Description

Key Characters:

Growth Form: Aromatic perennial herbs.

Stems: Stems erect or ascending, quadrangular, densely spreading long–pilose.

Roots:

Leaves: Leaves simple. Opposite. Blades ovate–cordate to narrowly ovate, (13–)16–24 cm long, (4–)6.5–10.5 cm wide. Upper surfaces moderately pilose; lower surfaces very densely pilose to subtomentose. Margins coarsely crenate or crenate–serrate. Petioles (5.5–)8–10.5 cm long, densely spreading long–pilose. Stipules absent.

Flowers: Flowers subsessile, in lax racemes 20–25 cm long, sparsely pilose and densely glandular puberulent, 2 flowers per verticillaster, bracts lanceolate, 4–7 mm long, lower ones foliaceous, up to 50 mm long. Calyx actinomorphic, narrowly funnelform, pendent, (11–)15–18 mm long, sparsely pilose and densely glandular puberulent, 5-toothed, the teeth deltate, 2.5–3.5 mm long, usually persistent or sometimes only ventral 1/2 persistent. Corolla zygomorphic, bilabiate, white or sometimes tinged purple externally, tube slightly curved, very narrowly funnelform, 20–28 mm long, sparsely pilose, upper lip 7–11 mm long, lower lip 11–20 mm long; nectary disk often present at base of ovary, annular or developed on anterior side only. Stamens 4, both pairs of similar length or the upper pair slightly longer, ascending under the upper lip, included; filaments inserted near the throat; anthers dithecal; anther sacs divaricate, opening by longitudinal slits. Ovary superior, 2-carpellate, placentation basal-axile; ovules 1 per ovary lobe; style 1, included, shortly 2-lobed, the lobes clavate, pubescent in upper part; stigmatic only at the truncate apex, 1 lobe often rudimentary; stigmas 2.

Fruit: Nutlets 1-seeded; dark brown; obovate; ca. 6–7 mm long; 2 abortive; inner surface near apex coarsely rugose and slightly glandular. Seeds with endosperm absent or scanty and oily.

Ploidy:

Habitat: Formerly occurring on the west end of Lānaʻi (Ahupua'a Ka'ā and Paoma'i).

Elevation Range: