A. Oahu BISH 1950-03-31
11 Spermacoce remota Kauai Preserved_Specimen Forbes, C.N. Kauai BISH 1940-02-12
12 Spermacoce remota Visitor Center and main ruins, Puuhonua o Honaunau National Historical Park Preserved_Specimen Higashino, P.K. 10294 Hawaii BISH 1984-04-08
13 Spermacoce remota Manele and Hulupoe Bays In lawns. Preserved_Specimen Oppenheimer, H.L. 40116 Lanai BISH 2001-04-15
14 Spermacoce remota Honolulu, Punahou area, 1940 Manoa Rd A weed in lawn and garden growing in full sun or partial shade. Preserved_Specimen The flowers are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
15 Spermacoce remota Manoa Campus, Univ. Hawaii, Honolulu Common roadside weed. Preserved_Specimen Flws: white-pink. Swarbrick, J.T. 7 Oahu BISH 1976-01-10
16 Spermacoce remota W Maui, Lahaina District, Napili Growing in unnamed gulch used as dump site. Preserved_Specimen Oppenheimer, H.L. 20128 Maui BISH 2001-02-21
17 Spermacoce remota Makawao District, disturbed roadside on Hwy side of Kanaha Pond (Haleakala Hwy) Dominated by grasses, Pluchea indica and P. symphytifolia. Preserved_Specimen Stems prostrate (perhaps due to continual mowing); corolla white, purple-tinged toward apex. Wagner, W.L. 5796 Maui BISH 1988-03-03
18 Spermacoce remota Mauna Kea (Puu Laau Road) In open mamane-naio forest. Preserved_Specimen Engilis Jr., A. 18 Hawaii BISH 1987-08-19
19 Spermacoce remota Punahou area, Honolulu. 1940 Manoa Rd In lawn and garden growing in full sun or partial shade. Preserved_Specimen A weed. The flws are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
20 Spermacoce remota Puna District, Keaau roadside Preserved_Specimen weed with white flowers Herbst, D.R. 5531 Hawaii BISH 1975-10-22
21 Spermacoce remota East Maui, Makawao District, Honokala, vicinity of Hana Hwy In waste area. Preserved_Specimen 1m tall herbs forming tangles; corollas white tinged with pink. Oppenheimer, H.L. 70203 Maui BISH 2002-07-14
22 Spermacoce remota U.S. Experimental Tree Planting site (Forest Reserve) within Halepuaa Forest Reserve in teh Puna District Common on the roadsides, trailsides, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
23 Spermacoce remota Along road at Haena State Park Area along road. Preserved_Specimen Weed. Corn, C. Kauai BISH 1978-06-15
24 Spermacoce remota Koloa Dist., Kalaheo, Kalawai Park Secondary vegetation of Syzygium cumini, Psidium guajava, and Ficus microcarpa. Common weed in open areas. Preserved_Specimen Perennial herb with suberect, purplish stems. Corolla pink. Lorence, D.H. 5210 Kauai BISH 1987-05-10
25 Spermacoce remota Kahuku, Honolulu Slope: 2. Soil: sandy. Vegetation type: Coastal plants. Dist: Dry and wet regions. Preserved_Specimen Slightly hairy plants spreading branches. Guinto, J. 32 Oahu BISH 1986-02-04
26 Spermacoce remota Kawaihau Dist., Nonou Forest Reserve, crest of Sleeping Giant Mountain Secondary forest of guava, Java plum, and Schinus, with forestry plantings and rare native remnants. Locally abundant in grassy clearing; population uniform. Naturalized. Preserved_Specimen Low, perennial herb, stems erect-spreading, corolla white. Lorence, D.H. 5550 Kauai BISH 1987-09-13
27 Spermacoce remota 2525 Nuuanu Ave Weed between coral pavement blocks, moist places. Preserved_Specimen Weed. Judd, C.S. Oahu BISH 1936-12-05
28 Spermacoce remota East Maui, Hana District, Paakea, along Hana Hwy Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 60318 Maui BISH 2003-06-22
29 Spermacoce remota Kahuku Training Area, from main Delta road, took left makai towards ICA-21, on ridge between Keaaulu and Lamaloa Gulches, a bit less than halfway to the big open red dirt clearing. Associated canopy: Leucaena leucocephala, Psidium cattleyanum, Syzygium cumini, Chrysophyllum oliviforme. Understory: Clidemia hirta, Dicranopteris linearis, Mimosa pudica, Paspalum conjugatum, Acacia confusa. Area shaded and damp. Preserved_Specimen Naturalized herb in lush patch ca 0.5 m tall, several meters wide. Upright but delicate and sprawling. Leaves opposite, up to 10 cm long. Flowers small, white, hairs on petals, buds prickly. Immature fruit. Inflorescence with faint pleasant scent. Thick patch represents either one massive clone or hundreds of individuals. Differs from Spermacoce assurgens in longer leaves and overall gestalt; growth in a dense patch also seemed unusual. Beachy, J. 512 Oahu BISH 2019-03-25
