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Psydrax odorata

(G.Forst.) A.C.Sm. & S.P.Darwin (1988)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Rubiaceae Genus: Psydrax

alahe‘e [alahee], ō‘he‘e, ʻō‘he‘e [ohee], walahe‘e [walahee]

Description

Key Characters:

Growth Form: Shrubs or small trees 3–6(–15) m tall.

Stems:

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly to broadly elliptic, elliptic–ovate, or lanceolate to ovate, 2.8–9 cm long, 0.9–4.1 cm wide. Apex obtuse to rounded, sometimes emarginate. Base cuneate. Surfaces glabrous, glossy on upper surface; lower surfaces less so or dull, somewhat thickened and coriaceous. Margins entire. Petioles 0–1 cm long. Stipules 2–5 mm long, interpetiolar, deltoid to ovate or lanceolate, lacking adaxial indument, apex abruptly acuminate to caudate, valvate in bud, usually bearing mucilate-producing colleters on inner surface that aid in protection of the growing shoot tip.

Flowers: Flowers 4- or 5-merous, numerous in corymbose cymes, cymes 2.5–3.5 cm long, pedicels 1–3 mm long, the bracts and bracteoles usually inconspicuous; hypanthium broadly ellipsoid to nearly hemispherical. Flowers bisexual (perfect) or functionally pistillate, protandrous, often heterostylous; fragrant, insect-pollinated. Calyx ca. 2 mm long, minutely dentate, limb truncate to dentate, 4-5-lobed with open aestivation, usually shorter than hypanthium, lobes usually small, rudimentary or absent. Corolla white, 4–6-lobed, 4–7 mm long, the lobes 3–4.5 mm long, reflexed, subequal to tube in length or longer or shorter, often thickened toward apex, valvate, imbricate, or convolute, usually obtuse to acute, tube broadly cylindric, often with deflexed hairs within and pubescent at throat; nectary disk cushion–shaped with a dense fringe of retrorse hairs. Stamens as many as and alternate with the corolla lobes, attached in corolla throat, exserted; filaments ca. 1.5–2.5 mm long, well developed; anthers ca. 1.5–2.5 mm long, dorsifixed near base, lanceolate to narrowly ovate, usually reflexed, dithecal, opening by longitudinal slits. Ovary inferior, 2-locular; ovules solitary in each locule, attached to the upper 1/3 of septum; style 5.5-7.5 mm long, slender, longer than corolla tube, glabrous, sometimes narrowing at apex, 1 or as many as carpels, terminal, the stigmatic knob cylindric, longer than broad, hollow to about the middle, bifid or deeply cleft at apex when mature; stigmas 2-lobed, dry or occasionally wet.

Fruit: Drupes 2-seeded black; juicy; obovoid; somewhat compressed; grooved on each side; emarginate; 8–10 mm long; the pyrenes cartilaginous to woody; rugulose to deeply furrowed; with a shallow apical cleft. Seeds incurved; with entire endosperm.

Ploidy:

Habitat: Dry shrubland and dry to mesic forest; often on slopes; also apparently in wet forest on O'ahu.

Elevation Range: 10–860(–1160) m.

Historical Distribution

Images

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Uses and Culture

USES

  • Black (leaves)

  • Durable wood used as cutting tool on softer woods and for farming tools, shark hooks, bait sticks (charred over fire); leaves for dye (Krauss 1993:25, Little & Skolmen 1989:296; Rock 1913:437; Wagner et al. 1990:1118); one of the hardest woods (Abbott 1992:12) that was fashioned into adzes (Kamakau 1976:122); sticks to keep pot lids open (Malo 1951:22). In the Ethnology Collection at Bishop Museum there is a post-contact example of the wood made into a bowl.

  • O'o - agricultural tool

PROPAGATION/CULTIVATION

  • The fruit of Psydrax odorata is about 3/8 inch in diameter, more or less round, and glossy black and juicy when ripe. The fruits are generally ripe in the late fall and winter. The pulpy fruits generally contain two seeds. Although many seeds are produced, they are often attacked by the larvae of the moth Orneodes objurgatella. These seeds have holes in them, are generally not viable, and should be discarded. Criley notes that fewer than 10% of seeds may be viable due to insect damage.
  • Psydrax odorata grows easily from healthy seeds. Culliney and Koebele note, however, that the seeds in unripe fruit will not germinate. If viable seeds are available, they should be removed from the pulpy fruit. The soft fruits can be broken up by hand in water; the lighter pulp will float and can be poured off. The seeds should then be washed thoroughly and dried on a paper towel. Before planting the seeds, soak them for 24 hours in cool tap water. Stratton's survey respondents recommended soaking for as long as 48 hours, changing the water daily. Discard seeds that float.
  • Culliney and Koebele recommend disinfecting the seeds by putting them in a solution of 1 part bleach and 9 parts water for 1/2 hour. After the seeds are removed from the bleach solution, soak them in tap water for a day using just enough water to cover the seeds.
  • Culliney and Koebele suggest sowing the seeds on the top of moistened vermiculite and covering them with a layer of moistened green sphagnum moss. As soon as the seeds begin to germinate, remove most of the moss layer. They write that germination takes about 1 month.
  • Other sources suggest planting the seeds at a depth of about twice their diameter in a soil-less, well-drained potting mix. NTBG suggests 1 part perlite to 1 part peat; Stratton's informants also suggested a mixture of 3 parts #2 perlite to 1 part Sunshine Mix #4. Keep the planting medium moist and place the containers in a shaded location. Again, these sources state that germination generally takes about 30 days, but it may take as long as 6 months.
  • In his germination studies, Obata found that untreated seeds had germination rates ranging from 5 to 30%. Yoshinaga found that fresh seeds of Psydrax odorata have a germination rate of 50%. The germination rate drops rapidly with storage at room temperature (55-84 degress F) and normal humidity of 58-98%. Some seeds remained viable after being stored for one year at 39 degrees F with a humidity of 10%. (Criley 1999; Culliney 1999; NTBG 1992; NTBG 1996; Obata 1967; Stratton 1998; Wagner 1990; Yoshinaga 1997; Yoshinaga 1998)
  • Reliable techniques for starting Psydrax odorata from cuttings have not yet been developed. Ginoza and Rauch tested 4 dilutions of rooting hormone with a 2 to 1 indolebutyric acid (IBA) to naphthaleneacetic acid (NAA) ratio (Dip'n Grow) on 4 inch cuttings. The dilutations ranged from 500 to 5,000 parts per million (ppm) of IBA. Vermiculite was used as the rooting medium and the cuttings were placed under intermittent mist of 6 seconds every 2 minutes. There was no significant difference in the results from the different treatments and the maximum rooting percentage obtained was 7%.
  • Criley reports limited success in rooting softwood cuttings under high humidity using 4,000 parts per million (ppm) indolebutyric acid (IBA). This work was done using an intermittent mist system which was on for 6 to 8 seconds every 5 or 6 minutes under 30% shade. He reports 10 percent of cuttings rooted with this treatment in 10 to 12 weeks. Increasing the IBA treatment to 5,000 and 10,000 ppm did not improve rooting success. (Bornhorst 1996; Criley 1998; Criley 1999; Culliney 1999; Ginoza 1997; NTBG 1992)
  • Successful techniques for propagating Psydrax odorata from air layers have not yet been developed. (Bornhorst 1996; NTBG 1992) [Data from Herring, E. C., & Criley, R. A. (2003). The Hawaiian Native Plant Propagation Web Site: Developing a Webbased Information Resource. HortTechnology, 13(3), 545-548. https://www.ctahr.hawaii.edu/hawnprop/]

