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Phyla nodiflora

(L.) Greene (1899)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Phyla

carpet grass, mat grass, phyla

Description

Key Characters:

Growth Form: Perennial herbs.

Stems: Branches many, creeping, rooting at distal nodes, minutely strigose.

Roots:

Leaves: Leaves simple. Opposite. Blades spatulate, 1-3 X 0.5-1.5 cm. Base cuneate. Blade papery, surfaces pubescent. Margins distally sharply serrate. Veins inconspicuously 4-paired. Subsessile. Stipules absent.

Flowers: Inflorescences cylindric to ovate capitula, 1-2.5 cm; peduncle 1-7 cm. Flowers perfect. Calyx membranous, 2-lipped, 4-lobed. Corolla pinkish purple or white, glabrous. Stamens 4, didynamous, inserted on distal part of corolla tube, included. Ovary superior, 2-locular; ovule 1 per locule; style short; stigma capitate.

Fruit: Capsules ca. 1.5 mm in diameter; splitting into 2 1-seeded cocci.

Ploidy: 2n = 36

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

  • Yes

Natural History

Statewide Status

Naturalized

Island Status

Kuaihelani (Midway Atoll) Naturalized
Kaua'i Potentially naturalizing
O'ahu Naturalized
Molokai Naturalized
Maui Naturalized
Hawai'i Potentially naturalizing

Dispersal Agents

Water/Ocean

Vegetative Reproduction

Pollinators

Specimens

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Bibliography

Name Published In: Pittonia 4: 46 (1899)

Other References

Wagner et al. 1990:1316 (noted as cult. and potential escape); Starr et al. 2002:26 (NEWNAT/Mi, WM); Staples & Herbst 2005:555 (KEY), 566 (DESCR); Kennedy et al. 2010:24 (O, Mo); Faccenda, Arthur, et al. 2025:217 (K)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Phyla nodiflora (L.) Greene MAUI: W. Maui, Wailuku District, Kahului. Growing in sandy soil on side of road with Cynodon, Cenchrus, Malva, Chamaesyce, Asystasia, Alternanthera, & Verbesina. PRESERVED_SPECIMEN Hank Oppenheimer H40137 Maui PTBG 4/23/2001
2 Phyla nodiflora (L.) Greene Kamiloloa. PRESERVED_SPECIMEN Hank Oppenheimer H70121 Molokai PTBG 7/14/2001
3 Phyla nodiflora (L.) Greene Next to a rock wall adjoining adjacent house lot, Kamoa State Historical Park. Along coastline. PRESERVED_SPECIMEN Carolyn Corn s.n. Hawaii PTBG 9/3/1985
4 Phyla nodiflora (L.) Greene KAUAI: Koloa District. Lawai Valley. National Tropical Botan ical Garden. Lawnweed around edge of lily pond in Big Valley . PRESERVED_SPECIMEN Tim Flynn 6635 Kauai PTBG 9/22/1999
5 Phyla nodiflora (L.) Greene Pupukea. PRESERVED_SPECIMEN Hank Oppenheimer H70920 Oahu PTBG 7/22/2009
6 Phyla nodiflora (L.) Greene kaelepulu pond, kailua, oahu. at the south end of the pond, devries property. Growing on an exposed mud flat. PRESERVED_SPECIMEN l. m. crago, c. mcguire 2005-235 WTU 11/18/2005
7 Phyla nodiflora (L.) Greene central maui, southern coast of isthmus, kealia pond national wildlife refuge PRESERVED_SPECIMEN c. imada 98-20 Maui US 10/6/1998
8 Phyla nodiflora (L.) Greene lawai valley. national tropical botanical garden, weed in lawn around edge of lilypond in big valley. PRESERVED_SPECIMEN t. w. flynn 6635 Kauai US 9/22/1999
9 Phyla nodiflora (L.) Greene east maui, wailuku district, at kanaha beach park Scattered in lawn. Associated vegetation: Indigofera suffruticosa, Cynodon dactylon. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011123-2 Maui BISH 11/23/2001
10 Phyla nodiflora (L.) Greene Sand Island PRESERVED_SPECIMEN Herbst, D.R. 6364 Midway Atoll BISH 6/29/1980
11 Phyla nodiflora (L.) Greene W Maui, Lahaina District, Launiupoko Beach Park Just above high tide on berm betw sand & lawn. Associated vegetation: Prosopis pallida, lawn. PRESERVED_SPECIMEN Starr, F. 000611-1 Maui BISH 6/11/2000
12 Phyla nodiflora (L.) Greene East Maui, Makawao District, Makawao PRESERVED_SPECIMEN Duvall, F. s.n. Maui BISH 7/18/2006
13 Phyla nodiflora (L.) Greene east maui, makawao district, makawao PRESERVED_SPECIMEN duvall, f. collector number: s.n. Maui BISH 7/18/2006
14 Phyla nodiflora (L.) Greene Central Maui, southern coast of isthmus, Kealia Pond National Wildlife Refuge Extensive creeper on dikes of impoundment basins adjacent to main office. Growing with Atriplex, Cynodon dactylon. PRESERVED_SPECIMEN Imada, C.T. 98-20 Maui BISH 10/6/1998
15 Phyla nodiflora (L.) Greene East Maui, Wailuku District, at Kanaha Beach Park Scattered in lawn. Associated vegetation: Indigofera suffruticosa, Cynodon dactylon. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011123-2 Maui BISH 11/23/2001
16 Phyla nodiflora (L.) Greene w maui, wailuku district, kahului PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40137 Oahu BISH 4/23/2001
17 Phyla nodiflora (L.) Greene W Maui, Wailuku District, Kahului Growing in sandy soil on side of rd w. Cynodon, Cenchrus, Malva, Chamaesyce, Asystasia, Alternanthera, & Verbesina. PRESERVED_SPECIMEN Oppenheimer, H.L. H40137 Maui BISH 4/23/2001
18 Phyla nodiflora (L.) Greene sand island, sector 5, near drainage ditch, along runway near water catchment A new naturalized record. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 990512-2 Midway Atoll BISH 5/12/1999
19 Phyla nodiflora (L.) Greene w maui, lahaina district, launiupoko beach park Just above high tide on berm betw sand & lawn. Associated vegetation: Prosopis pallida, lawn. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 000611-1 Maui BISH 6/11/2000
20 Phyla nodiflora (L.) Greene Kamoa State Historical Park, North Kona Growing along coastline next to a rock wall adjoining adjacent house lot. PRESERVED_SPECIMEN Corn, C. Collector Number: s.n. Hawaii BISH 9/3/1985
21 Phyla nodiflora (L.) Greene Sand Island, sector 5, near drainage ditch, along runway near water catchment A new naturalized record. PRESERVED_SPECIMEN Starr, F. 990512-2 Midway Atoll BISH 5/12/1999
22 Phyla nodiflora (L.) Greene Sand Island PreservedSpecimen Herbst, D.R. Takeuchi, W.N. Collector Number: 6364 Midway Atoll BISH Specimen 6/29/1980
23 Phyla nodiflora (L.) Greene manoa campus, u. of h PRESERVED_SPECIMEN herbst, d.r. collector number: 635 Oahu BISH 10/7/1967
24 Phyla nodiflora (L.) Greene Sand Island, sector 5, near drainage ditch, along runway near water catchment A new naturalized record. PreservedSpecimen Starr, F. Martz, K. Collector Number: 990512-2 Midway Atoll BISH Specimen 5/12/1999
25 Phyla nodiflora (L.) Greene Kamoa State Historical Park, North Kona Growing along coastline next to a rock wall adjoining adjacent house lot. PRESERVED_SPECIMEN Corn, C. s.n. Hawaii BISH 9/3/1985
26 Phyla nodiflora (L.) Greene kamoa state historical park, north kona Growing along coastline next to a rock wall adjoining adjacent house lot. PRESERVED_SPECIMEN corn, c. collector number: s.n. Hawaii BISH 9/3/1985
27 Phyla nodiflora (L.) Greene Koloa District. Lawai Valley. National Tropical Botanical Garden. Weed in lawn around edge of lilypond in Big Valley Mat-forming herb; stems prostrate, rooting at nodes. PRESERVED_SPECIMEN Flynn, T. 6635 Kauai BISH 9/22/1999
28 Phyla nodiflora (L.) Greene sand island PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6364 Midway Atoll BISH 6/29/1980
29 Phyla nodiflora (L.) Greene Kamiloloa Along roadside. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H70121 Molokai BISH 7/14/2001
30 Phyla nodiflora (L.) Greene kamiloloa PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h70121 Maro Reef BISH 7/14/2001
31 Phyla nodiflora (L.) Greene Kaelepulu Pond, Kailua, Oahu, Hawaii Growing on an exposed mud flat at the south end of the pond, Devries Property. PRESERVED_SPECIMEN Crago, L.M. 2005-235 Oahu BISH 11/18/2005
32 Phyla nodiflora (L.) Greene koloa district. lawai valley. national tropical botanical garden. weed in lawn around edge of lilypond in big valley rock cliffs PRESERVED_SPECIMEN flynn, t. collector number: 6635 Oahu BISH 9/22/1999
33 Phyla nodiflora (L.) Greene Kamiloloa Along roadside. PRESERVED_SPECIMEN Oppenheimer, H.L. H70121 Molokai BISH 7/14/2001
34 Phyla nodiflora (L.) Greene kaelepulu pond, kailua, oahu, hawaii Growing on an exposed mud flat at the south end of the pond, Devries Property. PRESERVED_SPECIMEN crago, l.m. mcguire, c. collector number: 2005-235 Oahu BISH 11/18/2005
35 Phyla nodiflora (L.) Greene Ko'olauloa District, Pupukea. Possibly naturalized in yard area. PRESERVED_SPECIMEN Oppenheimer, H.L. H70920 Oahu BISH 7/22/2009
36 Phyla nodiflora (L.) Greene East Maui, Wailuku District, at Kanaha Beach Park Scattered in lawn. Associated vegetation: Indigofera suffruticosa, Cynodon dactylon. PRESERVED_SPECIMEN Starr, F. 011123-2 Maui BISH 11/23/2001
37 Phyla nodiflora (L.) Greene W Maui, Wailuku District, Kahului Growing in sandy soil on side of rd w. Cynodon, Cenchrus, Malva, Chamaesyce, Asystasia, Alternanthera, & Verbesina. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40137 Maui BISH 4/23/2001
38 Phyla nodiflora (L.) Greene Manoa Campus, U. of H used as ground cover PRESERVED_SPECIMEN Herbst, D.R. 635 Oahu BISH 10/7/1967
