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Rubus macraei

A.Gray (1854)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Rosales Family: Rosaceae Genus: Rubus

‘ākala [akala], ‘ākalakala [akalakala], kala

Description

Key Characters:

Growth Form: Shrubs with shedding bark.

Stems: Stems prostrate or decumbent, sometimes rooting at the nodes, sparsely covered with slender prickles up to 3 mm long and densely tomentose, also with glandular hairs on younger parts. Primocanes and floricanes present and similar, somewhat thick, coriaceous when dry.

Roots:

Leaves: Leaves compound (palmate). Alternate. Blades very broadly ovate to suborbicular or rhombic–suborbicular, 5.1–8.5(–11) cm long, 3–6(–9) cm wide. leaflets 3, the terminal one largest, the lateral ones slightly asymmetrical. Upper surfaces uniformly sparsely pilose, occasionally the hairs more concentrated on or between the impressed veins; lower surfaces usually densely velvety tomentose, rarely only along the raised veins. Margins irregularly and usually coarsely serrate and usually somewhat lobed. Petiolate; petiolules 0.4–3 cm long, petiolules and petioles moderately to densely tomentose, sometimes with scattered small prickles, petioles 1.5–3.5(–7) cm long. Stipules narrowly oblong to linear–lanceolate, 0.8–1.5 cm long.

Flowers: Flowers 1–8 in corymbose inflorescences. Flowers bisexual (perfect); pedicels 8–55 mm long, densely tomentose and also usually covered with slender prickles. Calyx of 5 persistent sepals; Sepals ovate, 10–16 mm long, both surfaces densely tomentose, apex acuminate to caudate. Corolla of 5 petals, petals dark pink to dusky rose, obovate, 13–16 mm long. Stamens numerous. Ovary superior, carpels usually numerous, inserted on the raised receptacle, 1-celled; ovules 2 per cell; style filiform, subterminal, persistent.

Fruit: Fruit dark red to dark purple; juicy; ovoid; 2.54 cm long; densely pubescent. Seeds 1 per drupelet.

Ploidy:

Habitat: Uncommon to rare on steep rocky slopes and open rocky areas in wet forest and margins of bogs; sometimes in sheltered thickets of other plants in pastures or subalpine shrubland.

Elevation Range: 1,610–2,080 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: U.S. Expl. Exped., Phan. 1: 505 (1854)