Ethnobotanical Images

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Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Pollinators

Bibliography

Name Published In: Fl. Vit. Nova 4: 230 (1988)

Other References

Wagner et al. 1990:1119 (K, O, Mo, L, M, H [as Canthium odoratum]); Smith 1988:230/Herbst & Wagner 1999:31 (COMBNOV, Syn. C. odoratum = P. odorata); Staples & Herbst 2005:484 (KEY), 499 (DESCR)

Laau Hawaii (Abbott) p.12

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Psydrax odorata East Maui, Auwahi, lava flows Preserved_Specimen Rock, J.F.C. Maui BISH 1954-02-01
2 Psydrax odorata 9.0 mi W. of Naalehu Post Office on Hwy. 11 Preserved_Specimen Wagner, W.L. 5234 Hawaii BISH 1984-01-16
3 Psydrax odorata Haupu Mts., near Nawiliwili Bay Preserved_Specimen Forbes, C.N. 711 Kauai BISH 1910-10-31
4 Psydrax odorata Olowalu Valley Preserved_Specimen Forbes, C.N. 2261 Maui BISH 1920-05-08
5 Psydrax odorata Mahana Valley Preserved_Specimen Rock, J.F.C. 8097 Lanai BISH 1910-07-06
6 Psydrax odorata 7.7 mi northwest of Kalae turnoff Preserved_Specimen Ozaki, E.T. 1303 Hawaii BISH 1954-12-23
7 Psydrax odorata East Maui, Kaapahu Parce above roadside on east slope above bridge at Alalele rivermouth. Preserved_Specimen Welton, P. 1741 Maui BISH 1993-06-08
8 Psydrax odorata N Kona, Puuwaawaa, Huehue pipe line, water tanks 1-3 Preserved_Specimen Kondo, Y. 111.5 Hawaii BISH 1946-07-24
9 Psydrax odorata Molokai Preserved_Specimen Rock, J.F.C. 6152 Molokai BISH 1910-03-01
10 Psydrax odorata Koolau, Keeawaawa valley Preserved_Specimen Skottsberg, C.J.F. 19 Oahu BISH 1922-08-04
11 Psydrax odorata Piko trail, Keaau Makua FR Preserved_Specimen Davis, H. Oahu BISH 1932-11-25
12 Psydrax odorata Piko trail, Kahanahaiki Valley, Waianae Mts Preserved_Specimen Krauss, N.H. Oahu BISH 1933-12-30
13 Psydrax odorata Wahiawa, Poamoho trail, Paalaa, Koolau Mts Preserved_Specimen Kato, M. Oahu BISH 1955-11-30
14 Psydrax odorata Makua Valley, Waianae Mts Preserved_Specimen Krauss, N.H. Oahu BISH 1932-11-25
15 Psydrax odorata Halawa quarry trail Preserved_Specimen Kerr, M. Oahu BISH 1945-09-09
16 Psydrax odorata Puu Kau Preserved_Specimen Emerson, J.S. Hawaii BISH 1903-08-25
17 Psydrax odorata S. ridge of Kipapa Gulch, Waipio, Koolau Mts.; Zone 2 Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-05-15
18 Psydrax odorata Makaha Valley, Waianae Mts Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
19 Psydrax odorata N. Kona Preserved_Specimen Shea Hawaii BISH 1928-01-01
20 Psydrax odorata Mahana Preserved_Specimen Pekelo Jr., N. Molokai BISH 1972-03-20
21 Psydrax odorata Mahana, W. Molokai Preserved_Specimen Spaulding III, P. Molokai BISH 1982-08-20
22 Psydrax odorata Keawaula Preserved_Specimen MacCaughey, V. Oahu BISH 1915-03-27
23 Psydrax odorata Kakaauuku Gulch - W fork, west of Mahana site on east side of the gulch Preserved_Specimen Spaulding III, P. Molokai BISH 1982-06-09
24 Psydrax odorata W Maui, road to Kahakuloa Preserved_Specimen Hillebrand, W.B. Maui BISH 1836-08-04
25 Psydrax odorata Kaena Point. Preserved_Specimen Allison, A. Oahu BISH 2018-12-02
26 Psydrax odorata Kaua Preserved_Specimen UH student Oahu BISH 1958-04-26
27 Psydrax odorata Tantalus trail, toward Pali Preserved_Specimen Kerr, M. Oahu BISH 1945-09-02
28 Psydrax odorata Aiea Loop Trail, Koolau Mts. Preserved_Specimen Ziegler, M.F. Oahu BISH 1987-06-10
29 Psydrax odorata Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
30 Psydrax odorata Kaluaa, W Mts Preserved_Specimen Kondo, Y. Kauai BISH 1947-01-19
31 Psydrax odorata Waianae Mts., Palikea, firebreak trail Preserved_Specimen Pearsall, G.A. Oahu BISH 1953-07-25
32 Psydrax odorata Piko trail, Makua, Waianae Mts Preserved_Specimen Baker, R.J. Oahu BISH 1932-11-25
33 Psydrax odorata Maui Preserved_Specimen Wilder, G.P. Maui BISH 1936-08-04
34 Psydrax odorata Nanakuli Preserved_Specimen Mangelsdorf, A.J. Oahu BISH 1962-03-03
35 Psydrax odorata Kawaihapai Valley Preserved_Specimen Suehiro, A. Oahu BISH 1931-11-15
36 Psydrax odorata Kanepuu Preserved_Specimen Montgomery, S.L. Lanai BISH 1971-06-05
37 Psydrax odorata Leilehua Plain Preserved_Specimen Wilder, G.P. Oahu BISH 1912-01-01
38 Psydrax odorata 7 mi S of Kalae Terminal Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-23
39 Psydrax odorata Puuwaawaa Preserved_Specimen Corn, C. Hawaii BISH 1984-03-29
40 Psydrax odorata Hawaii Loa ridge trail. Preserved_Specimen Allison, A. Oahu BISH 2020-10-31
41 Psydrax odorata Maui Preserved_Specimen Wilder, G.P. Maui BISH 1913-01-01
42 Psydrax odorata Nanakuli, upper valley, between 2 ridges Preserved_Specimen Oahu BISH 1946-01-19
43 Psydrax odorata [Hawaiian Islands; U.S.A. Preserved_Specimen U.S. Exploring Expedition 407 BISH
44 Psydrax odorata Kaiholena Preserved_Specimen Munro, G.C. 14 Lanai BISH 1913-08-01
45 Psydrax odorata Kaloi Hana District Preserved_Specimen Woolford, H.A. 150 Maui BISH 1955-12-27