30 Spermacoce remota In tall grass in the Wainiha Hui lands, Haena, Kauai Wet depression. Spermacoce is common in the area as are California grass, Paspalum, Amaranthus, and tropical almond. Collection site is weedy. Preserved_Specimen Plants are very rangy with weak stems. Funk, E.J. 801 Kauai BISH 1990-05-03
31 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
32 Spermacoce remota University of Hawaii Campus, Maile Way, Honolulu Roadside. Preserved_Specimen Flowers white to pinkish. Fosberg, F.R. 10528 Oahu BISH 1933-04-09
33 Spermacoce remota Hana District; Hana, along Uwala road near Hana airport Road side weed Preserved_Specimen Alien veg. Weed, 1 foot ht with flowers and fruit Perlman, S. 22924 Maui BISH 2012-06-20
34 Spermacoce remota Lihue Dist., Sugar cane road E of Omoe Ruderal. Preserved_Specimen Herbs with erect branches to 40cm, flws: white with reddish tips of corolla lobes, seeds dark brown. Kiehn, M. 90091722 Kauai BISH 1990-09-17
35 Spermacoce remota Kalaupapa; Bayview, Quonset lawn, weedy Preserved_Specimen Wysong, M.L. 786 Molokai BISH 2005-08-12
36 Spermacoce remota East Maui, Hana District, Kipahulu Growing on side of rd. Preserved_Specimen 1m tall shrubs; flowers white w. pink tint. Oppenheimer, H.L. 60115 Maui BISH 2001-06-11
37 Spermacoce remota Kaunakakai, near the wharf In lawn. Preserved_Specimen Oppenheimer, H.L. 70111 Molokai BISH 2001-07-14
38 Spermacoce remota Limahuli Valley (Botanical Garden), Napali Coast Common along paths and around buildings. Preserved_Specimen Branched, stems ascending, lvs dark purplish green, flws: pinkish white, corollas well exserted. Fosberg, F.R. 57443 Kauai BISH 1978-03-31
39 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch Growing in lawn. Preserved_Specimen Oppenheimer, H.L. 110109 Maui BISH 2001-11-08
40 Spermacoce remota Herbarium of the H.L. Lyon Arboretum. On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
41 Spermacoce remota Woodlawn, Manoa, Honolulu Roadside weed patch. Preserved_Specimen Flowers pinkish white. Fosberg, F.R. 14111 Oahu BISH 1937-06-19
42 Spermacoce remota C.M.Cooke, Jr. Ranch, Maunawili, Koolau Mts., Kailua Near dairy buildings Preserved_Specimen Weed. Flowers white Fosberg, F.R. 10772 Oahu BISH 1935-01-23
43 Spermacoce remota East Maui, Hana District, Coastal pasture lands NW of Ka Iwi O Pele in Casuarina windbreak With Plantago, Digitaria and Oxalis. Preserved_Specimen Flws: white. Flynn, T.W. 2588 Maui BISH 1987-10-30
44 Spermacoce remota Hualalai (Olma Tract, Kaloka Mauka) Ohia-fern mesic forest Preserved_Specimen Engilis Jr., A. 31 Hawaii BISH 1987-08-19
45 Spermacoce remota On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
46 Spermacoce remota Woodlawn, Manoa Valley Naturalized roadside weed. Preserved_Specimen Degener, O. 27707 Oahu BISH 1929-01-13
47 Spermacoce remota P.T.B.G., Lawai Valley Common. Preserved_Specimen Herb with dark green foliage, white flws. Herbst, D.R. 2276 Kauai BISH 1971-12-30
48 Spermacoce remota U.S. Experimental Tree Planting Site ( Forest Reserve) within Halepuaa Forest Reserve in the Puna District On the roadside, trailside, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
49 Spermacoce remota Kualapuu, near road to gulf course Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 110127 Molokai BISH 2001-11-18
50 Spermacoce remota Tantalus Dr. ca 200m above the last house of the development Common plant on roadside Preserved_Specimen Common Faccenda, K. 1878 Oahu BISH 2021-05-22
