Description
Key Characters:
Growth Form:
Trees or shrubs.
Stems:
Roots:
Leaves:
Leaves simple.
Alternate.
Blade elliptic, oblong, obovate, or oblanceolate.
Apex acute to rounded, often emarginate.
Surfaces punctate or with secretory canals.
Margins entire to serrate or dentate.
Pinnately veined.
Petiolate.
Stipules absent.
Flowers:
Flowers few to numerous in axillary, fasciculate, umbellate, or glomerate inflorescences arising on short scaly branches, spurs, or woody knobs often below the leaves, bracteate, but bracteoles absent.
Flowers small, bisexual (perfect) or unisexual (and then plants usually dioecious).
Calyx 4–5(6)-lobed, usually deeply so, the lobes imbricate or valvate in bud, punctate or with secretory lines, margins usually papillose or ciliate, persistentpunctate or with secretory lines, margins usually papillose or ciliate, persistent.
Corolla 4–5(6)-lobed, the lobes nearly distinct or connate up to ½ their length, imbricate in bud, usually punctate or with secretory lines, margins usually ciliate.
Stamens inserted on corolla throat and filaments completely adnate to corolla tube, or inserted at base of lobes and filaments connate, forming a short tube that is partly distinct from corolla tube; anthers dithecal, introrse, reduced and sterile in pistillate flowers.
Ovary superior, 3–5(6)-carpellate, 1-celled; ovules few in a single row, ovary reduced and sterile or absent in staminate flowers; style very short or absent, terminal; stigma variously capitate-angled, conical, or sausage–shaped.
Fruit:
Fruit a subglobose drupe; endocarp crustaceous; coriaceous; or woody; the withered stigma persistent.
Seeds 1 per fruit; small; dark colored; subglobose; excavated at base.
Ploidy:
Habitat:
Elevation Range: