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Cyperus phleoides var. phleoides

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Cyperaceae Genus: Cyperus

Description

Key Characters:

Growth Form: Perennials with short, woody rhizomes.

Stems: culms 10–50 cm tall, (much shorter than leaves) 2–5 mm in diameter near base, base thickly clothed with leaf sheaths 3–20 cm long.

Roots:

Leaves: Leaves simple. Leaves numerous, crowded. Leaves numerous, crowded. Blades linear shorter than or longer than the culm, 3–8 mm wide. Blades coriaceous. Margins recurved. Veins parallel. Sheathing; sheaths tinged brown, pale brown, or reddish brown, eventually splitting into parallel fibers. Stipules absent.

Flowers: Flowers in inflorescences simple or partially compound, umbelliform or contracted and head-like, 1.5–4 cm long, 3– 8 cm wide, rays 3–11, 0.3–3 cm long, rarely further branched, spikes 5–12, ovoid or ellipsoid-cylindrical, 1–2 cm long, 0.8–1 cm in diameter, simple, rarely developing a smaller lateral spike at base, densely bearing numerous spikelets, the bracteoles hispid–scabrous on midnerve; involucral bracts 5–12, divergent to reflexed, 2–25 cm long, much longer than the inflorescence; spikelets bearing 2–3 glumes and 1–2 fertile flowers, erect to spreading, often curved upward, oblong-lanceolate or linear-lanceolate, moderately flattened, 3.5–4 mm long, 1–1.5 mm wide, apex acute; glumes herbaceous with rather broad hyaline margins, lanceolate to oblong-lanceolate, 3–4.5 mm long, 1–1.3 mm wide, strongly folded, 7–nerved or 9–nerved, the midnerve prominently to weakly spinulose–scabrous. Flowers bisexual (perfect). Corolla (petals) absent. Ovary superior.

Fruit: Achenes linear–oblong; trigonous; 1.5–2.2 mm long; ca. 0.4 mm wide. Seeds 1; seed coat thin; free from pericarp; embryo basal; endosperm abundant.

Ploidy:

Habitat: Occurring on shallow soil on coastal cliffs or among rocks on beaches.

Elevation Range: 0–10 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Kaho'olawe Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Other References

Wagner et al. 1990:1423 (K, Mo, L, M, H [as Mariscus phleoides subsp. phleoides]); Wagner & Herbst 1995:18 (Ka); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M. phleoides subsp. phleoides)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Cyperus phleoides var. phleoides 'Ale'ale stack, near Puu Koae. Elev. 10-90 m. Mixed native coastal shrubland. PRESERVED_SPECIMEN K. R. Wood 1726 Kahoolawe PTBG 3/18/1992
2 Cyperus phleoides var. phleoides 'Ale'ale stack, near Puu Koae. Elev. 10-90 m. Mixed Native Coastal Shrubland. PRESERVED_SPECIMEN K. R. Wood 1723 Kahoolawe PTBG 3/18/1992
3 Cyperus phleoides var. phleoides W. Maui; Hale Pohaku. Metrosideros-Dicranopteris Montane Wet Ridge. PRESERVED_SPECIMEN K. R. Wood 1612 Maui PTBG 2/4/1992
4 Cyperus phleoides var. phleoides Kahakuloa, Puu Kahulianapa, coastal sea cliffs and slopes. Coastal dry. PRESERVED_SPECIMEN K. R. Wood 7182 Maui PTBG 2/11/1998
5 Cyperus phleoides var. phleoides Waianae Mts. ,Lualualei, slopes of Puu Kaua,1840 ft. el., on cliffs. In Dodonaea shrubland. PRESERVED_SPECIMEN Steve Perlman 14060 Oahu PTBG 3/1/1994