51 Spermacoce remota W Maui, Wailuku District, Kahului In lawns and waste areas. Preserved_Specimen Weed. Oppenheimer, H.L. 40136 Maui BISH 2001-04-23
52 Spermacoce remota Kaimi Farm, Kailua Koolaupoko Growing in moist road back. Preserved_Specimen 1-1 1/2' tall, flws white. Hosaka, E.Y. 2438 Oahu BISH 1939-02-05
53 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
54 Spermacoce remota Kahuku. roadside Slope: 1%. Soil: Sandy loam. Vegetation type: tropical. Preserved_Specimen Flws: white, 1/12 across. Plant can be downy or smooth. Smith, M. 30 Oahu BISH 1986-02-04
55 Spermacoce remota East Maui, Halehaku, vicinity of Papalua Gulch along Hana Hwy Roadside. Preserved_Specimen 1m tall weed. Oppenheimer, H.L. 110118 Maui BISH 2001-11-12
56 Spermacoce remota 3rd, Mountain View, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/21/2007
57 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
58 Spermacoce remota Hilo Airport Preserved_Specimen Stemmermann, R.L. 7129 Hawaii BISH 1986-11-11
59 Spermacoce remota Kipahulu Preserved_Specimen Dnse patches sprawling over grass in rich, wet, lowland pastures. Hobdy, R.W. 3881 Maui BISH 1995-09-07
60 Spermacoce remota Kahana Valley State Park Preserved_Specimen Herb 0.2-0.3m tall; flower petals pink. Annable, C.R. 3105 Oahu BISH 1996-11-04
61 Spermacoce remota Hilo District, Coconut Island Preserved_Specimen flowers white, ripe fruit only on old (already dry) plants Kiehn, M. 89072711 Hawaii BISH 1989-07-27
62 Spermacoce remota East Maui, Kipahulu Preserved_Specimen Welton, P. 1672 Maui BISH 1992-08-27
63 Spermacoce remota W Maui, Lahaina District, Napili OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
64 Spermacoce remota Hilo, parking lot at Tropical Rent-a-Car, disturbed area Preserved_Specimen Wagner, W.L. 5684 Hawaii BISH 1986-06-12
65 Spermacoce remota near Olaa, Keaau Macadamia Orchard Preserved_Specimen Mangelsdorf, A.J. Hawaii BISH 1962-07-26
66 Spermacoce remota Stainback Highway Preserved_Specimen Degener, O. 31587 Hawaii BISH 1967-07-18
67 Spermacoce remota grounds of Pacific Tropical Botanical Garden Preserved_Specimen Wagner, W.L. 5139 Kauai BISH 1983-12-20
68 Spermacoce remota East Maui, Haleakala National Park in Pipiwai exclosure below bridges Preserved_Specimen Welton, P. 2457 Maui BISH 2020-10-27
69 Spermacoce remota Manoa, Cooper Rd., Honolulu Preserved_Specimen Egler, F.E. 37422 Oahu BISH 1937-06-03
70 Spermacoce remota roadside, Hilo Preserved_Specimen Degener, O. 32432 Hawaii BISH 1966-05-20
71 Spermacoce remota Kauman substation Preserved_Specimen Wagner, W.L. 4632 Hawaii BISH 1982-11-16
72 Spermacoce remota 2062A St. Louis Dr., Honolulu Preserved_Specimen Wagner, W.L. Oahu BISH 1988-04-27
73 Spermacoce remota Exp.. Sta. Grounds Preserved_Specimen Caum, E.L. Oahu BISH 1935-01-21
74 Spermacoce remota Honolulu Preserved_Specimen Woolford, H.A. Oahu BISH 1950-03-31
75 Spermacoce remota Kauai Preserved_Specimen Forbes, C.N. Kauai BISH 1940-02-12
76 Spermacoce remota Visitor Center and main ruins, Puuhonua o Honaunau National Historical Park Preserved_Specimen Higashino, P.K. 10294 Hawaii BISH 1984-04-08
77 Spermacoce remota Manele and Hulupoe Bays In lawns. Preserved_Specimen Oppenheimer, H.L. 40116 Lanai BISH 2001-04-15
78 Spermacoce remota Honolulu, Punahou area, 1940 Manoa Rd A weed in lawn and garden growing in full sun or partial shade. Preserved_Specimen The flowers are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
79 Spermacoce remota Manoa Campus, Univ. Hawaii, Honolulu Common roadside weed. Preserved_Specimen Flws: white-pink. Swarbrick, J.T. 7 Oahu BISH 1976-01-10
80 Spermacoce remota W Maui, Lahaina District, Napili Growing in unnamed gulch used as dump site. Preserved_Specimen Oppenheimer, H.L. 20128 Maui BISH 2001-02-21
81 Spermacoce remota Makawao District, disturbed roadside on Hwy side of Kanaha Pond (Haleakala Hwy) Dominated by grasses, Pluchea indica and P. symphytifolia. Preserved_Specimen Stems prostrate (perhaps due to continual mowing); corolla white, purple-tinged toward apex. Wagner, W.L. 5796 Maui BISH 1988-03-03
