Santalum haleakalae

Hillebr. (1888)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Santalopsida  Order: Santalales Family: Santalaceae Genus: Santalum

‘iliahi [iliahi], sandalwood

Description

Key Characters:

Growth Form: Small trees, evergreen, single or multiple stems; 1–8 m tall. Apparently hemiparasitic on roots of other plants.

Stems: Older trees readily spread vegetatively via root sprouts when surface roots are exposed or damaged, sometimes forming small stands.

Roots:

Leaves: Leaves simple. Opposite. Blades ovate, obovate, elliptic or orbicular, 2.2–8.4 cm x 1.2–4.9 cm. Apex acute or obtuse (rarely acuminate). Base cuneate. More glaucous and paler on lower surfaces, evergreen, simple, olive green, often purplish–tinged, usually glaucous, stiff and coriaceous. Margins flat to slightly recurved. Petioles 1.2–21.7 mm long. Stipules absent.

Flowers: Flowers in congested, terminal, compound cymes, rarely with a few additional axillary flower clusters. Flowers bisexual (perfect), weakly fragrant, reddish to cream-colored in bud, longer than wide. Pedicels 2–4 mm long, bracts persistent until flowering. Calyx (sepals) absent. Corolla of 4 (rarely 5) petals; petals deep pink to red throughout, or with white to pink interiors; floral–tubes campanulate to cylindrical, without vertical keel below each corolla lobe; disk lobes lingulate, apices acute or round, concave with thickened margins, papillate. Stamens 4, hairs proximal to the filaments (rarely hairs distal). Ovary partly inferior (approx. ⅔); drupe elliptic, with subapical ring; stigma lobes 3.

Fruit: Drupes black or purplish black at maturity; 9–16 mm long; with a subapical receptacular ring. Seeds muricate; with pointed beak on one end.

Ploidy:

Habitat: In subalpine and montane mesic forest plant communities and exposed or semishaded scrublands.

Elevation Range: 800–2,700 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Maui Endemic