6 Cyperus phleoides var. phleoides Wailau Valley; seacliffs of Wailau; Elev. 400-600 ft. PRESERVED_SPECIMEN K. R. Wood 3823 Molokai PTBG 12/8/1994
7 Cyperus phleoides var. phleoides Kalalau Rim; Kalahu side below and west of first Kalalau lookout; 300 m east of plane crash. Diverse Montane Mesic Precipitous Slopes with small pockets of forest and Vertical Cliffs. PRESERVED_SPECIMEN K. R. Wood 1398 Kauai PTBG 5/9/1991
8 Cyperus phleoides var. phleoides Offshore islet east of Kalaupapa Peninsula. Chamaesyce celastroides/Chenopodium oahuense coastal dry shrubland with Diospyros forest. PRESERVED_SPECIMEN K. R. Wood 8322 Mokapu Islet PTBG 3/8/2000
9 Cyperus phleoides var. phleoides Pohakuao; Hanging valley between Kalalau and Hanakoa; Collected on SW wall PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Lau, J. 1765 Kauai US 4/1/1992
10 Cyperus phleoides var. phleoides Hana, Ka'uiki head, on SE side of Hana Bay. Elev. 80-386 ft. Secondary forest. PRESERVED_SPECIMEN Tim Flynn 2412 Maui PTBG 10/10/1987
11 Cyperus phleoides var. phleoides Kalaupapa Trail, on basalt cliffs. PRESERVED_SPECIMEN Harold St. John 25239 Molokai PTBG 12/1/2025
12 Cyperus phleoides var. phleoides West Maui, Hale Pohaku. PRESERVED_SPECIMEN K. R. Wood 2078 Maui PTBG 9/2/1992
13 Cyperus phleoides var. phleoides Kalalau; west of first lookout; Kalahu area. Diverse mesic forest & cliffs. PRESERVED_SPECIMEN K. R. Wood 1596 Kauai PTBG 1/28/1992
14 Cyperus phleoides var. phleoides Na Pali Coast, Hono o Na Pali N.A.R.,Waiahuakua valley, back of valley by main waterfall, on cliffs west of main falls. Metrosideros lowland mesic cliff. PRESERVED_SPECIMEN Steve Perlman 13272 Kauai PTBG 1/1/2024
15 Cyperus phleoides var. phleoides makawao dist; 'ale'ale stack; near pu'u koa'e; 50 deg ne aspect. PRESERVED_SPECIMEN k. r. wood, r. hobdy & s. p. perlman 2152 Kahoolawe US 11/24/1992
16 Cyperus phleoides var. phleoides South point area, Kamilo, Kaalualu bay in beach strand. Shrubland PRESERVED_SPECIMEN Perlman, S. P.; Perry, L.; Bio, K. 21360 Hawaii US 1/20/2009
17 Cyperus phleoides var. phleoides Near main road, Kaaukuu Gulch. PRESERVED_SPECIMEN Degener, O. 22206 Molokai US 3/16/1952
18 Cyperus phleoides var. phleoides Olowalu Valley, W Maui Edge of stream PRESERVED_SPECIMEN Forbes, C. N. 2263.M Maui US 5/8/2020
19 Cyperus phleoides var. phleoides kalaupapa trail, on basalt cliffs. PRESERVED_SPECIMEN h. st. john 25239 Molokai US 12/25/1953
20 Cyperus phleoides var. phleoides w. maui, hulu, offshore islet south of pu`u koae, kahakuloa, west side of central ridge. Coastal vegetation. PRESERVED_SPECIMEN k. r. wood, f. starr & k. starr 11185 Maui US 4/4/2005
21 Cyperus phleoides var. phleoides W. Maui, Hulu, offshore islet south of Pu'u Koae, Kahakuloa, west side of central ridge. Coastal vegetation. PRESERVED_SPECIMEN Wood, K. R.; Starr, F.; Starr, K. 11185 Maui US 4/4/2005
22 Cyperus phleoides var. phleoides olowalu valley, w maui PRESERVED_SPECIMEN c. n. forbes 2263.m Oahu US 5/8/1920