39 Phyla nodiflora (L.) Greene Honolulu, Lower Palolo, 10th Avenue. PRESERVED_SPECIMEN W. M. Bush 226 Oahu PTBG 6/2/1975
40 Phyla nodiflora (L.) Greene MAUI: W. Maui, Wailuku District, Kahului. Growing in sandy soil on side of road with Cynodon, Cenchrus, Malva, Chamaesyce, Asystasia, Alternanthera, & Verbesina. PRESERVED_SPECIMEN Hank Oppenheimer H40137 Maui PTBG 4/23/2001
41 Phyla nodiflora (L.) Greene Kamiloloa. PRESERVED_SPECIMEN Hank Oppenheimer H70121 Molokai PTBG 7/14/2001
42 Phyla nodiflora (L.) Greene Next to a rock wall adjoining adjacent house lot, Kamoa State Historical Park. Along coastline. PRESERVED_SPECIMEN Carolyn Corn s.n. Hawaii PTBG 9/3/1985
43 Phyla nodiflora (L.) Greene KAUAI: Koloa District. Lawai Valley. National Tropical Botan ical Garden. Lawnweed around edge of lily pond in Big Valley . PRESERVED_SPECIMEN Tim Flynn 6635 Kauai PTBG 9/22/1999
44 Phyla nodiflora (L.) Greene Pupukea. PRESERVED_SPECIMEN Hank Oppenheimer H70920 Oahu PTBG 7/22/2009
45 Phyla nodiflora (L.) Greene kaelepulu pond, kailua, oahu. at the south end of the pond, devries property. Growing on an exposed mud flat. PRESERVED_SPECIMEN l. m. crago, c. mcguire 2005-235 WTU 11/18/2005
46 Phyla nodiflora (L.) Greene central maui, southern coast of isthmus, kealia pond national wildlife refuge PRESERVED_SPECIMEN c. imada 98-20 Maui US 10/6/1998
47 Phyla nodiflora (L.) Greene lawai valley. national tropical botanical garden, weed in lawn around edge of lilypond in big valley. PRESERVED_SPECIMEN t. w. flynn 6635 Kauai US 9/22/1999
48 Phyla nodiflora (L.) Greene east maui, wailuku district, at kanaha beach park Scattered in lawn. Associated vegetation: Indigofera suffruticosa, Cynodon dactylon. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011123-2 Maui BISH 11/23/2001
49 Phyla nodiflora (L.) Greene Sand Island PRESERVED_SPECIMEN Herbst, D.R. 6364 Midway Atoll BISH 6/29/1980
50 Phyla nodiflora (L.) Greene W Maui, Lahaina District, Launiupoko Beach Park Just above high tide on berm betw sand & lawn. Associated vegetation: Prosopis pallida, lawn. PRESERVED_SPECIMEN Starr, F. 000611-1 Maui BISH 6/11/2000
51 Phyla nodiflora (L.) Greene East Maui, Makawao District, Makawao PRESERVED_SPECIMEN Duvall, F. s.n. Maui BISH 7/18/2006
52 Phyla nodiflora (L.) Greene east maui, makawao district, makawao PRESERVED_SPECIMEN duvall, f. collector number: s.n. Maui BISH 7/18/2006
53 Phyla nodiflora (L.) Greene Central Maui, southern coast of isthmus, Kealia Pond National Wildlife Refuge Extensive creeper on dikes of impoundment basins adjacent to main office. Growing with Atriplex, Cynodon dactylon. PRESERVED_SPECIMEN Imada, C.T. 98-20 Maui BISH 10/6/1998
54 Phyla nodiflora (L.) Greene East Maui, Wailuku District, at Kanaha Beach Park Scattered in lawn. Associated vegetation: Indigofera suffruticosa, Cynodon dactylon. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011123-2 Maui BISH 11/23/2001
55 Phyla nodiflora (L.) Greene w maui, wailuku district, kahului PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40137 Oahu BISH 4/23/2001
56 Phyla nodiflora (L.) Greene W Maui, Wailuku District, Kahului Growing in sandy soil on side of rd w. Cynodon, Cenchrus, Malva, Chamaesyce, Asystasia, Alternanthera, & Verbesina. PRESERVED_SPECIMEN Oppenheimer, H.L. H40137 Maui BISH 4/23/2001
57 Phyla nodiflora (L.) Greene sand island, sector 5, near drainage ditch, along runway near water catchment A new naturalized record. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 990512-2 Midway Atoll BISH 5/12/1999
58 Phyla nodiflora (L.) Greene w maui, lahaina district, launiupoko beach park Just above high tide on berm betw sand & lawn. Associated vegetation: Prosopis pallida, lawn. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 000611-1 Maui BISH 6/11/2000