Dispersal Agents


Pollinators

Specimens

Bibliography

Name Published In: Fl. Hawaiian Isl.: 390 (1888)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Santalum haleakalae Hillebr. East Maui, Hana District, Kahikinui, south of Kahua, west of upper Kamaole drainage basin. Vegetation: Metrosideros, Acacia, Dodonaea, Leptecophylla, Myrsine, Ilex, Coprosma, Dicranopteris, Diplazium, Cibotium, Smilax, Astelia, Dryopteris, Sadleria, Pteris, Coniogramme, Cyrtomium, Peperomaia, Lobelia, Melicope, Carex, Pipturus, Urera, Perrottetia, Odontosoria, Asplenium, Descampsia, Hypolepis, Clermontia, Morelotia, Pennisetum, Holcus, Anthoxanthum, Senecio, Rubus, Axonopus, Verbena, Adiantum. PRESERVED_SPECIMEN Oppenheimer, H.L. H21605 Maui BISH 2/4/2016
2 Santalum haleakalae Hillebr. East Maui, Makawao District, Koolau Gap. Vegetation: Metrosideros, Cheirodendron, Broussaisia, Myrsine, Vaccinium, Dubautia, Perrottetia, Coprosma, Leptecophylla, Rubus, Sadleria, Diplazium, Sthyrium, Dryopteris, Pipturus, Uncinia, Carex, Peperomia, Pneumatopteris, Asplenium, Elaphoglossum, Coniogramme, Melicope, Lysimachia, Pteris, Pseudophegopteris. PRESERVED_SPECIMEN Oppenheimer, H.L. H40936 Maui BISH 4/30/2009
3 Santalum haleakalae Hillebr. East Maui, Hana District, Kahikinui, upper Kepuni drainage basin. Vegetation: Metrosideros, Dodonaea, Leptecophylla, Coprosma, Dryopteris, Holcus, Deschampsia, Anthoxanthum, Pteris, Pellaea, Hypocaeris. PRESERVED_SPECIMEN Oppenheimer, H.L. H91315 Maui BISH 9/17/2013
4 Santalum haleakalae Hillebr. East Maui, Makawao District, Kanaio ahu. On cinder/ash substrate. Vegetation: Sophora, Dodonaea, Rubus, Osteomeles, Dryopteris, Dubautia, Bidens, Carex, Luzula, Pellaea, Asplenium, Polypodium, Cirisum, Lythrum, Senecio, Holcus, Hypochoeris, Bidens pilosa. PRESERVED_SPECIMEN Oppenheimer, H.L. H40928 Maui BISH 4/22/2009
5 Santalum haleakalae Hillebr. Growing on talus slope PRESERVED_SPECIMEN s. carlquist 548 Kauai BISH
6 Santalum haleakalae Hillebr. West Maui, Lahaina District, Manawainui Gulch, ridge between west and east forks. Vegetation: Metrosideros, Acacia, Diospyros, Nestegis, Dodonaea, Myrsine, Leptecophylla, Wikstroemia, Osteomeles, Pouteria, Bidens, Eragrostis, Pteridium, cocculus, Asplenium, Asplenium, Panicum, Casuarina, Grevillea, Psidium, Erigeron, Senecio, Andropogon, Setaria. PRESERVED_SPECIMEN Oppenheimer, H.L. H101311 Maui BISH 10/17/2013
7 Santalum haleakalae Hillebr. East Maui, Hana District, East Wailua Iki. Emergent in small intermittent gully in Subalpine Shrubland. Vegetation: Sophora, Metrosideros, Leptecophylla, Coprosma, Dodonaea, Vaccinium, Dubautia, Myrsine, Deschampsia, Uncinia, Carex, Morelotia, Fragaria, Rubus, Pteris, Dryopteris, Polystichum, Pteris, Pteridium, Asplenium, Elaphoglossum, Astelia, Holcus, Anthoxanthum, Rumux, Hypochaeris. PRESERVED_SPECIMEN Oppenheimer, H.L. H41304 Maui BISH 4/9/2013
8 Santalum haleakalae Hillebr. E. Maui, Kula Forest Reserve. Waiohuli. Degraded subalpine shrubland. PRESERVED_SPECIMEN Hank Oppenheimer H21003 Maui PTBG 2/16/2010
9 Santalum haleakalae Hillebr. haleakala, e. maui PRESERVED_SPECIMEN h. mann & w. t. brigham 396 MO
10 Santalum haleakalae Hillebr. haleakala. below the start of the halemau trail. PRESERVED_SPECIMEN r. l. wilbur and g. l. webster 1021 NY 7/5/1948
11 Santalum haleakalae Hillebr. E. Maui, Waikamoi Preserve; Ainahou Bowl, upper Keanae, Koolau forest Reserve. Metrosideros-Cheirodendron. PRESERVED_SPECIMEN K. R. Wood 10178 Maui PTBG 5/22/2003
12 Santalum haleakalae Hillebr. sandwich islands, maui moist bank, gulch bottom PRESERVED_SPECIMEN wilkes expedition TENN
13 Santalum haleakalae Hillebr. Koolau Gap, Ainahou, Keanae valley, west side of gap, 5560 ft. elev. north facing slope , with Metrosideros, Styphelia, Sadleria cyathioides, Rubus hawaiiensis, Dubautia dolosa, Dubautia linearis, Cheirodendron, Coprosma, Holchus, Vaccinium, Lysimachia hillebrandii, Ageratina adenophora, Lythrum, PRESERVED_SPECIMEN Steve Perlman 16027 Maui PTBG 8/1/2014
14 Santalum haleakalae Hillebr. West Maui, Lahaina District, Kahana, ridge separating Kahana valley and Mailepai Gulch to the north. Vegetation: Metrosideros, Dicranopteris, Leptecophylla, Wikstroemia, Macharina. PRESERVED_SPECIMEN Oppenheimer, H.L. H81101 Maui BISH 8/19/2011
15 Santalum haleakalae Hillebr. PRESERVED_SPECIMEN leila m. shultz, john s. shultz 6394 USU
16 Santalum haleakalae Hillebr. PRESERVED_SPECIMEN leila m. shultz, john s. shultz 6449 USU
17 Santalum haleakalae Hillebr. E. Maui, Kula Forest Reserve, along Skyline Road, south of Puu Keokea. Sophora chrysophylla sub-alpine shrub. PRESERVED_SPECIMEN K. R. Wood 13704 Maui PTBG 4/22/2009
18 Santalum haleakalae Hillebr. Hawaiian Islands; Maui. Crater of Haleakala. PRESERVED_SPECIMEN C. N. Forbes m-301 SFV
19 Santalum haleakalae Hillebr. Maui Island, haleakala highway PRESERVED_SPECIMEN Vladimir J. Krajina 620415014 UBC 4/15/1962
20 Santalum haleakalae Hillebr. off the road going to the summit of Haleakala. PRESERVED_SPECIMEN Lani Stemmermann BISH 6/21/1975