46 Psydrax odorata East Maui, Auwahi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2012 Maui BISH 1920-03-18
47 Psydrax odorata E of Puu Kaupakuhale, NE Kaala, Waianae Mts., Mokuleia Preserved_Specimen St.John, H. 13186 Oahu BISH 1933-05-14
48 Psydrax odorata East Maui, SW slope of Haleakala, above Kanaio Preserved_Specimen Alexander, A.M. 5345 Maui BISH 1948-05-17
49 Psydrax odorata Hanapepe Preserved_Specimen Faurie, A. 313 Kauai BISH 1909-12-01
50 Psydrax odorata Drive down Chain of Craters Road about 11.9 miles toward Kalapana from intersection with Crater Rim Road; or about 1 mile before Naulu Picnic Area road. Plant was uphill from a paved pulloff. (plot 6) Metrosideros - native shrub - Andropogon savanna community Preserved_Specimen Newell, C.L. 837 Hawaii BISH 1967-01-23
51 Psydrax odorata Kau Rd. nr. Honuapo on ancient lava flow (Kipuka) Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-24
52 Psydrax odorata Hawaii In mountain forests Preserved_Specimen Large tee, flowers whitish. Nelson, D. BISH
53 Psydrax odorata Kamoku, Kaiholena Gulch, ridge N grassy slope Preserved_Specimen 1 m shrub; flowers white; fruit immature Fosberg, F.R. 12493 Lanai BISH 1935-11-30
54 Psydrax odorata Hanalei, Napali Coast State Park, just off of trail just past Hoolulu valley Coastal mesic forest w/ Psydrax odorata, Polyscias racemosum, Charpentiera densiflora, Pandanus, Aleurites, Pittosporum napaliense, Diospyros, Oplismenus hirtellus, Eleph... Weedy dry area above trail past Hoolulu Preserved_Specimen Tree 6 ft tall with white trunk. Leaves dark glossy green, seeds dark mature with moth holes. N. Tangalin 4076 Kauai BISH 2015-01-30
55 Psydrax odorata Peacock Flats, Mokuleia FR Metrosideros Preserved_Specimen 16' x 2 tree Little Jr., E.L. 31035 Oahu BISH 1976-08-10
56 Psydrax odorata Kanepuu dry forest Preserved_Specimen Desha, E. Lanai BISH 1963-05-04
57 Psydrax odorata Napali coast, W Hanakoa Valley steep slope, woods Preserved_Specimen 8 m x 2 dm tree St.John, H. 23220 Kauai BISH 1947-12-31
58 Psydrax odorata Kawaihau District. Anahola, Moloaa Forest Reserve, along motorcycle track. Disturbed Metrosideros-Dicranopteris lowland windward Forest W/Panicum maximum, Syzygium cumnii, Osteomeles, Psidium cattleianum, Waltheria, Andropogon. Preserved_Specimen 15 ft. tall next to trail with immature and mature fruit. Dark green glossy leaves. White trunk. Tangalin, N. 2826 Kauai BISH 2011-09-27
59 Psydrax odorata west Maui, Lahaina District, Ukumehame Valley, back of valley, east fork. Vegetation: Acacia, Dodonaea, Wikstroemia, Diospyros, Erythrina, Heteropogon, Osteomeles, Waltheria, Rauvolfia, Pleomele, Bidens, Carex, Eragrostis, Leptecophylla, Sida, Peperomia, Leucaena, Melinis, Opuntia, Syzygium, Aleurites, Lantana. Preserved_Specimen Common trees in this area. Oppenheimer, H.L. 81232 Maui BISH 2012-08-24
60 Psydrax odorata Makua Valley, Waianae Mts dry hillside Preserved_Specimen 4 m tree MacDaniels, L.H. 82 Oahu BISH 1926-10-29
61 Psydrax odorata Kaena, Nihoa Gulch thickets Preserved_Specimen 6-20' trees St.John, H. 11144 Oahu BISH 1931-11-15
62 Psydrax odorata Hulopoe Gulch, Mahana woods Preserved_Specimen 4 m tree St.John, H. 18741 Lanai BISH 1938-04-09
63 Psydrax odorata Lihue District. Kalepa Forest Reserve Nonnative Shrubland with weedy grasses and occasional native speices, forestry outplantings w/ Santalum album, Santalum album X ellipticum, Eucalyptus, Grevillea robusta, Dodonea, Osteomeles, Wikstroemia. Casuarina, Leucaena, Urochola maxima, Stachytarpheta, Passiflora laurifolia, Psidium ca Preserved_Specimen Tree about 7 ft tall with immature fruit. Skinny dk green lvs and white trunk. Tangalin, N. 4142 Kauai BISH 2015-02-04
64 Psydrax odorata Pohakea Pass, Halona, Lualualei FR wooded gulch Preserved_Specimen 20' x 6 tree St.John, H. 13128 Oahu BISH 1933-05-12
65 Psydrax odorata Waimano trail, Koolau Mts dry, open slope Preserved_Specimen 8 m tree, slender St.John, H. 20954 Oahu BISH 1945-05-13
66 Psydrax odorata Flow of 1800, N. Kona dry aa flow Preserved_Specimen bushy tree 4 m. tall Fosberg, F.R. 44 Hawaii BISH 1936-08-08
67 Psydrax odorata Kaalaiki, Kau along highway Preserved_Specimen flower buds Meinecke, W.H. Hawaii BISH 1970-05-16
68 Psydrax odorata Hy. sw. of Seven Pools Park. Alt. near sea level Shrub,wind-swept. Flowers white. Preserved_Specimen Little Jr., E.L. 31113 Maui BISH 1976-08-25
69 Psydrax odorata Waianae Mts., central Lualualei, below Kanehoa, valley 1 dry Preserved_Specimen shrub Christophersen, E. 3675 Oahu BISH 1932-06-01
70 Psydrax odorata peak at head Kalama Valley, Maunalua, W fork, main divide windswept Preserved_Specimen 2 m shrub; leaves glossy; fruit not mature Fosberg, F.R. 13614 Oahu BISH 1937-03-07
71 Psydrax odorata Hawaii In mountain forests Preserved_Specimen Large tee, flowers whitish. Nelson, D. BISH 1779-01-01
72 Psydrax odorata 2.3 miles n.e. of Huehue Ranch House, Kaupulehu, N. Kona Dist open scattered orest on aa lava, common tree Preserved_Specimen up to 7 m. X 2 dm. St.John, H. 22514 Hawaii BISH 1947-01-02