Other References

Wagner et al. 1990:1109 (EM, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Rubus macraei A.Gray 21 mi w of hilo on saddle road PRESERVED_SPECIMEN miller, j.s. merello, m. collector number: 7919 Hawaii BISH 8/5/1992
2 Rubus macraei A.Gray paliku, haleakala Steep shrubby slope. PRESERVED_SPECIMEN degener, o. ordonez; salucop, f. collector number: 12512 Maui BISH 8/20/1939
3 Rubus macraei A.Gray 21 mi W of Hilo on Saddle Road PRESERVED_SPECIMEN Miller, J.S. Merello, M. Collector Number: 7919 Hawaii BISH 8/5/1992
4 Rubus macraei A.Gray S Hilo DISTR. Waiakea AHU. Tr 26(3); W of Puu Oo Volcano Trail in kipuka betwn 1881 & 1852 flows Mesic ohia forest with other native trees and a native shrub, matted-fern understory. Growing on pahoehoe substrate. PRESERVED_SPECIMEN Warshauer, F.R. FRW 1599 Hawaii BISH 12/9/1977
5 Rubus macraei A.Gray kau forest reserve, kau district Epiphitic on moss covered fallen log in closed canopy `Ohi`a - Koa forest. PRESERVED_SPECIMEN anderson, s.j. collector number: 464 Hawaii BISH 1/20/1982
6 Rubus macraei A.Gray Between Ohia and Koa Mills opposite road leading to Hoopuloa On rocky bank in rain forest. PRESERVED_SPECIMEN Degener, O. 8167 Hawaii BISH 3/29/1930
7 Rubus macraei A.Gray east maui, hana district, hana forest reserve At margin of an open bog. Trailing Vaccinium & Metrosideros. PRESERVED_SPECIMEN harrison, b. collector number: 323 Maui BISH 7/10/1973
8 Rubus macraei A.Gray east maui, haleakala, kipahulu-kuhiwa divide, wai anapanapa Occasional in thickets on divide, with the erect Rubus hawaiensis. PRESERVED_SPECIMEN st.john, h. mitchell, a.l. collector number: 20978 Maui BISH 8/13/1945
9 Rubus macraei A.Gray keauhou 2, road up hualalai steep north facing Metrosideros montane mesic to wet forest; associated with Xylosma hawaiiensis, Psychotria greenwelliae, Pritchardia minor, Pleomele aurea, Tetraplasandra waialealae, Pouteria sandwicensis. PRESERVED_SPECIMEN f. r. fosberg 42117 Kauai BISH 9/9/1961
10 Rubus macraei A.Gray Humuula. Forest around Puu Kipu, Kulani Growing in edge of Metrosideros forest with Styphelia, Vaccinium, sedges and ferns. PRESERVED_SPECIMEN Jacobson, J.D. 1001 Hawaii BISH 9/2/1984
11 Rubus macraei A.Gray palakea (?) PRESERVED_SPECIMEN forbes, c.n. collector number: 968.h Hawaii BISH 6/27/1915
12 Rubus macraei A.Gray east maui, hana district, hana forest reserve, n of kipahulu-waihoi divide Bog. PRESERVED_SPECIMEN st.john, h. mitchell, a.l. collector number: 21105 Maui BISH 8/22/1945
13 Rubus macraei A.Gray Kapapala Forest Reserve Growing in Styphelia thicket in open spot of Metrosideros-Acacia forest, occasional Dodonaea & Myoporum. Ground cover includes Deschampia, Coprosma ernodeoides; few patches Fragaria chiloensis var. sandwicensis in vicinity. PRESERVED_SPECIMEN Davis, J. 933 Hawaii BISH 1/13/1983
14 Rubus macraei A.Gray Kipuka Ahiu PRESERVED_SPECIMEN Forbes, C.N. 896.H Hawaii BISH 6/21/1915
15 Rubus macraei A.Gray East Maui, Haleakala PRESERVED_SPECIMEN Mann, H. 425 Maui BISH 2/6/1905
16 Rubus macraei A.Gray PRESERVED_SPECIMEN remy, m.j. collector number: 649 BISH 1851-01-01
17 Rubus macraei A.Gray Slopes of Hualalai above Honokane (?) PRESERVED_SPECIMEN Forbes, C.N. 179.H Hawaii BISH 6/16/1911
18 Rubus macraei A.Gray koolau gap of haleakala crater PRESERVED_SPECIMEN degener, o. wiebke, h. collector number: 2258 Hawaii BISH 8/17/1927
19 Rubus macraei A.Gray Kau Forest Reserve, Kau District Epiphitic on moss covered fallen log in closed canopy `Ohi`a - Koa forest. PRESERVED_SPECIMEN Anderson, S.J. 464 Hawaii BISH 1/20/1982
20 Rubus macraei A.Gray Koolau Gap of Haleakala Crater Creeping on fog-swept eroded aa slope. PRESERVED_SPECIMEN Degener, O. 2258 Maui BISH 8/10/1927
21 Rubus macraei A.Gray keauhou ranch (cr-l) kau district In kipuka of tall Metrosideros bisected by jeep trail. Log jam beside jeep trail. Substrate `a`a. PRESERVED_SPECIMEN esp field crew collector number: esp 113 Hawaii BISH 9/17/1979
22 Rubus macraei A.Gray east maui, n of kuiki just outside of haleakala section of park boundary Foggy region. PRESERVED_SPECIMEN degener, o. ordonez; salucop, f. collector number: 12511 Maui BISH 8/9/1939
23 Rubus macraei A.Gray Waianapanapa, Haleakala PRESERVED_SPECIMEN Forbes, C.N. 1176.M Maui BISH 8/20/1919
24 Rubus macraei A.Gray Kapapala Forest Reserve Growing in Styphelia thicket in open spot of Metrosideros - Acacia forest, occasional Dodonaea and Myoporum: ground cover includes Deschampsia, Coprosma ernodeoides and few patches of Fragaria chiloensis var. snadwicensis in vicinty. PRESERVED_SPECIMEN Davis, J. Collector Number: 933 Hawaii BISH 1/13/1983
25 Rubus macraei A.Gray East Maui, Haleakala, Kipahulu-Kuhiwa divide, Wai Anapanapa Occasional in thickets on divide, with the erect Rubus hawaiensis. PRESERVED_SPECIMEN St. John, H. 20978 Maui BISH 8/13/1945
26 Rubus macraei A.Gray east maui, hana district, hana forest reserve, n of kipahulu-waihoi divide Schinus and guava invested slopes, Pisonia sandwicensis the only native sp. in the vicinity. PRESERVED_SPECIMEN st.john, h. mitchell, a.l. collector number: 21105 Oahu BISH 8/22/1945
27 Rubus macraei A.Gray Near hill NE of Olinda Sheltered open thicket in cattle range. PRESERVED_SPECIMEN Degener, O. 2260 Maui BISH 6/15/1927
28 Rubus macraei A.Gray koolau gap of haleakala crater Creeping on fog-swept eroded `a`a lava slope. PRESERVED_SPECIMEN degener, o. wiebke, h. collector number: 2258 Maui BISH 8/10/1927
29 Rubus macraei A.Gray mauna kea, hawaii island PRESERVED_SPECIMEN j. macrae GH
30 Rubus macraei A.Gray mauna loa strip road, slopes of mauna loa. PRESERVED_SPECIMEN f.r. fosberg 57484 PH 3/14/1978
31 Rubus macraei A.Gray Keauhou 2, Road up Hualalai PRESERVED_SPECIMEN Fosberg, F. R. 42117 Hawaii US 9/9/1961
32 Rubus macraei A.Gray E. Maui, Koolau Gap, Kalialinui. subalpine shrubland/forest understory PRESERVED_SPECIMEN Hank Oppenheimer H80906 Maui PTBG 8/7/2009
33 Rubus macraei A.Gray E. Maui, Piinaau drainage basin. Subalpine shrubland. PRESERVED_SPECIMEN Hank Oppenheimer H71005 Maui PTBG 7/8/2010
34 Rubus macraei A.Gray PRESERVED_SPECIMEN J. Remy 649 PTBG 1851-01-01
35 Rubus macraei A.Gray E. Maui, Hanawi; east ridge from Frisbee Meadow. Elev. 6100-6800 ft. Metrosideros-Cheirodendron Montane Wet Forest PRESERVED_SPECIMEN K. R. Wood 4301 Maui PTBG 6/23/1995
36 Rubus macraei A.Gray Kipahulu valley, National Park lands, gulch below Kuiki. Deschampsia nubigena subalpine mesic grassland with forest patches on steep slopes, and headwater riparian zones. PRESERVED_SPECIMEN K. R. Wood 6784 Maui PTBG 9/26/1997
37 Rubus macraei A.Gray E. Maui, Ainahou area of Koolau Gap. Metrosideros wet forest PRESERVED_SPECIMEN K. R. Wood 6596 Maui PTBG 8/13/1997
38 Rubus macraei A.Gray puu huluhulu, humuula saddle PRESERVED_SPECIMEN f. r. fosberg 41751 Hawaii US 3/24/1961