82 Spermacoce remota Mauna Kea (Puu Laau Road) In open mamane-naio forest. Preserved_Specimen Engilis Jr., A. 18 Hawaii BISH 1987-08-19
83 Spermacoce remota Punahou area, Honolulu. 1940 Manoa Rd In lawn and garden growing in full sun or partial shade. Preserved_Specimen A weed. The flws are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
84 Spermacoce remota Puna District, Keaau roadside Preserved_Specimen weed with white flowers Herbst, D.R. 5531 Hawaii BISH 1975-10-22
85 Spermacoce remota East Maui, Makawao District, Honokala, vicinity of Hana Hwy In waste area. Preserved_Specimen 1m tall herbs forming tangles; corollas white tinged with pink. Oppenheimer, H.L. 70203 Maui BISH 2002-07-14
86 Spermacoce remota U.S. Experimental Tree Planting site (Forest Reserve) within Halepuaa Forest Reserve in teh Puna District Common on the roadsides, trailsides, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
87 Spermacoce remota Along road at Haena State Park Area along road. Preserved_Specimen Weed. Corn, C. Kauai BISH 1978-06-15
88 Spermacoce remota Koloa Dist., Kalaheo, Kalawai Park Secondary vegetation of Syzygium cumini, Psidium guajava, and Ficus microcarpa. Common weed in open areas. Preserved_Specimen Perennial herb with suberect, purplish stems. Corolla pink. Lorence, D.H. 5210 Kauai BISH 1987-05-10
89 Spermacoce remota Kahuku, Honolulu Slope: 2. Soil: sandy. Vegetation type: Coastal plants. Dist: Dry and wet regions. Preserved_Specimen Slightly hairy plants spreading branches. Guinto, J. 32 Oahu BISH 1986-02-04
90 Spermacoce remota Kawaihau Dist., Nonou Forest Reserve, crest of Sleeping Giant Mountain Secondary forest of guava, Java plum, and Schinus, with forestry plantings and rare native remnants. Locally abundant in grassy clearing; population uniform. Naturalized. Preserved_Specimen Low, perennial herb, stems erect-spreading, corolla white. Lorence, D.H. 5550 Kauai BISH 1987-09-13
91 Spermacoce remota 2525 Nuuanu Ave Weed between coral pavement blocks, moist places. Preserved_Specimen Weed. Judd, C.S. Oahu BISH 1936-12-05
92 Spermacoce remota East Maui, Hana District, Paakea, along Hana Hwy Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 60318 Maui BISH 2003-06-22
93 Spermacoce remota Kahuku Training Area, from main Delta road, took left makai towards ICA-21, on ridge between Keaaulu and Lamaloa Gulches, a bit less than halfway to the big open red dirt clearing. Associated canopy: Leucaena leucocephala, Psidium cattleyanum, Syzygium cumini, Chrysophyllum oliviforme. Understory: Clidemia hirta, Dicranopteris linearis, Mimosa pudica, Paspalum conjugatum, Acacia confusa. Area shaded and damp. Preserved_Specimen Naturalized herb in lush patch ca 0.5 m tall, several meters wide. Upright but delicate and sprawling. Leaves opposite, up to 10 cm long. Flowers small, white, hairs on petals, buds prickly. Immature fruit. Inflorescence with faint pleasant scent. Thick patch represents either one massive clone or hundreds of individuals. Differs from Spermacoce assurgens in longer leaves and overall gestalt; growth in a dense patch also seemed unusual. Beachy, J. 512 Oahu BISH 2019-03-25
94 Spermacoce remota In tall grass in the Wainiha Hui lands, Haena, Kauai Wet depression. Spermacoce is common in the area as are California grass, Paspalum, Amaranthus, and tropical almond. Collection site is weedy. Preserved_Specimen Plants are very rangy with weak stems. Funk, E.J. 801 Kauai BISH 1990-05-03
95 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
96 Spermacoce remota University of Hawaii Campus, Maile Way, Honolulu Roadside. Preserved_Specimen Flowers white to pinkish. Fosberg, F.R. 10528 Oahu BISH 1933-04-09
97 Spermacoce remota Hana District; Hana, along Uwala road near Hana airport Road side weed Preserved_Specimen Alien veg. Weed, 1 foot ht with flowers and fruit Perlman, S. 22924 Maui BISH 2012-06-20
