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Psidium cattleyanum

Sabine (1821)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Myrtales Family: Myrtaceae Genus: Psidium

strawberry guava, waiawī, waiawī ‘ula‘ula [waiawi ulaula]

Description

Key Characters:

Growth Form: Shrubs or small trees 2–6 m tall, bark usually reddish, peeling.

Stems: Young branches terete (cylindrical), sparsely puberulent when young, soon glabrate.

Roots:

Leaves: Leaves simple. Opposite. Blades obovate to elliptic–obovate, 3.5–13.5 cm long, (2.5–)3–6 cm wide. Apex bluntly cuspidate. Base attenuate to cuneate. Surfaces glabrous or glabrate; aromatic, leathery; upper surfaces glossy dark green; lower surfaces paler. Margins entire. Pinnately veined, both surfaces with scarcely raised lateral veins. Petioles 5–10 mm long. Stipules vestigial or absent.

Flowers: Flowers usually solitary in the leaf axils, peduncles 4–8 mm long. Flowers bisexual (perfect), actinomorphic. Hypanthium well–developed above ovary, splitting when flower opens. Calyx of 4–5 sepals; sepals ca.4–5 mm long, persistent in fruit. Corolla of 4–5 petals, white, obovate, ca. 5–7 mm long. Stamens white, numerous, distinct; anthers dithecal. Ovary inferior, (2)3–4(–7)-celled; ovules numerous, placentation axile; style terminal, simple.

Fruit: Berries red to purplish red or occasionally yellow; globose to obovoid or ellipsoid; glossy and smooth; 2–3 cm in diameter; the pulp white. Seeds numerous; ca. 5 mm long; smooth.

Ploidy: 2n = 88

Habitat: Disturbed mesic forest and wet forest.

Elevation Range: 15–1,220 m.

Historical Distribution

Images

Synonyms (38)

Episyzygium oahuense Suess. & A.Ludw. (1950)
Eugenia ferruginea Sieber ex C.Presl (1828)
Eugenia oxygona Koidz. (1918)
Eugenia pseudovenosa H.Perrier (1953)
Eugenia urceolata Cordem. (1895)
Guajava arborea (Vell.) Kuntze (1891)
Guajava buxifolia (Nutt.) Kuntze (1891)
Guajava cattleyana (Sabine) Kuntze (1891)
Guajava humilis (Vell.) Kuntze (1891)
Guajava obovata (Mart. ex DC.) Kuntze (1891)
Psidium arboreum Vell. (1829)
Psidium buxifolium Nutt. (1842)
Psidium cattleianum Afzel. ex Sabine
Psidium cattleianum f. cattleianum
Psidium cattleianum f. lucidum O.Deg. (1939)
Psidium cattleianum var. littorale (Raddi) Fosberg (1962)
Psidium cattleyanum f. lucidum O.Deg. (1939)
Psidium cattleyanum var. cattleyanum
Psidium cattleyanum var. coriaceum Kiaersk. (1893)
Psidium cattleyanum var. littorale (Raddi) Fosberg (1962)
Psidium cattleyanum var. purpureum Mattos (2007)
Psidium cattleyanum var. pyriformis Mattos (1981)
Psidium chinense Lodd. ex Chardonier (1880)
Psidium coriaceum O.Berg (1856)
Psidium coriaceum var. grandifolium O.Berg (1857)
Psidium coriaceum var. longipes O.Berg (1857)
Psidium coriaceum var. obovatum O.Berg (1857)
Psidium ferrugineum C.Presl (1828)
Psidium gaudichaudianum Proença & Faria (2017)
Psidium humile Vell. (1829)
Psidium indicum Bojer (1837)
Psidium littorale Raddi (1821)
Psidium littorale var. longipes (O.Berg) Fosberg (1941)
Psidium littorale var. lucidum (O.Deg.) Fosberg (1941)
Psidium obovatum Mart. ex DC. (1828)
Psidium sellowianum O.Berg (1856)
Psidium ubatubense Mattos (1967)
Psidium variabile O.Berg (1856)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents

Birds

Vegetative Reproduction

Pollinators

Bibliography

Name Published In: Trans. Hort. Soc. London 4: 317 (1821)