39 Rubus macraei A.Gray Near hill NE of Olinda PRESERVED_SPECIMEN Degener, O. 8161 Maui US 7/27/2027
40 Rubus macraei A.Gray PRESERVED_SPECIMEN Remy, M. J. 649 Hawaii US 1851-00-00
41 Rubus macraei A.Gray Mouna [Mauna] Kea. PRESERVED_SPECIMEN Wilkes Explor. Exped. s.n. Hawaii US 1/13/1905
42 Rubus macraei A.Gray koolau gap of haleakala crater PRESERVED_SPECIMEN o. degener & h. wiebke 2258 Oahu US 8/11/1927
43 Rubus macraei A.Gray hana forest reserve, e maui PRESERVED_SPECIMEN b. harrison 323 Maui US 7/10/1973
44 Rubus macraei A.Gray koolau gap of haleakala crater PRESERVED_SPECIMEN o. degener & h. wiebke 2258 Oahu US 8/17/1927
45 Rubus macraei A.Gray haleakala, e maui PRESERVED_SPECIMEN h. mann & w. t. brigham s.n. Maui US
46 Rubus macraei A.Gray PRESERVED_SPECIMEN m. j. remy 649 Hawaii US 1851-08-08
47 Rubus macraei A.Gray Koolau Gap of Haleakala Crater PRESERVED_SPECIMEN Degener, O.; Wiebke, H. 2258 Oahu US 8/11/2027
48 Rubus macraei A.Gray Puu Huluhulu, Humuula Saddle PRESERVED_SPECIMEN Fosberg, F. R. 41751 Hawaii US 3/24/1961
49 Rubus macraei A.Gray Koolau Gap of Haleakala Crater PRESERVED_SPECIMEN Degener, O.; Wiebke, H. 2258 Oahu US 8/17/2027
50 Rubus macraei A.Gray Koolau Gap, Haleakala PRESERVED_SPECIMEN Degener, O.; Fleming, W. 25302 Maui US 5/28/1959
51 Rubus macraei A.Gray mouna [mauna] kea. PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US 1840-08-08
52 Rubus macraei A.Gray koolau gap, haleakala PRESERVED_SPECIMEN o. degener & w. fleming 25302 Maui US 5/28/1959
53 Rubus macraei A.Gray Hana Forest Reserve, E Maui PRESERVED_SPECIMEN Harrison, B. 323 Maui US 7/10/1973
54 Rubus macraei A.Gray near hill ne of olinda PRESERVED_SPECIMEN o. degener 8161 Maui US 7/27/1927
55 Rubus macraei A.Gray USA, Hawaiian Islands, Maui, east Maui, Hana District, Kahikinui, upper Kamole drainage basin, NNW of Pu'u Pane, SE of Kahua On steep gulch wall in shady area. Vegetation: Metrosideros/Acacia Montane Mesic Forest. PRESERVED_SPECIMEN Oppenheimer, H.L. H111404 Maui BISH 11/24/2014
56 Rubus macraei A.Gray e of ukulele PRESERVED_SPECIMEN forbes, c.n. collector number: 639.m Maui BISH 7/17/1919
57 Rubus macraei A.Gray puuouli, s slope of haleakala PRESERVED_SPECIMEN forbes, c.n. collector number: 2152.m Maui BISH 4/5/1920
58 Rubus macraei A.Gray Keauhou Ranch (Cr-l) Kau District In kipuka of tall Metrosideros bisected by jeep trail. Log jam beside jeep trail. Substrate `a`a. PRESERVED_SPECIMEN ESP Field Crew Collector Number: ESP 113 Hawaii BISH 9/17/1979
59 Rubus macraei A.Gray koolau gap of haleakala crater Creeping on fog-swept eroded aa slope. PRESERVED_SPECIMEN degener, o. wiebke, h. collector number: 2258 Maui BISH 8/10/1927
60 Rubus macraei A.Gray Koolau Gap, Haleakala PRESERVED_SPECIMEN Rock, J.F.C. 8654 Maui BISH 10/1/1910
61 Rubus macraei A.Gray kapapala forest reserve Growing in Styphelia thicket in open spot of Metrosideros - Acacia forest, occasional Dodonaea and Myoporum: ground cover includes Deschampsia, Coprosma ernodeoides and few patches of Fragaria chiloensis var. snadwicensis in vicinty. PRESERVED_SPECIMEN davis, j. collector number: 933 Hawaii BISH 1/13/1983
62 Rubus macraei A.Gray East Maui, Ainahou, Koolau Gap PRESERVED_SPECIMEN Hobdy, R.W. 772 Maui BISH 2/16/1980
63 Rubus macraei A.Gray waianapanapa, haleakala PRESERVED_SPECIMEN forbes, c.n. collector number: 1176.m Maui BISH 8/20/1919
64 Rubus macraei A.Gray NW side of Koolau Gap below cliffs, Haleakala Rainy `a`a lava. PRESERVED_SPECIMEN Degener, O. 12535 Maui BISH 8/20/1939