98 Spermacoce remota Lihue Dist., Sugar cane road E of Omoe Ruderal. Preserved_Specimen Herbs with erect branches to 40cm, flws: white with reddish tips of corolla lobes, seeds dark brown. Kiehn, M. 90091722 Kauai BISH 1990-09-17
99 Spermacoce remota Kalaupapa; Bayview, Quonset lawn, weedy Preserved_Specimen Wysong, M.L. 786 Molokai BISH 2005-08-12
100 Spermacoce remota East Maui, Hana District, Kipahulu Growing on side of rd. Preserved_Specimen 1m tall shrubs; flowers white w. pink tint. Oppenheimer, H.L. 60115 Maui BISH 2001-06-11
101 Spermacoce remota Kaunakakai, near the wharf In lawn. Preserved_Specimen Oppenheimer, H.L. 70111 Molokai BISH 2001-07-14
102 Spermacoce remota Limahuli Valley (Botanical Garden), Napali Coast Common along paths and around buildings. Preserved_Specimen Branched, stems ascending, lvs dark purplish green, flws: pinkish white, corollas well exserted. Fosberg, F.R. 57443 Kauai BISH 1978-03-31
103 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch Growing in lawn. Preserved_Specimen Oppenheimer, H.L. 110109 Maui BISH 2001-11-08
104 Spermacoce remota Herbarium of the H.L. Lyon Arboretum. On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
105 Spermacoce remota Woodlawn, Manoa, Honolulu Roadside weed patch. Preserved_Specimen Flowers pinkish white. Fosberg, F.R. 14111 Oahu BISH 1937-06-19
106 Spermacoce remota C.M.Cooke, Jr. Ranch, Maunawili, Koolau Mts., Kailua Near dairy buildings Preserved_Specimen Weed. Flowers white Fosberg, F.R. 10772 Oahu BISH 1935-01-23
107 Spermacoce remota East Maui, Hana District, Coastal pasture lands NW of Ka Iwi O Pele in Casuarina windbreak With Plantago, Digitaria and Oxalis. Preserved_Specimen Flws: white. Flynn, T.W. 2588 Maui BISH 1987-10-30
108 Spermacoce remota Hualalai (Olma Tract, Kaloka Mauka) Ohia-fern mesic forest Preserved_Specimen Engilis Jr., A. 31 Hawaii BISH 1987-08-19
109 Spermacoce remota On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
110 Spermacoce remota Woodlawn, Manoa Valley Naturalized roadside weed. Preserved_Specimen Degener, O. 27707 Oahu BISH 1929-01-13
111 Spermacoce remota P.T.B.G., Lawai Valley Common. Preserved_Specimen Herb with dark green foliage, white flws. Herbst, D.R. 2276 Kauai BISH 1971-12-30
112 Spermacoce remota U.S. Experimental Tree Planting Site ( Forest Reserve) within Halepuaa Forest Reserve in the Puna District On the roadside, trailside, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
113 Spermacoce remota Kualapuu, near road to gulf course Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 110127 Molokai BISH 2001-11-18
114 Spermacoce remota Tantalus Dr. ca 200m above the last house of the development Common plant on roadside Preserved_Specimen Common Faccenda, K. 1878 Oahu BISH 2021-05-22
115 Spermacoce remota W Maui, Wailuku District, Kahului In lawns and waste areas. Preserved_Specimen Weed. Oppenheimer, H.L. 40136 Maui BISH 2001-04-23
116 Spermacoce remota Kaimi Farm, Kailua Koolaupoko Growing in moist road back. Preserved_Specimen 1-1 1/2' tall, flws white. Hosaka, E.Y. 2438 Oahu BISH 1939-02-05
117 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
118 Spermacoce remota Kahuku. roadside Slope: 1%. Soil: Sandy loam. Vegetation type: tropical. Preserved_Specimen Flws: white, 1/12 across. Plant can be downy or smooth. Smith, M. 30 Oahu BISH 1986-02-04
119 Spermacoce remota East Maui, Halehaku, vicinity of Papalua Gulch along Hana Hwy Roadside. Preserved_Specimen 1m tall weed. Oppenheimer, H.L. 110118 Maui BISH 2001-11-12
120 Spermacoce remota 3rd, Mountain View, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/21/2007
121 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
122 Spermacoce remota Hilo Airport Preserved_Specimen Stemmermann, R.L. 7129 Hawaii BISH 1986-11-11
123 Spermacoce remota Kipahulu Preserved_Specimen Dnse patches sprawling over grass in rich, wet, lowland pastures. Hobdy, R.W. 3881 Maui BISH 1995-09-07