73 Psydrax odorata Kealia trail, Waianae Mts dry; koa haole Preserved_Specimen 8' shrub Ozaki, E.T. 2 Oahu BISH 1952-10-05
74 Psydrax odorata Kealia trail, Waianae Mts dry, brushy Preserved_Specimen 4 m tree; leaves dark green glossy; fruit immature Fosberg, F.R. 12852 Oahu BISH 1936-02-02
75 Psydrax odorata Valley E of Hahaione dry Preserved_Specimen Judd Jr., A.F. 1619 Oahu BISH 1931-02-25
76 Psydrax odorata North Kona Distr., Puuanahulu dry forest on aa Preserved_Specimen tree 5 m. high Webster, G.L. 14003 Hawaii BISH 1968-06-30
77 Psydrax odorata Kuokala plateau, Keawaula heights Erythrina sandwicensis Preserved_Specimen small shrub, flowering Takeuchi, W.N. 2777 Oahu BISH 1986-08-23
78 Psydrax odorata Kau, Punaluu on aa flow Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-08
79 Psydrax odorata Hoalulu Valley, Hanakapiai steep slope Preserved_Specimen 7 m x 2 dm tree; leaves shiny St.John, H. 23160 Kauai BISH 1948-01-01
80 Psydrax odorata Pololu Valley, lower center part of valley, where Waiakalae and Pololu streams diverge rare in the valley, abundant on ridges above. Preserved_Specimen Terry, R. Hawaii BISH 1985-07-01
81 Psydrax odorata S of Kaiwikoele Stream between koa and ohia zones Preserved_Specimen Degener, O. 30495 Oahu BISH 1966-03-29
82 Psydrax odorata Manoa, Haena, valley wet, under mango, dappled sun Preserved_Specimen 3-5 m tree Earle, E. 33 Kauai BISH 1972-06-25
83 Psydrax odorata Puna: Wahaula heion Shade. Preserved_Specimen Young shoot from shade. Stone, B.C. 2901 Hawaii BISH 1959-06-21
84 Psydrax odorata Lihue, Hoary Head Mts., N Haupu Diospyros, Canthium, Bobea, Pisonia, Aleurites, Caesalpinia, Psidium Preserved_Specimen 8 m x 20 cm tree; crown dense; leaves shiny green; fruit dark green Lorence, D.H. 5833 Kauai BISH 1988-03-04
85 Psydrax odorata 6 mi S of Manuka Park aa lava; dry, open Metrosideros forest Preserved_Specimen 15'x3.5 tree with horizontal branches; bark smooth, grayish; foliage dark green, glossy above, light below; fruit dark green Herbst, D.R. 943 Hawaii BISH 1968-02-23
86 Psydrax odorata Koolau Mts., Waimea Falls, canyon to N, ~2 miles from confluence dry basalt; Canthium, Osteomeles, Wikstroemia, Diospyros Preserved_Specimen Gillett, G.W. 1514 Oahu BISH 1964-03-07
87 Psydrax odorata Kaaweiki ridge dry scrub Preserved_Specimen Degener, O. 27703 Kauai BISH 1960-11-24
88 Psydrax odorata Haleakala side of ridge, between Lahainaluna School and David Malo's grave Mountains above Lahaina. Preserved_Specimen Tree 10 ft. high with dense foliage and profuse with fragrant white flowers. Gillett, G.W. 2009 Maui BISH 1966-10-11
89 Psydrax odorata Isaac Hale Beach Park, 20 mi se. of Hilo common on lava flow, roadsides Preserved_Specimen Small tree 15 ft. Little Jr., E.L. 31064 Hawaii BISH 1976-08-17
90 Psydrax odorata Nihoa Gulch, Mokuleia dry Preserved_Specimen tall shrub Hume, E.P. 313 Oahu BISH 1931-11-15
91 Psydrax odorata ridge 3/4 mi N of Laaukahi, Koloa, NW facing slope edge of forest Preserved_Specimen 7 m x 15 cm tree St.John, H. 23016 Kauai BISH 1947-12-24
92 Psydrax odorata Along Waimano Trail Just past tree plantings. Preserved_Specimen Corn, C. Oahu BISH 1978-02-10
93 Psydrax odorata Hulopoe Gulch, head of, at Munro Trail forest Preserved_Specimen spindly tree Degener, O. 28580 Lanai BISH 1963-08-11
94 Psydrax odorata Kawiwi-Kaala ridge, Makaha-Waianae Kai wooded Preserved_Specimen 6 m tree St.John, H. 17606 Oahu BISH 1935-03-31
95 Psydrax odorata Napali coast, Hanakapiai, ridge woods Preserved_Specimen St.John, H. 10987 Kauai BISH 1931-12-02
96 Psydrax odorata Kipu, crest of Hoary Heads between the jeep road and Queen Victoria's profile Dry mixed native and exotic forest. Preserved_Specimen Small tree with light gray bark; glossy, dark green foliage. Common; all with mature fruit. Herbst, D.R. 2362 Kauai BISH 1972-03-08
97 Psydrax odorata Kahelunui Valley Reminant ohia-koa forest heavily grazed with guava and lantana. North face of gulch. Preserved_Specimen Corn, C. Kauai BISH 1977-06-22
98 Psydrax odorata Kealakomo, Hawaii National Park Found in aa lava Preserved_Specimen A spreading tree 15 ft. tall with shiny foliage Fagerlund, G.O. 376 Hawaii BISH 1943-02-18
99 Psydrax odorata Volcano Road below Pahala aa lava Preserved_Specimen Fosberg, F.R. 10144 Hawaii BISH 1933-08-31
100 Psydrax odorata Along Haena-to-Kalalau Trail, ca. 2 1/2 miles in from Haena On ledges above trail Preserved_Specimen Shrubs to ca. 3 m high with ner-vertical main branches giving the whole the form of a candelabra. Fl;s. white; fruit not yet ripe. Fay, J.J. 154 Kauai BISH 1973-12-18
101 Psydrax odorata Located down Chain of Craters Road about 28.5 miles toward Kalapana from intersection with Crater Rim Road; or 0.2 miles before Queens Bath. Plant was about 0.25 miles up the Royal Estates Road, not far from HVNP boundary. (plot 27) Mixed lowland scrub community Preserved_Specimen Newell, C.L. 111 Hawaii BISH 1966-07-20