65 Rubus macraei A.Gray East Maui, Haleakala Crater, Koolau Gap, 2 mi N of Holua Cave Thicket at edge of woods. PRESERVED_SPECIMEN St. John, H. 21297 Maui BISH 9/4/1945
66 Rubus macraei A.Gray slopes of hualalai above honokane (?) PRESERVED_SPECIMEN forbes, c.n. collector number: 179.h Hawaii BISH 6/16/1911
67 Rubus macraei A.Gray 21 mi W of Hilo on Saddle Road PRESERVED_SPECIMEN Miller, J.S. 7919 Hawaii BISH 8/5/1992
68 Rubus macraei A.Gray between ohia and koa mills opposite road leading to hoopuloa On rocky bank in rain forest. PRESERVED_SPECIMEN degener, o. collector number: 8167 Hawaii BISH 3/29/1930
69 Rubus macraei A.Gray East Maui, N of Kuiki just outside of Haleakala Section of Park boundary Foggy region. PRESERVED_SPECIMEN Degener, O. 12511 Maui BISH 8/9/1939
70 Rubus macraei A.Gray East Maui, upper Hanawi Natural Area Reserve. Subalpine shrubland. PRESERVED_SPECIMEN Hobdy, R.W. 3282 Maui BISH 2/8/1991
71 Rubus macraei A.Gray near hill ne of olinda Sheltered open thicket in cattle range. PRESERVED_SPECIMEN degener, o. wiebke, h. collector number: 2260 Maui BISH 6/15/1927
72 Rubus macraei A.Gray E of Ukulele PRESERVED_SPECIMEN Forbes, C.N. 639.M Maui BISH 7/17/1919
73 Rubus macraei A.Gray Between Ohia and Koa Mills opposite road leading to Hoopuloa On rocky bank in rain-forest. PRESERVED_SPECIMEN Degener, O. 8166 Hawaii BISH 3/30/1930
74 Rubus macraei A.Gray S Hilo District. Upper Waiakea Forest Reserve. PowerLine Road, Pole 27, ca 2 mi S of the junction with Saddle Road. Kipuka just E of the road Assoc. with Vaccinium, Styphelia, Dryopteris, Metrosideros, Coprosma, Carex, Sadleria, etc. PRESERVED_SPECIMEN Gustafson, R. 2386 Hawaii BISH 7/21/1981
75 Rubus macraei A.Gray haleakala PRESERVED_SPECIMEN hillebrand, w.b. lydgate, j.m. collector number: s.n. Maui BISH
76 Rubus macraei A.Gray East Maui, Hana District, Hana Forest Reserve, N of Kipahulu-Waihoi divide Bog. PRESERVED_SPECIMEN St. John, H. 21105 Maui BISH 8/22/1945
77 Rubus macraei A.Gray s hilo district. upper waiakea forest reserve. powerline road, pole 27, ca 2 mi s of the junction with saddle road. kipuka just e of the road Assoc. with Vaccinium, Styphelia, Dryopteris, Metrosideros, Coprosma, Carex, Sadleria, etc. PRESERVED_SPECIMEN gustafson, r. collector number: 2386 Hawaii BISH 7/21/1981
78 Rubus macraei A.Gray nw side of koolau gap below cliffs, haleakala Rainy `a`a lava. PRESERVED_SPECIMEN degener, o. ordonez; salucop, f. collector number: 12535 Maui BISH 8/20/1939
79 Rubus macraei A.Gray East Maui, Hana District, Hana Forest Reserve, above N rim of Kipahulu At margin of a bog. Trailing over Vaccinium, with Plantago sp. & greensword. PRESERVED_SPECIMEN Harrison, B. 481 Maui BISH 9/23/1973
80 Rubus macraei A.Gray Kau Forest Reserve, Kau District Epiphitic on moss covered fallen log in closed canopy `Ohi`a - Koa forest. PRESERVED_SPECIMEN Anderson, S.J. Collector Number: 464 Hawaii BISH 1/20/1982
81 Rubus macraei A.Gray between ohia and koa mills opposite road leading to hoopuloa On rocky bank in rain-forest. PRESERVED_SPECIMEN degener, o. collector number: 8166 Hawaii BISH 3/30/1930
82 Rubus macraei A.Gray koolau gap, haleakala Rainswept plateau among other shrubs PRESERVED_SPECIMEN rock, j.f.c. collector number: 8654 Hawaii BISH 10/1/1910