Other References

Wagner et al. 1990:971 (K, O, Mo, L, M, H [as P. cattleianum]); Staples & Herbst 2005:428 (DESCR); Tuler et al. 2018:1196 (spelling change to cattleyanum)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Psidium cattleyanum Koolau Range, 47-285 Pulama Rd., Kaneohe, Senator Fong's Plantation and Garden with Schefflera and Schinus; lowland mesic forest Preserved_Specimen tree to 4 m; bark smooth and red; fruit red Annable, C.R. 3034 Oahu BISH 1996-09-25
2 Psidium cattleyanum Keauhou Ranch, Kau dist Several plants scattered for 50 ft. in pasture with remnant forest scrub trees. Preserved_Specimen Ht. 2', spread 3'. Fruit green. Flowers cream white and fragrant. Davis, J. 413 Hawaii BISH 1980-10-23
3 Psidium cattleyanum Koloa District: Kalaheo, Kalawai Park; secondary vegetation around park with Ficus microcarpa, Syzygium cumini, Psidium, and Schinus. Preserved_Specimen Lorence, D.H. 5567 Kauai BISH 1987-09-26
4 Psidium cattleyanum Honolulu, Lanipo Trail Common along trail. Preserved_Specimen Tree l7 ft ht, in few fruits and flowers seen. Kennedy, B. 18 Oahu BISH 1995-11-21
5 Psidium cattleyanum Lanaihale, N end Munro Trail, 3.4 mi in Growing as understory in planted Eucalyptus forest off jeep road. Preserved_Specimen Tree to 3 meters tall. Leaves glossy, glabrous. Fruit to 4 cm diam., yellow when mature. Spence, G. 310 Lanai BISH 1973-10-18
6 Psidium cattleyanum Kamokumui gulch, Kamananui, Waianae Rang dry forest Preserved_Specimen Stone, B.C. 893 Oahu BISH 1955-07-17
7 Psidium cattleyanum NW Part; Mokuleia Forest Reserve; Peacock Flats Medtrosideros forest. Preserved_Specimen Small tree. Fruit dark-reddish purple, edible. Little Jr., E.L. 31037 Oahu BISH 1976-08-10
8 Psidium cattleyanum Mahana, Hulopoe Gulch, below native rain forest. Taking over gulch below native rain forest. Preserved_Specimen Fruit yellow. Degener, O. 28613 Lanai BISH 1963-08-13
9 Psidium cattleyanum Wailua Game Reserve, above Wailua Stockpile of topsoil. Preserved_Specimen Invasive shrub, to 6 ft, leaves dark green shining above, paler below; fruit bright red. Howard, R.A. 20172 Kauai BISH 1985-10-20
10 Psidium cattleyanum Waipio-Waiawa, S ridge of Kipapa Gulch lower rain forest Preserved_Specimen 8' tree Ozaki, E.T. 50 Oahu BISH 1952-11-23
11 Psidium cattleyanum 6.7 miles from junction of Stainback w/Hwy 11 (Unit G) In dense clumps along highway; Found associated w/Metrosideros, Cibotium, Melastoma and grasses. Preserved_Specimen Small tree to 20 feet w/dbh 1 & 1/2 inches. Fruits yellow. Ishikawa, S. 224 Hawaii BISH 1973-01-13
12 Psidium cattleyanum ahupuaa in the Puna District of Hawaii The plants occur from areas exposed to direct sunlight to those areas that are moderately shaded. The soil is Aa with some organic material. Preserved_Specimen Yoshida, L. Hawaii BISH 1979-05-10
13 Psidium cattleyanum Laie Trail lower edge of forest Preserved_Specimen 3 m shrub Egler, F.E. 37302 Oahu BISH 1937-07-24
14 Psidium cattleyanum Drive up Kulani Prison Road about 7.4 miles toward Kulani Prison from intersection with Kona-Hilo Highway near Hilo; or up Kulani Prison Road 2.4 miles toward Kulani Prison from intersection with Volcano Road (shortcut to Kulani Prison from HVNP) Metrosideros - Cibotium - Pritchardia forest community Preserved_Specimen Tree 5-10 meters tall Newell, C.L. 325 Hawaii BISH 1966-08-09
15 Psidium cattleyanum Honolulu Distr., Wiliwilinui Trail, alongside open dirt track soon after leaving paved road portion of hike. Disturbed habitat with Casuarina equisetifolia and Acacia confusa overstory along with Psidium cattleianum and Schinus terebinthifolia. Preserved_Specimen Trees 8-10ft height, fruits green Kennedy, B. 152 Oahu BISH 2024-04-28
16 Psidium cattleyanum Kipapa Gulch, SE Ridge, Waipio Dist., Oahu Cultivated Preserved_Specimen 5 ft. tall Grant, L.G. 7091 Oahu BISH 1934-07-13
17 Psidium cattleyanum Hanalei District. Limahuli Valley Secondary forest along stream in valley bottom. Dominated by Psidium, Aleurites and Schefflera with scattered Metrosideros, Pisonia and Hedyotis. Preserved_Specimen Flynn, T.W. 2315 Kauai BISH 1987-08-20
18 Psidium cattleyanum Kipapa Trail, several meters beyond entrance, N side of trail disturbed wet lowland forest; Metrosideros, Acacia, Dicranopteris Preserved_Specimen 7-8 m tree; bark smooth, light reddish brown Reynolds, L.S. 58 Oahu BISH 2006-05-30
19 Psidium cattleyanum Kalaupapa National Historical Park trail switchback 3 M:xt,nt(ns-xg-xs) Preserved_Specimen medium sized tree,common to dominant in some areas Wysong, M.L. 790 Molokai BISH 2005-08-13
20 Psidium cattleyanum Located down Chain of Craters Road about 28.5 miles toward Kalapana from intersection with Crater Rim Road; or 0.2 miles before Queens' Bath. Plant was about .25 miles up the Royal Estates Road, not far from HVNP boundary. (plot 27) Mixed lowland scrub community. Preserved_Specimen Newell, C.L. 120 Hawaii BISH 1966-07-20
21 Psidium cattleyanum Oahu Forest NWR, Koa ridge, above Mililani Mauka. Site of October 30, 2023 fire. On ridge, edge of burned area, full sun, moist, formerly dominated by uluhe. Associated; Miconia crenata, Acacia koa, Trema orientale, Falcataria falcata, Dianella sandwicensis, Budleja asiatica, Cecropia obtusifolia, Rubus rosifolius. Preserved_Specimen Many plants with pretty well established Tectococcus. Arthur, K. 49 Oahu BISH 2025-02-10
22 Psidium cattleyanum Molokai Forest Reserve. At junction of Waikolu Valley Lookout road and Puukauwa Trail Growing in planted Eucalyptus grove w/undergrowth of fern and grasses. Preserved_Specimen Tree 20 ft. tall; dbh 3 inches. Leaves opposite; obovate; glossy green on top, lighter below. Fruits small- 2.5 cm diam.-red when mature. Spence, G. 59 Molokai BISH 1972-08-29
23 Psidium cattleyanum Kahuku clay loam Preserved_Specimen 10-20' tree; light bark; dark green leaves Weber, L. Oahu BISH 1977-03-26
24 Psidium cattleyanum Olaa; Olaa Forest Reserve, Site 08 OS Ohia-treefern dieback. Soil a deep and moderately well drained ash. Primary vegetation: Metrosideros polymorpha, Coprosma spp., Ilex anomala, Gouldia sp.& Psychotria hawaiiensis. Preserved_Specimen Adee, K.T. Hawaii BISH
25 Psidium cattleyanum Drive up Kulani Prison Road about 7.4 miles toward Kulani Prison from intersection with Kona-Hilo Highway near Hilo; or up Kulani Prison Road 2.4 miles toward Kulani Prison from intersection with Volcano Road (shortcut to Kulani Prison from HVNP) Metrosideros - Cibotium - Pritchardia forest community Preserved_Specimen Tree about 5-10 meters tall Newell, C.L. 326 Hawaii BISH 1966-08-09
26 Psidium cattleyanum Niu along the W Niu Stream Preserved_Specimen Egler, F.E. 37140 Oahu BISH 1937-06-25
27 Psidium cattleyanum Located down Chain of Craters Road about 28.5 miles toward Kalapana from intersection with Crater Rim Road; or 0.2 miles before Queens Bath. Plant was about 0.25 miles up the Royal Estates Road, not far from HVNP boundary. (plot 27) Mixed lowland scrub community Preserved_Specimen Shrub about 1 meter tall Newell, C.L. 122 Hawaii BISH 1966-07-20
28 Psidium cattleyanum Puna; Wahaula Hot, shrubby, coastal plain. Preserved_Specimen Degener, O. 31542 Hawaii BISH 1966-11-25
29 Psidium cattleyanum Laie sandy Preserved_Specimen flower white Kinikini, T.V. 34 Oahu BISH 1967-04-08
30 Psidium cattleyanum Hawaii Volcanoes National Park drive down Kona-Hilo Highway about 4 miles past Hongo Store towards Hilo. Turn right on Fern Estates Road and drive about 1.1 miles. Plant was about 25 meters east (left side) of road. (plot 56) Closed Metrosideros - Cibotium forest community. Preserved_Specimen Tree. Newell, C.L. 613 Hawaii BISH 1966-08-28
31 Psidium cattleyanum Keauhou Ranch, Kau District Several plants scattered for 50 ft. in pasture with remnant forest scrub trees. Preserved_Specimen Height 2', spread 3'. Fruit green, flowers cream white and fragrant. Davis, J. 413 Hawaii BISH 1980-10-23
32 Psidium cattleyanum Near Hilo, lava flow of 1881 a`a lava. Preserved_Specimen Small tree, 4 m. high. MacDaniels, L.H. 232 Hawaii BISH 1926-11-12
33 Psidium cattleyanum Along Highway 132, 2 miles east of Pahoa Found associated w/Metrosideros, Albizia and ferns. Preserved_Specimen Slender tree 15 ft in height, dbh 2 inches. Fruits globose, small red outside, w/white pulp. Ishikawa, S. 216 Hawaii BISH 1973-01-12
34 Psidium cattleyanum Limahuli, Haena wet gulch on bottom near pahoi, ditch and stream Preserved_Specimen tree, 4-8m tall, flowers white-yellow, no fruits. Bark smooth, mottled. Earle, E. 20 Kauai BISH 1972-06-04
35 Psidium cattleyanum Laie volcanic clay Preserved_Specimen fruit reddish; flower small, white Kamae, K.K. 46 Oahu BISH 1967-05-30
36 Psidium cattleyanum Keaiwa Heiau State Rec. Area, 10 mi NW of Honolulu, Aiea Loop Trail, ~3 miles from beginning in dense stands along with Melaleuca, Tristania Preserved_Specimen Fruit yellow, ovoid; large, spreading tree Ishikawa, S. 116 Oahu BISH 1972-06-22
37 Psidium cattleyanum East Maui, Ulupalakua, adjacent to Tedeschi Winery, NW of Puu Mahoe Open mesic slope of Grevillea robusta, Schinus terebinthifolius, Psidium guajava; groundcover dominated by Bryophyllum pinnatum. Preserved_Specimen Tree 15 ft. tall, bark smooth, mottled brown; fruit yellow, oblong. Imada, C. 200226 Maui BISH 2002-07-16
38 Psidium cattleyanum ahupuaa of Halepuaa in the Puna District The plants occur from areas exposed to direct sunlight to those areas that are moderately shaded. The soil is `a`a with some organic material. Preserved_Specimen Shrub or tree. Yoshida, L. Hawaii BISH 1979-05-10
39 Psidium cattleyanum Punaluu thicket Preserved_Specimen small tree; fruit red St.John, H. 10585 Oahu BISH 1930-09-28
40 Psidium cattleyanum Kona District: Koolau Mts., Honolulu Forest Reserve, north of Honolulu. Waahila (St. Louis Heights) Trail, running northeast along Waahila Ridge from Waahila State Park Dry open forest. Preserved_Specimen Shrub, 3 m.; fruit maroon-purple. Lammers, T.G. 5348 Oahu BISH 1983-08-28
41 Psidium cattleyanum Laie Hills clay Preserved_Specimen Shrubby tree, smooth leaves, fruit red Ziegelbauer, K. 33 Oahu BISH 1975-03-13
42 Psidium cattleyanum Lanai City, mauka Naturalized. Preserved_Specimen Fruit red. Degener, O. 28663 Lanai BISH 1963-07-25
43 Psidium cattleyanum Waimea State Park, Kauai. Along Hwy 55, one mile from Waimea Canyon Lookout Growing in assos. with Koa-Eucalyptus in scattered clumbs along Hwy. Preserved_Specimen Tree 15ft tall; dbh 2 inches. Fruits small and red when mature, 2.5 cm dia. Leaves opposite, obovate, glossy green on top, lighter below. Spence, G. 97 Kauai BISH 1972-11-07
44 Psidium cattleyanum Lyon Arboretum, Middle Lawn Makai, next to the road between Cottages A and B Preserved_Specimen ~4' shrub; red fruit Baker, R.F. 97 Oahu BISH 1975-04-26