124 Spermacoce remota Kahana Valley State Park Preserved_Specimen Herb 0.2-0.3m tall; flower petals pink. Annable, C.R. 3105 Oahu BISH 1996-11-04
125 Spermacoce remota Hilo District, Coconut Island Preserved_Specimen flowers white, ripe fruit only on old (already dry) plants Kiehn, M. 89072711 Hawaii BISH 1989-07-27
126 Spermacoce remota East Maui, Kipahulu Preserved_Specimen Welton, P. 1672 Maui BISH 1992-08-27
127 Spermacoce remota W Maui, Lahaina District, Napili OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
128 Spermacoce remota Hilo, parking lot at Tropical Rent-a-Car, disturbed area Preserved_Specimen Wagner, W.L. 5684 Hawaii BISH 1986-06-12
129 Spermacoce remota near Olaa, Keaau Macadamia Orchard Preserved_Specimen Mangelsdorf, A.J. Hawaii BISH 1962-07-26
130 Spermacoce remota Stainback Highway Preserved_Specimen Degener, O. 31587 Hawaii BISH 1967-07-18
131 Spermacoce remota grounds of Pacific Tropical Botanical Garden Preserved_Specimen Wagner, W.L. 5139 Kauai BISH 1983-12-20
132 Spermacoce remota East Maui, Haleakala National Park in Pipiwai exclosure below bridges Preserved_Specimen Welton, P. 2457 Maui BISH 2020-10-27
133 Spermacoce remota Manoa, Cooper Rd., Honolulu Preserved_Specimen Egler, F.E. 37422 Oahu BISH 1937-06-03
134 Spermacoce remota roadside, Hilo Preserved_Specimen Degener, O. 32432 Hawaii BISH 1966-05-20
135 Spermacoce remota Kauman substation Preserved_Specimen Wagner, W.L. 4632 Hawaii BISH 1982-11-16
136 Spermacoce remota 2062A St. Louis Dr., Honolulu Preserved_Specimen Wagner, W.L. Oahu BISH 1988-04-27
137 Spermacoce remota Exp.. Sta. Grounds Preserved_Specimen Caum, E.L. Oahu BISH 1935-01-21
138 Spermacoce remota Honolulu Preserved_Specimen Woolford, H.A. Oahu BISH 1950-03-31
139 Spermacoce remota Kauai Preserved_Specimen Forbes, C.N. Kauai BISH 1940-02-12
140 Spermacoce remota Visitor Center and main ruins, Puuhonua o Honaunau National Historical Park Preserved_Specimen Higashino, P.K. 10294 Hawaii BISH 1984-04-08
141 Spermacoce remota Manele and Hulupoe Bays In lawns. Preserved_Specimen Oppenheimer, H.L. 40116 Lanai BISH 2001-04-15
142 Spermacoce remota Honolulu, Punahou area, 1940 Manoa Rd A weed in lawn and garden growing in full sun or partial shade. Preserved_Specimen The flowers are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
143 Spermacoce remota Manoa Campus, Univ. Hawaii, Honolulu Common roadside weed. Preserved_Specimen Flws: white-pink. Swarbrick, J.T. 7 Oahu BISH 1976-01-10
144 Spermacoce remota W Maui, Lahaina District, Napili Growing in unnamed gulch used as dump site. Preserved_Specimen Oppenheimer, H.L. 20128 Maui BISH 2001-02-21
145 Spermacoce remota Makawao District, disturbed roadside on Hwy side of Kanaha Pond (Haleakala Hwy) Dominated by grasses, Pluchea indica and P. symphytifolia. Preserved_Specimen Stems prostrate (perhaps due to continual mowing); corolla white, purple-tinged toward apex. Wagner, W.L. 5796 Maui BISH 1988-03-03
146 Spermacoce remota Mauna Kea (Puu Laau Road) In open mamane-naio forest. Preserved_Specimen Engilis Jr., A. 18 Hawaii BISH 1987-08-19
147 Spermacoce remota Punahou area, Honolulu. 1940 Manoa Rd In lawn and garden growing in full sun or partial shade. Preserved_Specimen A weed. The flws are white with a rose pink tinge. Lau, J. 1475 Oahu BISH 1985-09-30
148 Spermacoce remota Puna District, Keaau roadside Preserved_Specimen weed with white flowers Herbst, D.R. 5531 Hawaii BISH 1975-10-22
149 Spermacoce remota East Maui, Makawao District, Honokala, vicinity of Hana Hwy In waste area. Preserved_Specimen 1m tall herbs forming tangles; corollas white tinged with pink. Oppenheimer, H.L. 70203 Maui BISH 2002-07-14
150 Spermacoce remota U.S. Experimental Tree Planting site (Forest Reserve) within Halepuaa Forest Reserve in teh Puna District Common on the roadsides, trailsides, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