59 Phyla nodiflora (L.) Greene Kamoa State Historical Park, North Kona Growing along coastline next to a rock wall adjoining adjacent house lot. PRESERVED_SPECIMEN Corn, C. Collector Number: s.n. Hawaii BISH 9/3/1985
60 Phyla nodiflora (L.) Greene Sand Island, sector 5, near drainage ditch, along runway near water catchment A new naturalized record. PRESERVED_SPECIMEN Starr, F. 990512-2 Midway Atoll BISH 5/12/1999
61 Phyla nodiflora (L.) Greene Sand Island PreservedSpecimen Herbst, D.R. Takeuchi, W.N. Collector Number: 6364 Midway Atoll BISH Specimen 6/29/1980
62 Phyla nodiflora (L.) Greene manoa campus, u. of h PRESERVED_SPECIMEN herbst, d.r. collector number: 635 Oahu BISH 10/7/1967
63 Phyla nodiflora (L.) Greene Sand Island, sector 5, near drainage ditch, along runway near water catchment A new naturalized record. PreservedSpecimen Starr, F. Martz, K. Collector Number: 990512-2 Midway Atoll BISH Specimen 5/12/1999
64 Phyla nodiflora (L.) Greene Kamoa State Historical Park, North Kona Growing along coastline next to a rock wall adjoining adjacent house lot. PRESERVED_SPECIMEN Corn, C. s.n. Hawaii BISH 9/3/1985
65 Phyla nodiflora (L.) Greene kamoa state historical park, north kona Growing along coastline next to a rock wall adjoining adjacent house lot. PRESERVED_SPECIMEN corn, c. collector number: s.n. Hawaii BISH 9/3/1985
66 Phyla nodiflora (L.) Greene Koloa District. Lawai Valley. National Tropical Botanical Garden. Weed in lawn around edge of lilypond in Big Valley Mat-forming herb; stems prostrate, rooting at nodes. PRESERVED_SPECIMEN Flynn, T. 6635 Kauai BISH 9/22/1999
67 Phyla nodiflora (L.) Greene sand island PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6364 Midway Atoll BISH 6/29/1980
68 Phyla nodiflora (L.) Greene Kamiloloa Along roadside. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H70121 Molokai BISH 7/14/2001
69 Phyla nodiflora (L.) Greene kamiloloa PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h70121 Maro Reef BISH 7/14/2001
70 Phyla nodiflora (L.) Greene Kaelepulu Pond, Kailua, Oahu, Hawaii Growing on an exposed mud flat at the south end of the pond, Devries Property. PRESERVED_SPECIMEN Crago, L.M. 2005-235 Oahu BISH 11/18/2005
71 Phyla nodiflora (L.) Greene koloa district. lawai valley. national tropical botanical garden. weed in lawn around edge of lilypond in big valley rock cliffs PRESERVED_SPECIMEN flynn, t. collector number: 6635 Oahu BISH 9/22/1999
72 Phyla nodiflora (L.) Greene Kamiloloa Along roadside. PRESERVED_SPECIMEN Oppenheimer, H.L. H70121 Molokai BISH 7/14/2001
73 Phyla nodiflora (L.) Greene kaelepulu pond, kailua, oahu, hawaii Growing on an exposed mud flat at the south end of the pond, Devries Property. PRESERVED_SPECIMEN crago, l.m. mcguire, c. collector number: 2005-235 Oahu BISH 11/18/2005
74 Phyla nodiflora (L.) Greene Ko'olauloa District, Pupukea. Possibly naturalized in yard area. PRESERVED_SPECIMEN Oppenheimer, H.L. H70920 Oahu BISH 7/22/2009
75 Phyla nodiflora (L.) Greene East Maui, Wailuku District, at Kanaha Beach Park Scattered in lawn. Associated vegetation: Indigofera suffruticosa, Cynodon dactylon. PRESERVED_SPECIMEN Starr, F. 011123-2 Maui BISH 11/23/2001
76 Phyla nodiflora (L.) Greene W Maui, Wailuku District, Kahului Growing in sandy soil on side of rd w. Cynodon, Cenchrus, Malva, Chamaesyce, Asystasia, Alternanthera, & Verbesina. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40137 Maui BISH 4/23/2001
77 Phyla nodiflora (L.) Greene Manoa Campus, U. of H used as ground cover PRESERVED_SPECIMEN Herbst, D.R. 635 Oahu BISH 10/7/1967
78 Phyla nodiflora (L.) Greene Honolulu, Lower Palolo, 10th Avenue. PRESERVED_SPECIMEN W. M. Bush 226 Oahu PTBG 6/2/1975
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