23 Cyperus phleoides var. phleoides Makawao Dist; 'Ale'ale stack; near Pu'u Koa'e; 50 deg NE aspect. PRESERVED_SPECIMEN Wood, K. R.; Hobdy, R.; Perlman, S. P. 2152 Kahoolawe US 11/24/1992
24 Cyperus phleoides var. phleoides Keopukaloa PRESERVED_SPECIMEN St. John, H.; Baker, R. J.; Coulter, J. W.; Fosberg, F. R.; Yuncker, T. G. 12739 Molokai US 12/27/1932
25 Cyperus phleoides var. phleoides keopukaloa PRESERVED_SPECIMEN h. st. john, r. j. baker, j. w. coulter, f. r. fosberg & t. g. yuncker 12739 Molokai US 12/27/1932
26 Cyperus phleoides var. phleoides Kalalau; W of first lookout; Kalahu area. PRESERVED_SPECIMEN Wood, K. R. 1596 Kauai US 1/28/1992
27 Cyperus phleoides var. phleoides south point area, kamilo, kaalualu bay in beach strand. Shrubland PRESERVED_SPECIMEN s. p. perlman, l. perry & k. bio 21360 Hawaii US 1/20/2009
28 Cyperus phleoides var. phleoides west maui, kahakuloa PRESERVED_SPECIMEN o. degener 21959 Maui US 2/28/1952
29 Cyperus phleoides var. phleoides Kalaupapa Trail, on basalt cliffs. PRESERVED_SPECIMEN St. John, H. 25239 Molokai US 12/25/1953
30 Cyperus phleoides var. phleoides kalalau; w of first lookout; kalahu area. PRESERVED_SPECIMEN k. r. wood 1596 Kauai US 1/28/1992
31 Cyperus phleoides var. phleoides maunalei PRESERVED_SPECIMEN munro, g.c. collector number: 515 Lanai BISH 2/11/1930
32 Cyperus phleoides var. phleoides pohakuao; hanging valley between kalalau and hanakoa; collected on sw wall PRESERVED_SPECIMEN k. r. wood, s. p. perlman & j. lau 1765 Kauai US 4/1/1992
33 Cyperus phleoides var. phleoides West Maui, Kahakuloa PRESERVED_SPECIMEN Degener, O. 21959 Maui US 2/28/1952
34 Cyperus phleoides var. phleoides E Maui District, Alau Island, easternmost coast of E Maui PRESERVED_SPECIMEN Hobdy, R.W. 1304 Maui BISH 3/7/1982
35 Cyperus phleoides var. phleoides Near main road, Kaaukuu [sp?] Gulch, West Molokai Dry grassy slope far from ocean. PRESERVED_SPECIMEN Degener, O. 22206 Molokai BISH 3/16/1952
36 Cyperus phleoides var. phleoides near main road, kaaukuu gulch. PRESERVED_SPECIMEN o. degener 22206 Molokai US 3/16/1952
37 Cyperus phleoides var. phleoides Kalaupapa Trail, on basalt cliffs Tufts on grassy ledges. PRESERVED_SPECIMEN St. John, H. 25239 Molokai BISH 12/25/1953
38 Cyperus phleoides var. phleoides West of Kauhola Light, Kohala On earth within few feet of edge of ocean cliffs PRESERVED_SPECIMEN Degener, O. 21925 Hawaii BISH 2/23/1952
39 Cyperus phleoides var. phleoides kalaupapa pali trail, near top PRESERVED_SPECIMEN munro, g.c. collector number: s.n. Molokai BISH 7/25/1922
40 Cyperus phleoides var. phleoides Kailikaula Cliffs and Stream; Elev. 750-900 ft. Metrosideros-Osteomeles-Chamaesyce Lowland Mesic Forest and cliff w/ riparian communities. PRESERVED_SPECIMEN K. R. Wood 4868 Hawaii PTBG 12/14/1995
41 Cyperus phleoides var. phleoides 'Ale'ale stack, near Puu Koae. Elev. 10-90 m. Mixed Native Coastal Shrubland. PRESERVED_SPECIMEN K. R. Wood 1731 Kahoolawe PTBG 3/18/1992
42 Cyperus phleoides var. phleoides kalaupapa trail, on basalt cliffs Tufts on grassy ledges. PRESERVED_SPECIMEN st.john, h. collector number: 25239 Molokai BISH 12/25/1953