151 Spermacoce remota Along road at Haena State Park Area along road. Preserved_Specimen Weed. Corn, C. Kauai BISH 1978-06-15
152 Spermacoce remota Koloa Dist., Kalaheo, Kalawai Park Secondary vegetation of Syzygium cumini, Psidium guajava, and Ficus microcarpa. Common weed in open areas. Preserved_Specimen Perennial herb with suberect, purplish stems. Corolla pink. Lorence, D.H. 5210 Kauai BISH 1987-05-10
153 Spermacoce remota Kahuku, Honolulu Slope: 2. Soil: sandy. Vegetation type: Coastal plants. Dist: Dry and wet regions. Preserved_Specimen Slightly hairy plants spreading branches. Guinto, J. 32 Oahu BISH 1986-02-04
154 Spermacoce remota Kawaihau Dist., Nonou Forest Reserve, crest of Sleeping Giant Mountain Secondary forest of guava, Java plum, and Schinus, with forestry plantings and rare native remnants. Locally abundant in grassy clearing; population uniform. Naturalized. Preserved_Specimen Low, perennial herb, stems erect-spreading, corolla white. Lorence, D.H. 5550 Kauai BISH 1987-09-13
155 Spermacoce remota 2525 Nuuanu Ave Weed between coral pavement blocks, moist places. Preserved_Specimen Weed. Judd, C.S. Oahu BISH 1936-12-05
156 Spermacoce remota East Maui, Hana District, Paakea, along Hana Hwy Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 60318 Maui BISH 2003-06-22
157 Spermacoce remota Kahuku Training Area, from main Delta road, took left makai towards ICA-21, on ridge between Keaaulu and Lamaloa Gulches, a bit less than halfway to the big open red dirt clearing. Associated canopy: Leucaena leucocephala, Psidium cattleyanum, Syzygium cumini, Chrysophyllum oliviforme. Understory: Clidemia hirta, Dicranopteris linearis, Mimosa pudica, Paspalum conjugatum, Acacia confusa. Area shaded and damp. Preserved_Specimen Naturalized herb in lush patch ca 0.5 m tall, several meters wide. Upright but delicate and sprawling. Leaves opposite, up to 10 cm long. Flowers small, white, hairs on petals, buds prickly. Immature fruit. Inflorescence with faint pleasant scent. Thick patch represents either one massive clone or hundreds of individuals. Differs from Spermacoce assurgens in longer leaves and overall gestalt; growth in a dense patch also seemed unusual. Beachy, J. 512 Oahu BISH 2019-03-25
158 Spermacoce remota In tall grass in the Wainiha Hui lands, Haena, Kauai Wet depression. Spermacoce is common in the area as are California grass, Paspalum, Amaranthus, and tropical almond. Collection site is weedy. Preserved_Specimen Plants are very rangy with weak stems. Funk, E.J. 801 Kauai BISH 1990-05-03
159 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
160 Spermacoce remota University of Hawaii Campus, Maile Way, Honolulu Roadside. Preserved_Specimen Flowers white to pinkish. Fosberg, F.R. 10528 Oahu BISH 1933-04-09
161 Spermacoce remota Hana District; Hana, along Uwala road near Hana airport Road side weed Preserved_Specimen Alien veg. Weed, 1 foot ht with flowers and fruit Perlman, S. 22924 Maui BISH 2012-06-20
162 Spermacoce remota Lihue Dist., Sugar cane road E of Omoe Ruderal. Preserved_Specimen Herbs with erect branches to 40cm, flws: white with reddish tips of corolla lobes, seeds dark brown. Kiehn, M. 90091722 Kauai BISH 1990-09-17
163 Spermacoce remota Kalaupapa; Bayview, Quonset lawn, weedy Preserved_Specimen Wysong, M.L. 786 Molokai BISH 2005-08-12
164 Spermacoce remota East Maui, Hana District, Kipahulu Growing on side of rd. Preserved_Specimen 1m tall shrubs; flowers white w. pink tint. Oppenheimer, H.L. 60115 Maui BISH 2001-06-11
165 Spermacoce remota Kaunakakai, near the wharf In lawn. Preserved_Specimen Oppenheimer, H.L. 70111 Molokai BISH 2001-07-14
166 Spermacoce remota Limahuli Valley (Botanical Garden), Napali Coast Common along paths and around buildings. Preserved_Specimen Branched, stems ascending, lvs dark purplish green, flws: pinkish white, corollas well exserted. Fosberg, F.R. 57443 Kauai BISH 1978-03-31