102 Psydrax odorata Mokuleia, N Waianae Mts, Kealia Trail above Dillingham Airfield. On fully exposed N facing rocky substrate. With Syzygium cumini, Urochloa maxima, Leucaena leucocephala. Preserved_Specimen Tree, 3 m tall. Kennedy, B. 116 Oahu BISH 2021-01-10
103 Psydrax odorata W Maui, Lahaina District, Lihau Lowland Dry Forest at 1350 ft on a north facing cliff face nestled with Dodonaea, Gouania hillebrandii, Pittosporum, Diospyros, Metrosideros, Alyxia, Bidens, Achyranthes, Chenopodium, and an assortment of native and alien grasses and herbs Preserved_Specimen Welton, P. 1356 Maui BISH 1991-10-27
104 Psydrax odorata Kaaawa-Kamokunui ridge, Mokuleia brush savanna Preserved_Specimen 8' shrub Grant, M.L. 7009 Oahu BISH 1934-07-11
105 Psydrax odorata Kapua on old lava flow, associated with Opuntia, Cassia glauca and Lantana Preserved_Specimen Small tree 4-1/2 m. high; flower small yellow green, fragrant. Bryan Jr., E.H. 749 Hawaii BISH 1932-11-24
106 Psydrax odorata Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road. Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Metrosideros - Diospyros mixed forest with patchy stands of Aleurites on aa lava Preserved_Specimen Shrub about 6 feet tall Newell, C.L. 491 Hawaii BISH 1966-08-20
107 Psydrax odorata Hill above Kepapa Springs North facing slope. Preserved_Specimen Corn, C. Kauai BISH 1977-07-18
108 Psydrax odorata Na Pali Coast, Nualolo Kai Growing in old Hawaiian terrace with Leaucena leucocephala and Nototrichium sandwicense. Preserved_Specimen Small tree or shrub. Clarke, G. 424 Kauai BISH 1979-07-20
109 Psydrax odorata West side of Halawa Valley hArid rocky slope Preserved_Specimen Degener, O. 9386 Molokai BISH 1928-06-20
110 Psydrax odorata Puu Kaua, Waianae Mts., Honouliuli wooded ridge Preserved_Specimen 15' tree St.John, H. 12201 Oahu BISH 1932-11-06
111 Psydrax odorata West side of 1907 flow moist forest Preserved_Specimen Fosberg, F.R. 48 Hawaii BISH 1936-08-10
112 Psydrax odorata Mahanaloa Valley, jct. and above Kuia Valley, ~2.5 mi W of Hwy. 55 kukui, Diospyros, Pelea, Pleomele Preserved_Specimen 3 m tree, leaves shiny, fruit green Lorence, D.H. 5110 Kauai BISH 1987-03-10
113 Psydrax odorata 2nd gulch, E of Puu Kaupakuhale, Mokuleia, Kaala, Waianae Mts dry wooded ridge, E of gulch Preserved_Specimen 4 m tree Fosberg, F.R. Oahu BISH 1932-10-23
114 Psydrax odorata Napali Coast, Maunapuluo, Hanakapiai cliff Preserved_Specimen St.John, H. 10869 Kauai BISH 1930-12-30
115 Psydrax odorata ridges above Aiea Acacia koa, Metrosideros Preserved_Specimen Iltis, H.H. 608 Oahu BISH 1967-08-13
116 Psydrax odorata Upper Makakilo, adjacent to Palehua Heights subdivision, on southwestern slope above Kalo‘i Gulch On steep side slope within overstory of Erythrina sandwicensis population, groundcover dominated by Panicum maximum; associates include Solanum americanum, Leucaena leucocephala, Sida fallax, Plumbago zeylanicum, Desmanthus. Preserved_Specimen Single healthy 14 ft tall tree, dbh 5 inches. Imada, C. 200435 Oahu BISH 2004-03-30
117 Psydrax odorata Kaupulehu Forest Reserve, Section B. Open bog, dry a'a', native forest at 1950 ft. Preserved_Specimen Rare, few small, stunted trees. Herbst, D.R. 2993 Hawaii BISH 1973-05-03
118 Psydrax odorata Hanalei, Napali Coast State Park, just off of trail just past Hoolulu valley Coastal mesic forest w/ Psydrax odorata, Polyscias racemosum, Charpentiera densiflora, Pandanus, Aleurites, Pittosporum napaliense, Diospyros, Oplismenus hirtellus, Eleph... Preserved_Specimen 7 dt tall, lvs [illegible] raised mid and [illegible] N. Tangalin 4082 Kauai BISH 2015-01-30
119 Psydrax odorata Lihue, Niumalu, Hoary Head Mts., below Keopaweo Syzygium cumini, Chrysophyllum oliviforme, Schinus, Desmodium, Oplismenus, Phymatosorus Preserved_Specimen 10' tree; bark whitish; leaves glossy, yellowish-green avove, dull green below; mature fruit dark brown, ~black, 1-2 seeded Flynn, T.W. 3179 Kauai BISH 1988-10-14
120 Psydrax odorata Punaluu near shore Preserved_Specimen small trees St.John, H. 11314 Hawaii BISH 1931-12-25
121 Psydrax odorata Haupu, ridge dry Preserved_Specimen 4 m tree MacDaniels, L.H. 875 Kauai BISH 1927-02-26
122 Psydrax odorata Kaala rainy woods Preserved_Specimen Kraynik, W.R. 30760 Oahu BISH 1962-12-30
123 Psydrax odorata Manoa-Palolo ridge forest Preserved_Specimen small trees St.John, H. Oahu BISH 1933-03-26
124 Psydrax odorata Behind Keawaula Beach Underneath kiawe. Preserved_Specimen Corn, C. Oahu BISH 1977-09-30
125 Psydrax odorata Waimea, Koaie Canyon Melia, Diospyros, Hibiscadelphus Preserved_Specimen 15' tree; bark whitish; branching layered or planar; leaves dark, glossy green with whitish-green midvein above, below dull, pale green; immature fruit dark green Flynn, T.W. 3751 Kauai BISH 1990-01-31