43 Cyperus phleoides var. phleoides Aukuu, slopes of PRESERVED_SPECIMEN Munro, G.C. 403 Lanai BISH 2/22/1915
44 Cyperus phleoides var. phleoides Pohakuao; hanging valley between Kalalau and Hanakoa; Elev. 400-600 m. Metrosideros-Diospyros lowland mesic forest. PRESERVED_SPECIMEN K. R. Wood 1765 Kauai PTBG 4/1/1992
45 Cyperus phleoides var. phleoides maliku, near paia Grassy ledges facing ocean. PRESERVED_SPECIMEN degener, o. collector number: 21947 Maui BISH 2/26/1952
46 Cyperus phleoides var. phleoides west of kauhola light, kohala Gulch PRESERVED_SPECIMEN degener, o. collector number: 21925 Oahu BISH 2/23/1952
47 Cyperus phleoides var. phleoides maunalei PRESERVED_SPECIMEN munro, g.c. collector number: 827b Lanai BISH 2/11/1930
48 Cyperus phleoides var. phleoides near main road, kaaukuu gulch, west molokai PRESERVED_SPECIMEN degener, o. collector number: 22206 Oahu BISH 3/16/1952
49 Cyperus phleoides var. phleoides Alau, Maui on north wall and sheer bowl facing east and east ridge of large hill PRESERVED_SPECIMEN Starr, F. 050405-26 Maui BISH 4/5/2005
50 Cyperus phleoides var. phleoides Paokalani islet, 150 m offshore of Honokea Stream, 270x85 m Lantana camara, Pluchea carolinensis, Osteomeles anthyllidifolia, Schiedea globosa, Chenopodium oahuense, Cocculus orbiculatus, Eragrostis variabilis, Plectranthus parviflorus, Dodonaea viscosa, Psydrax odorata, Chamaesyce celastroides PRESERVED_SPECIMEN Wood, K.R. 11216 Hawaii BISH 4/14/2005
51 Cyperus phleoides var. phleoides Aukuu PRESERVED_SPECIMEN Munro, G.C. 258 Lanai BISH 2/22/1915
52 Cyperus phleoides var. phleoides Wailau PRESERVED_SPECIMEN Rock, J.F.C. 7058 Molokai BISH 4/20/1910
53 Cyperus phleoides var. phleoides Nahiku Along the beach, between stones. PRESERVED_SPECIMEN Rock, J.F.C. 8755 Maui BISH 5/10/1911
54 Cyperus phleoides var. phleoides east slope of koko crater PRESERVED_SPECIMEN bergman, h.f. collector number: s.n. Oahu BISH 10/1/1927
55 Cyperus phleoides var. phleoides w maui, kahakuloa Windswept grassland near ocean. PRESERVED_SPECIMEN degener, o. collector number: 21959 Maui BISH 2/28/1952
56 Cyperus phleoides var. phleoides haleakala, lower ditch trail PRESERVED_SPECIMEN skottsberg, c.j.f. collector number: 798 Kauai BISH 10/14/1922
57 Cyperus phleoides var. phleoides keopukaloa Tufts on cliff in gulch. PRESERVED_SPECIMEN st.john, h. baker, r.j.; coulter, j.w.; fosberg, f.r.; yuncker, t.g. collector number: 12739 Molokai BISH 12/27/1932
58 Cyperus phleoides var. phleoides hamakua distr., kailikaula cliffs and stream; on open basalt walls PRESERVED_SPECIMEN wood, k.r. wood, p.; perlman, s. collector number: 4868 Maui BISH 12/14/1995
59 Cyperus phleoides var. phleoides By trailhead of Upper Wailuku River Trail PRESERVED_SPECIMEN Adee, K.T. s.n. Hawaii BISH 6/21/1979
60 Cyperus phleoides var. phleoides Near Sisal PRESERVED_SPECIMEN Forbes, C.N. 1757.O Oahu BISH 2/16/1912
61 Cyperus phleoides var. phleoides W Maui, Kahakuloa, Hulu Island PRESERVED_SPECIMEN Kepler, A.K. 45 Maui BISH 8/20/1981
62 Cyperus phleoides var. phleoides E Maui District, Keopuka Rock, off the N coast of E Maui PRESERVED_SPECIMEN Hobdy, R.W. 1399 Maui BISH 5/16/1982