83 Rubus macraei A.Gray humuula. forest around puu kipu, kulani Growing in edge of Metrosideros forest with Styphelia, Vaccinium, sedges and ferns. PRESERVED_SPECIMEN jacobson, j.d. collector number: 1001 Hawaii BISH 9/2/1984
84 Rubus macraei A.Gray east maui, hana district, haleakala crater, pass n of kuiki Rare in thicket. PRESERVED_SPECIMEN st.john, h. mitchell, a.l. collector number: 21114 Maui BISH 8/25/1945
85 Rubus macraei A.Gray east maui, haleakala On Natatorium outer wall and rock at base. PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 425 Oahu BISH
86 Rubus macraei A.Gray kipuka ahiu PRESERVED_SPECIMEN forbes, c.n. collector number: 896.h Molokai BISH 6/21/1915
87 Rubus macraei A.Gray haleakala PRESERVED_SPECIMEN forbes, c.n. collector number: s.n. Maui BISH 7/17/1919
88 Rubus macraei A.Gray kapapala forest reserve Growing in Styphelia thicket in open spot of Metrosideros-Acacia forest, occasional Dodonaea & Myoporum. Ground cover includes Deschampia, Coprosma ernodeoides; few patches Fragaria chiloensis var. sandwicensis in vicinity. PRESERVED_SPECIMEN davis, j. collector number: 933 Hawaii BISH 1/13/1983
89 Rubus macraei A.Gray s hilo distr. waiakea ahu. tr 26(3); w of puu oo volcano trail in kipuka betwn 1881 & 1852 flows Mesic ohia forest with other native trees and a native shrub, matted-fern understory. Growing on pahoehoe substrate. PRESERVED_SPECIMEN warshauer, f.r. collector number: frw 1599 Hawaii BISH 12/9/1977
90 Rubus macraei A.Gray Paliku, Haleakala Steep shrubby slope. PRESERVED_SPECIMEN Degener, O. 12512 Maui BISH 8/20/1939
91 Rubus macraei A.Gray Near hill NE of Olinda Sheltered open thicket in cattle range. PRESERVED_SPECIMEN Degener, O. 8161 Maui BISH 7/27/1927
92 Rubus macraei A.Gray Koolau Gap of Haleakala Crater Creeping on fog-swept eroded `a`a lava slope. PRESERVED_SPECIMEN Degener, O. 2258 Maui BISH 8/17/1927
93 Rubus macraei A.Gray near hill ne of olinda Sheltered open thicket in cattle range. PRESERVED_SPECIMEN degener, o. collector number: 8161 Maui BISH 7/27/1927
94 Rubus macraei A.Gray east maui, ainahou, koolau gap PRESERVED_SPECIMEN hobdy, r.w. collector number: 772 Maui BISH 2/16/1980
95 Rubus macraei A.Gray east maui, hana district, hana forest reserve, above n rim of kipahulu beach drift PRESERVED_SPECIMEN harrison, b. collector number: 481 Midway Atoll BISH 9/23/1973
96 Rubus macraei A.Gray Palakea (?) PRESERVED_SPECIMEN Forbes, C.N. 968.H Hawaii BISH 6/27/1915
97 Rubus macraei A.Gray PRESERVED_SPECIMEN Remy, M.J. 649 BISH 1/24/1905
98 Rubus macraei A.Gray Kapapala Forest Reserve Growing in Styphelia thicket in open spot of Metrosideros - Acacia forest, occasional Dodonaea and Myoporum: ground cover includes Deschampsia, Coprosma ernodeoides and few patches of Fragaria chiloensis var. snadwicensis in vicinty. PRESERVED_SPECIMEN Davis, J. 933 Hawaii BISH 1/13/1983
99 Rubus macraei A.Gray Koolau Gap of Haleakala Crater Creeping on fog-swept eroded `a`a lava slope. PRESERVED_SPECIMEN Degener, O. 2258 Maui BISH 8/10/1927
100 Rubus macraei A.Gray East Maui, Hana District, Haleakala Crater, Pass N of Kuiki Rare in thicket. PRESERVED_SPECIMEN St. John, H. 21114 Maui BISH 8/25/1945
101 Rubus macraei A.Gray east maui, haleakala crater, koolau gap, 2 mi n of holua cave Thicket at edge of woods. PRESERVED_SPECIMEN st.john, h. mitchell, a.l. collector number: 21297 Maui BISH 9/4/1945
102 Rubus macraei A.Gray Haleakala PRESERVED_SPECIMEN Forbes, C.N. s.n. Maui BISH 7/17/1919
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