126 Psydrax odorata Polihale State Park, mouth of Kaulaula vally, south of Haeleele, Secondary non-native vegetation dominated by leucaena leucocephala and Prosopis palida, w/ some relic trees of Erythrina sandwicensis, Psydrax odorata Preserved_Specimen Tree, 6 m tall, trunk gray-white, well branched, leaves dark green, in early fruit, uncommon Wood, K.R. 15853 Kauai BISH 2014-02-27
127 Psydrax odorata Kipu, crest of Hoary Heads between the jeep road and Queen Victoria's profile Dry mixed native and exotic forest. Preserved_Specimen Small tree with light gray bark; glossy, dark green foliage. Common; all with mature fruit. Herbst, D.R. 2362 Kauai BISH 1972-03-08
128 Psydrax odorata Koolau Mts., Wilhelmina Hts. trail koa Preserved_Specimen small tree, white flowers Herat, T. 489 Oahu BISH 1973-08-16
129 Psydrax odorata Waimano trail lower open forest Preserved_Specimen 4 m bush Kuykendall, J.R. 70 Oahu BISH 1943-06-10
130 Psydrax odorata Naulu Forest, Kealamomo Common, making undergrowth in open forest on aa lava Preserved_Specimen seedling/slender tree 3 m. tall (others much taller, fruit green Fosberg, F.R. 46039 Hawaii BISH 1965-03-19
131 Psydrax odorata Hoalulu Stream, Hanakapiai moist forest Preserved_Specimen 15' tree; flower white, sweet scented St.John, H. 13877 Kauai BISH 1933-12-30
132 Psydrax odorata Puna. Kamoamoa, on the coast On the coast. Preserved_Specimen Small trees - erries green; fls. green. Stone, B.C. 2849 Hawaii BISH 1989-06-20
133 Psydrax odorata Polihale State Park, mouth of Kaulaula valley, south of Haeleele. Secondary non-native vegetation dominated by Leucaena leucocephala and Prosopis pallida, w/ some relic trees of Erythrina sandwicensis, Psydrax odorata. Preserved_Specimen Tree, 6.5 m tall, trunk gray-white, well branched, leaves dark green, in fruit, uncommon, SEED COLL. NTBG ACC #20140059 Wood, K.R. 15854 Kauai BISH 2014-02-27
134 Psydrax odorata Waialae Nui Ridge, Koolau Mts Eucalyptus, Psidium guajava, Acacia koa Preserved_Specimen 7' x 1.5 tree; leaves glossy above, 4-6 cm long; flower white in cymose corymbs; fruit obovoid drupe, black when ripe Spence, G. 2 Oahu BISH 1971-10-17
135 Psydrax odorata Waianae Kai FR moist gully; under Acacia, Psidium Preserved_Specimen 3-4 m shrub Takeuchi, W.N. 9 Oahu BISH 1983-12-11
136 Psydrax odorata 2 mkiles n.e. of Huehue Ranch House, Kaupulehu, N. Kona Dist in scrub on aa lava Preserved_Specimen 5 m. tree St.John, H. 22450 Hawaii BISH 1946-12-28
137 Psydrax odorata Kanepuu-Kaa Osmanthus-Diospyros dry forest Preserved_Specimen 2 m tree with branching at base; leaves opposite, elliptic-oblong, 4x2 cm; glossy green on top, dull beneath. Inflors, a cymose corymb. Fruit a fleshy drupe, obovid, 7-8 mm diam. Spence, G. 163 Lanai BISH 1973-07-02
138 Psydrax odorata Kau Forest Reserve on rough lava Preserved_Specimen Tree 6 m. high MacDaniels, L.H. 265 Hawaii BISH 1926-11-16
139 Psydrax odorata Kawaa Kau along highway Preserved_Specimen flowers Meinecke, W.H. Hawaii BISH 1970-05-17
140 Psydrax odorata Kapuna Below Mokuleia trail. Preserved_Specimen Welton, P. 588 Oahu BISH 1990-10-19
141 Psydrax odorata N Kona, N of Puuwaawaa, near main road dry lava Preserved_Specimen Cranwell, L.M. 3219 Hawaii BISH 1938-09-11
142 Psydrax odorata Near Halawa Preserved_Specimen Shrub 10 ft. Dry shrub type. Little Jr., E.L. 31223 Molokai BISH 1976-09-16
143 Psydrax odorata ridge, Palolo to Waialae Nui dry Preserved_Specimen 3 m Fosberg, F.R. 9713 Oahu BISH 1933-08-03
144 Psydrax odorata Kau on rough lava Preserved_Specimen Tree 6 m. high, fls. very fragrant MacDaniels, L.H. 256 Hawaii BISH 1926-11-27
145 Psydrax odorata 15 mi n.w. of Kailua, on road to Waimea common in dry open forest on aa lava Preserved_Specimen Small tree 6 m. tall; leaves glossy; fruit immature. Fosberg, F.R. 41747 Hawaii BISH 1961-03-24
146 Psydrax odorata Kaiholena Valley, NE ridge lower foggy forest Preserved_Specimen Degener, O. 24203 Lanai BISH 1957-08-17
147 Psydrax odorata 6 mi S of Manuka Park aa lava; dry, open Metrosideros forest Preserved_Specimen 15'x3.5 tree with horizontal branches; bark smooth, grayish; foliage dark green, glossy above, light below; fruit dark green Herbst, D.R. 943 Hawaii BISH 1968-02-23
148 Psydrax odorata 6 mi S of Manuka Park aa lava; dry, open Metrosideros forest Preserved_Specimen 15'x3.5 tree with horizontal branches; bark smooth, grayish; foliage dark green, glossy above, light below; fruit dark green Herbst, D.R. 943 Hawaii BISH 1968-02-23
149 Psydrax odorata Waimea Falls Park dry forest Preserved_Specimen 12' x 2 tree, corolla white Little Jr., E.L. 31310 Oahu BISH 1978-10-20
150 Psydrax odorata Koolau Mts koa, Metrosideros Preserved_Specimen 8-12' x 3 bush Woolford, H.A. 245 Oahu BISH 1956-01-08
151 Psydrax odorata Honolulu, Lanipo Trail Near utility pole at first valley portion of trail. Preserved_Specimen Tree, 10 ft ht, with black heart-shaped fruit, flowers not seen. Kennedy, B. 10 Oahu BISH 1995-11-21