63 Cyperus phleoides var. phleoides aukuu PRESERVED_SPECIMEN munro, g.c. collector number: 258 Lanai BISH 2/22/1915
64 Cyperus phleoides var. phleoides wailau PRESERVED_SPECIMEN rock, j.f.c. collector number: 7058 Molokai BISH 4/20/1910
65 Cyperus phleoides var. phleoides hoolehua Dry cliffs overlooking ocean. PRESERVED_SPECIMEN degener, o. collector number: 8433 Molokai BISH 5/4/1928
66 Cyperus phleoides var. phleoides maliku, on the shore PRESERVED_SPECIMEN rock, j.f.c. collector number: 8755 Kauai BISH 5/1/1911
67 Cyperus phleoides var. phleoides east maui, hana district, hana, kauiki head, on se side of hana bay Secondary forest dominated by Casuarina, Terminalia, Syzygium, Ardisia, and Lantana. On open, steep cinder faces on northeast slope. With Scaevola, Ipomoea, Bidens, Morinda, and Vigna. PRESERVED_SPECIMEN flynn, t. lewis, c. collector number: 2412 Maui BISH 10/10/1987
68 Cyperus phleoides var. phleoides West Maui, Wailuku District, Kahakuloa, between Puu Koae and Puu Kahulianapa. Associated vegetation: Scaevola Coastal Shrubland with Lycium, Sesuvium, Fimbristylis, Cyperus, Chenopodium, Schiedea, Osteomeles, Sida, Lipochaeta, Panicum, Eragrostis, Schenkia, Lysimachia, Pseudognaphalium, Schinus, Digitaria, Lantana, Anagallis, Desdomium. PRESERVED_SPECIMEN Oppenheimer, H.L. H41509 Maui BISH 4/2/2015
69 Cyperus phleoides var. phleoides Niulii, Kohala Dry exposed coast. PRESERVED_SPECIMEN Degener, O. 8461 Hawaii BISH 8/6/1928
70 Cyperus phleoides var. phleoides Maliku, near Paia Grassy ledges facing ocean. PRESERVED_SPECIMEN Degener, O. 21947 Maui BISH 2/26/1952
71 Cyperus phleoides var. phleoides windward side of island, anapuhi to wainene, long coastal boulder-strewn beach with vertical cliff at back 10-30 m from water Cliff with Artemisia australis, Chenopodium oahuense, Schiedea globosa, Kalanchoe pinnata, Tetramolopium sylvae, Lythrum maritimum, and Hedyotis littoralis, among others. Back of rocky area near hala stand. PRESERVED_SPECIMEN wagner, w.l. lorence, d. collector number: 5742 Molokai BISH 8/1/1987
72 Cyperus phleoides var. phleoides W Maui, Kahakuloa Windswept grassland near ocean. PRESERVED_SPECIMEN Degener, O. 21959 Maui BISH 2/28/1952
73 Cyperus phleoides var. phleoides East slope of Koko Crater PRESERVED_SPECIMEN Bergman, H.F. s.n. Oahu BISH 10/1/1927
74 Cyperus phleoides var. phleoides Windward side PRESERVED_SPECIMEN Rock, J.F.C. 6166 Molokai BISH 4/1/1910
75 Cyperus phleoides var. phleoides pauwela Sandy soil. PRESERVED_SPECIMEN krauss, n.l.h. collector number: 1104 Oahu BISH 4/4/1967
76 Cyperus phleoides var. phleoides w maui, honokohau On side of cliff. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1779 Maui BISH 2/10/1937
77 Cyperus phleoides var. phleoides niulii, kohala PRESERVED_SPECIMEN degener, o. collector number: 8461 Hawaii BISH 8/6/1928
78 Cyperus phleoides var. phleoides Maunalei PRESERVED_SPECIMEN Munro, G.C. 827b Lanai BISH 2/11/1930
79 Cyperus phleoides var. phleoides Keopukaloa Tufts on cliff in gulch. PRESERVED_SPECIMEN St. John, H. 12739 Molokai BISH 12/27/1932