167 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch Growing in lawn. Preserved_Specimen Oppenheimer, H.L. 110109 Maui BISH 2001-11-08
168 Spermacoce remota Herbarium of the H.L. Lyon Arboretum. On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
169 Spermacoce remota Woodlawn, Manoa, Honolulu Roadside weed patch. Preserved_Specimen Flowers pinkish white. Fosberg, F.R. 14111 Oahu BISH 1937-06-19
170 Spermacoce remota C.M.Cooke, Jr. Ranch, Maunawili, Koolau Mts., Kailua Near dairy buildings Preserved_Specimen Weed. Flowers white Fosberg, F.R. 10772 Oahu BISH 1935-01-23
171 Spermacoce remota East Maui, Hana District, Coastal pasture lands NW of Ka Iwi O Pele in Casuarina windbreak With Plantago, Digitaria and Oxalis. Preserved_Specimen Flws: white. Flynn, T.W. 2588 Maui BISH 1987-10-30
172 Spermacoce remota Hualalai (Olma Tract, Kaloka Mauka) Ohia-fern mesic forest Preserved_Specimen Engilis Jr., A. 31 Hawaii BISH 1987-08-19
173 Spermacoce remota On bank behind Cottage A Fairly open, well drained. Preserved_Specimen Baker, R.F. 92 Oahu BISH 1975-04-26
174 Spermacoce remota Woodlawn, Manoa Valley Naturalized roadside weed. Preserved_Specimen Degener, O. 27707 Oahu BISH 1929-01-13
175 Spermacoce remota P.T.B.G., Lawai Valley Common. Preserved_Specimen Herb with dark green foliage, white flws. Herbst, D.R. 2276 Kauai BISH 1971-12-30
176 Spermacoce remota U.S. Experimental Tree Planting Site ( Forest Reserve) within Halepuaa Forest Reserve in the Puna District On the roadside, trailside, and in disturbed areas. Preserved_Specimen Clarke, G. 79.223 Hawaii BISH 1979-03-08
177 Spermacoce remota Kualapuu, near road to gulf course Roadside. Preserved_Specimen Weed. Oppenheimer, H.L. 110127 Molokai BISH 2001-11-18
178 Spermacoce remota Tantalus Dr. ca 200m above the last house of the development Common plant on roadside Preserved_Specimen Common Faccenda, K. 1878 Oahu BISH 2021-05-22
179 Spermacoce remota W Maui, Wailuku District, Kahului In lawns and waste areas. Preserved_Specimen Weed. Oppenheimer, H.L. 40136 Maui BISH 2001-04-23
180 Spermacoce remota Kaimi Farm, Kailua Koolaupoko Growing in moist road back. Preserved_Specimen 1-1 1/2' tall, flws white. Hosaka, E.Y. 2438 Oahu BISH 1939-02-05
181 Spermacoce remota ahupuaa of Halepuaa in the Puna District The plant is common growing on the side of the road. The area is exposed to moderate sunlight. Preserved_Specimen Kubo, R. 5 Hawaii BISH 1979-03-07
182 Spermacoce remota Kahuku. roadside Slope: 1%. Soil: Sandy loam. Vegetation type: tropical. Preserved_Specimen Flws: white, 1/12 across. Plant can be downy or smooth. Smith, M. 30 Oahu BISH 1986-02-04
183 Spermacoce remota East Maui, Halehaku, vicinity of Papalua Gulch along Hana Hwy Roadside. Preserved_Specimen 1m tall weed. Oppenheimer, H.L. 110118 Maui BISH 2001-11-12
184 Spermacoce remota 3rd, Mountain View, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/21/2007
185 Spermacoce remota W Maui, Lahaina District, Alaeloa, N of Kaopala Gulch OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
186 Spermacoce remota Hilo Airport Preserved_Specimen Stemmermann, R.L. 7129 Hawaii BISH 1986-11-11
187 Spermacoce remota Kipahulu Preserved_Specimen Dnse patches sprawling over grass in rich, wet, lowland pastures. Hobdy, R.W. 3881 Maui BISH 1995-09-07
188 Spermacoce remota Kahana Valley State Park Preserved_Specimen Herb 0.2-0.3m tall; flower petals pink. Annable, C.R. 3105 Oahu BISH 1996-11-04
189 Spermacoce remota Hilo District, Coconut Island Preserved_Specimen flowers white, ripe fruit only on old (already dry) plants Kiehn, M. 89072711 Hawaii BISH 1989-07-27
190 Spermacoce remota East Maui, Kipahulu Preserved_Specimen Welton, P. 1672 Maui BISH 1992-08-27
191 Spermacoce remota W Maui, Lahaina District, Napili OBSERVATION Curtis Daehler Maui HAPI 3/19/2007
192 Spermacoce remota Hilo, parking lot at Tropical Rent-a-Car, disturbed area Preserved_Specimen Wagner, W.L. 5684 Hawaii BISH 1986-06-12
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