152 Psydrax odorata No. Kona District: Hwy. 190, about 1-1/2 miles S of the turnoff to the paved road that leads to Puuwaawaa Dry forest area. Associated with Erythrina taihitensis, Nototrichium sandwicense, Peperomia, Argemone glauca, Diospyros ferrea, etc. Preserved_Specimen large shrub, 10' tall; flws, white, fragrant Gustafson, R. 1081 Hawaii BISH 1978-07-13
153 Psydrax odorata Manoa, Haena, edge of pali sun, small trees Preserved_Specimen 3-4 m tree Earle, E. 31 Kauai BISH 1972-07-01
154 Psydrax odorata Kipapa Old lava flow. Preserved_Specimen Small tree 2 m ht, flowers white, no fruit. Kirch, P.V. 550 Maui BISH 1996-07-21
155 Psydrax odorata ridge above Woodlawn, Manoa dry cliff Preserved_Specimen 2 m tree Fosberg, F.R. 9293 Oahu BISH 1933-03-19
156 Psydrax odorata mauka of Kaluanui Flats, above bench dry forest Preserved_Specimen Degener, O. 30733 Lanai BISH 1967-01-24
157 Psydrax odorata Makaha Valley, Waianae Mts thicket Preserved_Specimen 5 m tree St.John, H. 24863 Oahu BISH 1952-11-30
158 Psydrax odorata East Maui, S side of Haleakala, 2 mi beyond Ulupalakua In dry forest area on aa lava, with Diospyros ferrea, Erythrina sandwicensis, Reynoldsia sandwicensis, Rauwolfia sandwicensis, Dodonaea viscosa, Cocculus ferrandianus. Preserved_Specimen Rounded tree 15 feet tall, foliage dark green. Carlquist, S. 2390 Maui BISH 1967-03-14
159 Psydrax odorata 2 mi E. of Honuapo on Govt. Road old aa flow; very dry, windy, hot Preserved_Specimen Tree 12', branching. Diam, breast high 6. Flower white-small-pleasant odor. Fruit 2-seeded dark green; 4 grooved berry, no odor. Bark fairly smooth, gray-black. Leaves very glossy and smooth Russ, G.W. Hawaii BISH 1927-04-16
160 Psydrax odorata On dry aa lava along Piilani Highway, about 1/2 mile E of Ulupalakua Growing in a small gully with Nesoluma, Rauvolfia, Ipomoea. Preserved_Specimen Crooked, gnarled shrub about 6 ft. tall, with leathery, glossy, dark green leaves, and white flowers. Herbst, D.R. 839 Maui BISH 1968-01-01
161 Psydrax odorata Hanalei, Kalalau, Lower valley, up Southwestern drainage below Kalahu Psydrax degraded mesic forest dominated by invasive Syzygium cumini, w/ cliff species of Lobelia niihauensis, Doryopteris decipiens, Carex wahuensis, Carex meyenii, Wikstroemia oahuensis, Lipochaeta connata acris, Artemisia australis, Euphorbia celastroides hanapepen... Preserved_Specimen Tree, 6 m tall, with fruit, dark purple-black, common K.R. Wood 19494 Kauai BISH 2024-05-21
162 Psydrax odorata Honopu Beach Growing on talus slopes near cliff. Preserved_Specimen ESP Field Crew 20 Kauai BISH 1979-07-20
163 Psydrax odorata Haupu, Kipu Preserved_Specimen 15' x 3 tree St.John, H. 13649 Kauai BISH 1933-12-25
164 Psydrax odorata Waianae Mts., Dupont trail Preserved_Specimen 2..7 m shrub Chock, A.K. 812 Oahu BISH 1952-12-30
165 Psydrax odorata bluff above Kaena Pt., ridge Preserved_Specimen fruit dark green Fosberg, F.R. 13154 Oahu BISH 1936-05-31
166 Psydrax odorata Honolulu, Bishop Museum Campus, native Hawaiian garden. Preserved_Specimen Tree 12-15 ft ht; flowers white; fruits black Kennedy, B. 64 Oahu BISH 2018-07-25
167 Psydrax odorata Kaiholena Ridge grassy, shrubby slope Preserved_Specimen Degener, O. 24201 Lanai BISH 1957-08-01
168 Psydrax odorata Auwahi, Ulupalakua-Hana Road, 4 miles SE of Ulupalakua ranch house Preserved_Specimen Tree 18 ft. high, 2dm. d.b.h. Chock, A.K. 321 Maui BISH 1951-12-26
169 Psydrax odorata Kealia Pali Preserved_Specimen 2-3 m x 2-3 cm tree; flower white, fragrant; fruit green to blackish Bryan Jr., E.H. 782 Oahu BISH 1934-09-25
170 Psydrax odorata bluff above Kaena Pt., ridge Preserved_Specimen fruit dark green Fosberg, F.R. 13154 Oahu BISH 1936-05-31
171 Psydrax odorata East Maui, Auwahi, in scrub ona a lava Preserved_Specimen St.John, H. 24736 Maui BISH 1951-12-26
172 Psydrax odorata Kaala Preserved_Specimen Mann, H. 346 Oahu BISH 1864-01-01
173 Psydrax odorata Kamohamoa, near the beach Preserved_Specimen Shrub to 3 m.; flowers white; fruits unripe, green Kiehn, M. 89072831 Hawaii BISH 1989-07-28
174 Psydrax odorata Kalaupapa; Papapaiki, outside fence Preserved_Specimen shrub or small tree Hughes, G.D. 1118 Molokai BISH 2005-01-01
175 Psydrax odorata Honolulu, Bishop Museum Campus, native Hawaiian garden. Preserved_Specimen Tree 15 ft ht with fruit Kennedy, B. 20176 Oahu BISH 2017-11-09
176 Psydrax odorata Kalalau Trail, near first ridge beyond (W of) Hanakapiai Preserved_Specimen Sm. glossy-lvd. tree on precipitious slope. Stone, B.C. 1469 Kauai BISH 1956-12-24
177 Psydrax odorata East Maui, Hana District, Kaloi, dry scrub. Roadside in old aa lava flow, with Lantana Camara, Opuntia maga cantha and Hunnemannia Preserved_Specimen Large scrub 5m. alt. and 5m. crown. Diam of trunk ldm. Flower white, fragrant. Woolford, H.A. 150 Maui BISH 1955-12-27
178 Psydrax odorata Maui Preserved_Specimen Forbes, C.N. 2079 Maui BISH 1920-03-23
179 Psydrax odorata Makaleha Valley, central ridge forest Preserved_Specimen Degener, O. 21269 Oahu BISH 1951-01-12