80 Cyperus phleoides var. phleoides paokalani islet, 150 m offshore of honokea stream, 270x85 m Lantana camara, Pluchea carolinensis, Osteomeles anthyllidifolia, Schiedea globosa, Chenopodium oahuense, Cocculus orbiculatus, Eragrostis variabilis, Plectranthus parviflorus, Dodonaea viscosa, Psydrax odorata, Chamaesyce celastroides PRESERVED_SPECIMEN wood, k.r. polhemus, j. collector number: 11216 Hawaii BISH 4/14/2005
81 Cyperus phleoides var. phleoides puu koae, kahakuloa, exposed grassy windward slope PRESERVED_SPECIMEN st.john, h. collector number: 24719 Maui BISH 12/23/1951
82 Cyperus phleoides var. phleoides Camp 10 Road, just past Waineke Swamp PRESERVED_SPECIMEN Wagner, W.L. 6044 Kauai BISH 4/9/1988
83 Cyperus phleoides var. phleoides Hamakua Distr., Kailikaula Cliffs and Stream; on open basalt walls Metrosideros-Osteomeles-Chamaesyce Lowland Mesic Forest and cliff with riparian communities. Associated species: Carex spp., Dianella, Canthium, Eragrostis, Diospyros, Dodonaea, Alyxia. Threats: Psidium spp., Oplismenus, Coix, Rubus rosifolius, Schinus, Lantana. PRESERVED_SPECIMEN Wood, K.R. 4868 Hawaii BISH 12/14/1995
84 Cyperus phleoides var. phleoides west of kauhola light, kohala On earth within few feet of edge of ocean cliffs. PRESERVED_SPECIMEN degener, o. collector number: 21925 Hawaii BISH 2/23/1952
85 Cyperus phleoides var. phleoides aukuu, slopes of PRESERVED_SPECIMEN munro, g.c. collector number: 403 Lanai BISH 2/22/1915
86 Cyperus phleoides var. phleoides near sisal PRESERVED_SPECIMEN forbes, c.n. cooke, c.m. collector number: 1757.o Oahu BISH 2/16/1912
87 Cyperus phleoides var. phleoides Hanalei Distr., Pilaa, Kepuhi Point Growing in seepage area on steep, shaded slope back of beach. PRESERVED_SPECIMEN Herbst, D.R. 2400 Kauai BISH 4/27/1972
88 Cyperus phleoides var. phleoides Pauwela PRESERVED_SPECIMEN Krauss, N.L.H. 1104 Maui BISH 4/4/1967
89 Cyperus phleoides var. phleoides by trailhead of upper wailuku river trail PRESERVED_SPECIMEN adee, k.t. collector number: s.n. Hawaii BISH 6/21/1979
90 Cyperus phleoides var. phleoides e maui district, alau island, easternmost coast of e maui PRESERVED_SPECIMEN hobdy, r.w. collector number: 1304 Maui BISH 3/7/1982
91 Cyperus phleoides var. phleoides Maunalei PRESERVED_SPECIMEN Munro, G.C. 515 Lanai BISH 2/11/1930
92 Cyperus phleoides var. phleoides makawao district, aleale stack, near puu koae Mixed Native Coastal Shrubland with Portulaca molokiniensis, P. villosa, Senna gaudichaudii, Chamaesyce celastroides var. amplectens, Chamaesyce sp., Capparis, Lipochaeta lavarum, Bidens mauiensis, Sida fallax, Doryopteris decipiens, Panicum torridum, P. fauriei var. latius, Eragrostis atropioides, Ipomoea tuboides; indications of Bulwers Petrel and Wedgetail Shearwater nesting sites. Weeds: Waltheria, Emilia fosbergii, Chloris barbata, Nicotiana glauca. PRESERVED_SPECIMEN wood, k.r. perlman, s.; lau, j.; rowland, c. collector number: 1726 Kahoolawe BISH 3/18/1992
93 Cyperus phleoides var. phleoides mui zoological & botanical gardens. originally from pauwela point PRESERVED_SPECIMEN davis, j. sylva, r. collector number: 36 Maui BISH 7/3/1978
94 Cyperus phleoides var. phleoides Kalaupapa; Papapaiki, E coastal side, fence PRESERVED_SPECIMEN Wysong, M.L. 578 Molokai BISH 1/12/2005
95 Cyperus phleoides var. phleoides growing between pohaku malumalu and kepuhi point In wet pocket by seepage area. PRESERVED_SPECIMEN herbst, d.r. collector number: 2400 Kauai BISH 4/27/1972
96 Cyperus phleoides var. phleoides wailau valley PRESERVED_SPECIMEN forbes, c.n. collector number: 525.mo Molokai BISH 9/1/1912
97 Cyperus phleoides var. phleoides Hoolehua Dry cliffs overlooking ocean. PRESERVED_SPECIMEN Degener, O. 8433 Molokai BISH 5/4/1928
98 Cyperus phleoides var. phleoides w maui district, coastal cliffs by puu koae PRESERVED_SPECIMEN hobdy, r.w. collector number: 385 Maui BISH 10/30/1978
99 Cyperus phleoides var. phleoides Wailau Valley PRESERVED_SPECIMEN Forbes, C.N. 525.Mo Molokai BISH 9/1/1912
100 Cyperus phleoides var. phleoides Kaaukuu Gulch, West Molokai, near main road Dry grassy slope far from ocean. PRESERVED_SPECIMEN Degener, O. 22206 Molokai BISH 3/16/1952
101 Cyperus phleoides var. phleoides camp 10 road, just past waineke swamp PRESERVED_SPECIMEN wagner, w.l. takeuchi, w.; imada, c. collector number: 6044 Kauai BISH 4/9/1988
102 Cyperus phleoides var. phleoides windward side PRESERVED_SPECIMEN rock, j.f.c. collector number: 6166 Molokai BISH 4/1/1910
103 Cyperus phleoides var. phleoides Kalaupapa pali trail, near top PRESERVED_SPECIMEN Munro, G.C. s.n. Molokai BISH 7/25/1922
104 Cyperus phleoides var. phleoides windward side On denuded slope. PRESERVED_SPECIMEN rock, j.f.c. collector number: 6166 Oahu BISH 4/1/1910
105 Cyperus phleoides var. phleoides olowalu valley, edge of stream PRESERVED_SPECIMEN forbes, c.n. collector number: 2263.m Midway Atoll BISH 5/8/1920
106 Cyperus phleoides var. phleoides Makawao District, Aleale stack; near Puu Koae 50 deg NE aspect; mixed native coastal shrubland w/Senna gaudichaudii, Lipochaeta lavarum, Waltheria, Chamaesyce celastroides var. amplectens, Bidens mauiensis, Panicum torridum, Eragrostis cf. atropioides; Occuring in shallow soil amidst talus; upper section of stack. PRESERVED_SPECIMEN Wood, K.R. 2152 Kahoolawe BISH 11/24/1992
107 Cyperus phleoides var. phleoides Kau Forest Reserve, Mt. House Trail Common along roadside. Wet w/ tree ferns, Metrosideros, Tibouchina PRESERVED_SPECIMEN Herbst, D.R. 203 Hawaii BISH 8/4/1966
108 Cyperus phleoides var. phleoides Puu Koae, Kahakuloa, exposed grassy windward slope PRESERVED_SPECIMEN St. John, H. 24719 Maui BISH 12/23/1951
109 Cyperus phleoides var. phleoides Growing between Pohaku Malumalu and Kepuhi Point In wet pocket by seepage area. PRESERVED_SPECIMEN Herbst, D.R. 2400 Kauai BISH 4/27/1972
110 Cyperus phleoides var. phleoides Maliku, on the shore PRESERVED_SPECIMEN Rock, J.F.C. 8755 Maui BISH 5/1/1911
111 Cyperus phleoides var. phleoides Olowalu Valley, edge of stream PRESERVED_SPECIMEN Forbes, C.N. 2263.M Maui BISH 5/8/1920
112 Cyperus phleoides var. phleoides West of Kauhola Light, Kohala On earth within few feet of edge of ocean cliffs. PRESERVED_SPECIMEN Degener, O. 21925 Hawaii BISH 2/23/1952