180 Psydrax odorata Bishop Museum, Atherton Halau Preserved_Specimen 2.5 m tree, single trunk, dense, roundish crown; flower white, sweet (afternoon) Lau, J. 1652 Oahu BISH 1985-11-18
181 Psydrax odorata Kau Road, Honuapo ancient lava flow Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-28
182 Psydrax odorata Makua shrubbery Preserved_Specimen Tsuji, N. Oahu BISH 1932-11-25
183 Psydrax odorata Puu Haupu, Ss. part of id Preserved_Specimen Small tree on W slope Stone, B.C. 1633 Kauai BISH 1956-12-31
184 Psydrax odorata Waialae Nui rocky ledge Preserved_Specimen Takeuchi, W.N. 2194 Oahu BISH 1985-05-10
185 Psydrax odorata Waianae Mts., Dupont Trail, opposite Kaupakuhale Preserved_Specimen Krajina, V.J. 620225320 Oahu BISH 1962-02-25
186 Psydrax odorata Wailupe Valley Preserved_Specimen Forbes, C.N. 2510 Oahu BISH 1917-05-04
187 Psydrax odorata Mokuleia, Waianae Mts., E Makaleha valley, W branch Preserved_Specimen Chock, A.K. 225 Oahu BISH 1951-04-29
188 Psydrax odorata Wailupe Valley, R fork Preserved_Specimen Garber, D.W. 149 Oahu BISH 1920-01-12
189 Psydrax odorata Kaalaiki, Kau; along highway Preserved_Specimen Meinecke, W.H. Hawaii BISH 1970-05-16
190 Psydrax odorata Waiahole ditch trail Preserved_Specimen Meebold, A. Oahu BISH 1932-06-01
191 Psydrax odorata Halawa trail, quarry to divide Preserved_Specimen Oahu BISH 1945-09-09
192 Psydrax odorata Waimea Canyon Hwy Preserved_Specimen Little Jr., E.L. 23025 Kauai BISH 1968-05-30
193 Psydrax odorata Puuwaawaa Preserved_Specimen Forbes, C.N. 32 Hawaii BISH 1911-06-08
194 Psydrax odorata ridge, mauka gap, Lihue Preserved_Specimen Forbes, C.N. 15 Kauai BISH 1909-07-08
195 Psydrax odorata Kona, Huehue Preserved_Specimen Rock, J.F.C. 3499 Hawaii BISH 1909-06-07
196 Psydrax odorata Laaukahi, Haiku, ridge Preserved_Specimen 15' tree, compact St.John, H. 13475 Kauai BISH 1933-12-22
197 Psydrax odorata Lanai Preserved_Specimen Munro, G.C. 18 Lanai BISH
198 Psydrax odorata Kau side of Kilauea on dry lava Preserved_Specimen Guild, E. Hawaii BISH 1957-03-01
199 Psydrax odorata Kau Desert Preserved_Specimen Skottsberg, C.J.F. 600 Hawaii BISH 1922-09-29
200 Psydrax odorata Kawainui Stream, ~8 mi E of Haleiwa Preserved_Specimen 10' shrub; flower buds; leaves shiny dark green above, dull light green below Little Jr., E.L. 31233 Oahu BISH 1978-10-08
201 Psydrax odorata Wiliwilinui ridge, Waialae Iki - Wailupe, Koolau Mts Preserved_Specimen bush; flower white Shinohara, B. 244 Oahu BISH 1947-05-11
202 Psydrax odorata N. slope of Hualalai, near flow of 1801 Preserved_Specimen Krajina, V.J. 611230040 Hawaii BISH 1961-12-30
203 Psydrax odorata Kauai Preserved_Specimen Large Shrub Jones, J. Kauai BISH 1965-03-03
204 Psydrax odorata Lanai Preserved_Specimen Mann, H. 346 Lanai BISH 1864-01-01
205 Psydrax odorata Makaha Valley, ridges above Preserved_Specimen MacDaniels, L.H. 597 Oahu BISH 1927-01-31
206 Psydrax odorata E Palolo ridge Preserved_Specimen Christophersen, E. 1431 Oahu BISH 1931-01-16
207 Psydrax odorata Kalapana Preserved_Specimen Gerhing, P. 3006 Hawaii BISH 1966-01-25
208 Psydrax odorata Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2425 Kauai BISH 1895-09-10
209 Psydrax odorata Halemanu Preserved_Specimen Rock, J.F.C. 2046 Kauai BISH 1909-02-14
210 Psydrax odorata N Kona, Mamaloha Hwy., near mile 28, Kaupulehu Preserved_Specimen Flynn, T.W. 1295 Hawaii BISH 1985-10-16
211 Psydrax odorata Kaena, Uluhulu Gulch Preserved_Specimen St.John, H. 10004 Oahu BISH 1929-11-03
212 Psydrax odorata Fenced area in Ka'upulehu Forest Reserve; W. of Ka'upulehu lava flow, N of road Preserved_Specimen Wagner, W.L. 4897 Hawaii BISH 1983-07-28
213 Psydrax odorata Maloaa Gulch Preserved_Specimen Forbes, C.N. 549 Kauai BISH 1916-10-09
214 Psydrax odorata N. slope of Hualalai, near flow of 1801 Preserved_Specimen Krajina, V.J. 611230037 Hawaii BISH 1961-12-30
215 Psydrax odorata Mahana, W. Molokai Preserved_Specimen Spaulding III, P. 1982.439 Molokai BISH 1982-08-20
216 Psydrax odorata Kapua, Kona Preserved_Specimen Forbes, C.N. 349 Hawaii BISH 1911-07-24
217 Psydrax odorata West Molokai Preserved_Specimen Degener, O. 22133 Molokai BISH 1952-03-17
218 Psydrax odorata Lihue Preserved_Specimen Forbes, C.N. 491 Kauai BISH 1916-10-04
219 Psydrax odorata Kaala, road Preserved_Specimen Lamoureux, C.H. 2432 Oahu BISH 1963-10-20
220 Psydrax odorata Waimea Arboretum Preserved_Specimen Miller, J.S. 7811 Oahu BISH 1992-07-28
221 Psydrax odorata Waianae Mts. Puu Hapapa Preserved_Specimen Krajina, V.J. 6020401128 Oahu BISH 1962-04-01
222 Psydrax odorata Halawa Quarry trail to divide Preserved_Specimen Kerr, M. 18 Oahu BISH 1945-09-09
223 Psydrax odorata Kau Preserved_Specimen Giffard, W.M. 364 Hawaii BISH 1928-07-01
224 Psydrax odorata Kaholuamanu Preserved_Specimen Rock, J.F.C. 1926 Kauai BISH 1909-03-03
225 Psydrax odorata N-S Halawa Ridge, Koolau Mts Preserved_Specimen Ozaki, E.T. 71 Oahu BISH 1952-12-14
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