113 Cyperus phleoides var. phleoides w maui, kahakuloa, hulu island PRESERVED_SPECIMEN kepler, a.k. collector number: 45 Maui BISH 8/20/1981
114 Cyperus phleoides var. phleoides hanalei distr., pilaa, kepuhi point PRESERVED_SPECIMEN herbst, d.r. collector number: 2400 Kauai BISH 4/27/1972
115 Cyperus phleoides var. phleoides W Maui, Honokohau On side of cliff. PRESERVED_SPECIMEN Hosaka, E.Y. 1779 Maui BISH 2/10/1937
116 Cyperus phleoides var. phleoides e maui district, keopuka rock, off the n coast of e maui PRESERVED_SPECIMEN hobdy, r.w. collector number: 1399 Oahu BISH 5/16/1982
117 Cyperus phleoides var. phleoides nahiku moist forest, steep slope PRESERVED_SPECIMEN rock, j.f.c. collector number: 8755 Lanai BISH 5/10/1911
118 Cyperus phleoides var. phleoides makawao district, aleale stack; near puu koae 50 deg NE aspect; mixed native coastal shrubland w/Senna gaudichaudii, Lipochaeta lavarum, Waltheria, Chamaesyce celastroides var. amplectens, Bidens mauiensis, Panicum torridum, Eragrostis cf. atropioides; Occuring in shallow soil amidst talus; upper section of stack. PRESERVED_SPECIMEN wood, k.r. hobdy, r.w.; perlman, s. collector number: 2152 Kahoolawe BISH 11/24/1992
119 Cyperus phleoides var. phleoides Makawao District, Aleale Stack, near Puu Koae Mixed Native Coastal Shrubland with Portulaca molokiniensis, P. villosa, Senna gaudichaudii, Chamaesyce celastroides var. amplectens, Chamaesyce sp., Capparis, Lipochaeta lavarum, Bidens mauiensis, Sida fallax, Doryopteris decipiens, Panicum torridum, P. fauriei var. latius, Eragrostis atropioides, Ipomoea tuboides; indications of Bulwers Petrel and Wedgetail Shearwater nesting sites. Weeds: Waltheria, Emilia fosbergii, Chloris barbata, Nicotiana glauca. PRESERVED_SPECIMEN Wood, K.R. 1726 Kahoolawe BISH 3/18/1992
120 Cyperus phleoides var. phleoides W Maui District, coastal cliffs by Puu Koae PRESERVED_SPECIMEN Hobdy, R.W. 385 Maui BISH 10/30/1978
121 Cyperus phleoides var. phleoides Windward side of island, Anapuhi to Wainene, long coastal boulder-strewn beach with vertical cliff at back 10-30 m from water Cliff with Artemisia australis, Chenopodium oahuense, Schiedea globosa, Kalanchoe pinnata, Tetramolopium sylvae, Lythrum maritimum, and Hedyotis littoralis, among others. Back of rocky area near hala stand. PRESERVED_SPECIMEN Wagner, W.L. 5742 Molokai BISH 8/1/1987
122 Cyperus phleoides var. phleoides East Maui, Hana District, Hana, Kauiki Head, on SE side of Hana Bay Secondary forest dominated by Casuarina, Terminalia, Syzygium, Ardisia, and Lantana. On open, steep cinder faces on northeast slope. With Scaevola, Ipomoea, Bidens, Morinda, and Vigna. PRESERVED_SPECIMEN Flynn, T. 2412 Maui BISH 10/10/1987
123 Cyperus phleoides var. phleoides kaaukuu gulch, west molokai, near main road Dry grassy slope far from ocean. PRESERVED_SPECIMEN degener, o. collector number: 22206 Molokai BISH 3/16/1952
124 Cyperus phleoides var. phleoides Haleakala, lower ditch trail PRESERVED_SPECIMEN Skottsberg, C.J.F. 798 Maui BISH 10/14/1922
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