45 Psidium cattleyanum Waimea State Park, Kauai. Along Hwy 55, 1/2 mile from Waimea Canyon Lookout growing with Psidium cattleianum in clumps of a Koa forest Preserved_Specimen Tree 15ft tall; dbh 3 inches. Fruits yellow when mature, 3 cm dia. Leaves opposite; glabrous; dark green on top, lighter below. Spence, G. 99 Kauai BISH 1971-11-07
46 Psidium cattleyanum Piihonua; Hilo Forest Reserve, Site 38 OS Ohia-koa dieback. Soil a somewhat poorly drained ash found over an unknown substrate. Primary vegetation: Acacia koa, Metrosideros polymorpha, Psidium cattleianum, Cheirodendron trigynum, Antidesma platyphyllum & Vaccinium calycinum. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-04-19
47 Psidium cattleyanum Limahuli, Haena wet gulch on bottom near pahoi, ditch and stream Preserved_Specimen tree, 4-8m tall, flowers white-yellow, no fruits. Bark smooth, mottled. Earle, E. 20 Kauai BISH 1972-06-04
48 Psidium cattleyanum Olaa; Olaa Forest Reserve, Site 08 OS Ohia-treefern dieback. Soil a deep and moderately well drained ash. Primary vegetation: Metrosideros polymorpha, Coprosma spp., Ilex anomala, Gouldia sp.& Psychotria hawaiiensis. Preserved_Specimen Adee, K.T. Hawaii BISH
49 Psidium cattleyanum East Maui, along Hana Hwy., 1 mi past Kailua with Melaleuca, Eucalyptus, Cocos, Zingiber, Nephrolepis and Dicranopteris understory Preserved_Specimen 10' tree. Leaves obovate, dark green. Fruit red outside, white pulp, ~1 diam. Ishikawa, S. 156 Maui BISH 1972-10-14
50 Psidium cattleyanum Manoa, E ridge to Olympus, Koolau Mts Preserved_Specimen Hume, E.P. 414 Oahu BISH 1931-12-10
51 Psidium cattleyanum near Kahili Swamp, Wahiawa, Lihue-Koloa Forest Reserve woods Preserved_Specimen St.John, H. 10857 Kauai BISH 1930-12-29
52 Psidium cattleyanum Waipio, SE ridge of Kipapa Gulch Preserved_Specimen 5 ft. Grant, M.L. 7091 Oahu BISH 1934-07-13
53 Psidium cattleyanum Kapalama Valley Preserved_Specimen Shrub (naturalized). Christophersen, E. 1630 Oahu BISH 1931-02-28
54 Psidium cattleyanum Manoa Cliff Trail HUMAN_OBSERVATION Friday, J.B. Oahu 10/19/2019
55 Psidium cattleyanum Puna; Wahaula Preserved_Specimen Scrub. Fruit red. Degener, O. 31540 Hawaii BISH 1966-11-25
56 Psidium cattleyanum Aihualama, near swimming pool Preserved_Specimen Russell, P. 94 Oahu BISH 1932-07-19
57 Psidium cattleyanum Honolulu? Preserved_Specimen (Not on sheet) Oahu BISH
58 Psidium cattleyanum West Maui, Makanukaole Gulch Preserved_Specimen fruit purple Gagné, W.C. Maui BISH
59 Psidium cattleyanum Preserved_Specimen Tree roughly 2 meters tall with immature and mature fruit, along trailside Wolkis, D. Kauai BISH 2020-08-16
60 Psidium cattleyanum Waiakea Forest Reserve; 7 miles SW of State Route 11 on Mamki Road to Kulani Prison Preserved_Specimen Tree 4 m. tall; fruits green. Miller, J.S. 7936 Hawaii BISH 1992-08-06
61 Psidium cattleyanum Kilauea Preserved_Specimen Tree 20 feet tall, with bark like Platanus. Fagerlund, G.O. 407 Hawaii BISH 1943-02-24
62 Psidium cattleyanum Lyon Arboretum, Middle Lawn Makai, next to the road between Cottages A and B Preserved_Specimen ~4' shrub; red fruit Baker, R.F. 97 Oahu BISH 1975-04-26
63 Psidium cattleyanum Eastern part of island, Kapu Ranch Preserved_Specimen Shrub ca 4 m tall. Fl sepals green, petals white; numerous white stamens; style long and emerging from stamens; stigma green & capitate with irregular, angular margins. Frt green, maturing to cranberry-red outside, white inside; remnant calyx present on top of frt. Sees to 5 mm long, tan. Pell, S.K. 758 Molokai BISH 2008-07-23
64 Psidium cattleyanum Near Waipahee Falls Preserved_Specimen Tree 50ft, 1ft diam., forming thickets. Fruits yellow, edible. Little Jr., E.L. 31181 Kauai BISH 1976-09-02
65 Psidium cattleyanum Maunalaha Trail, Makiki Valley, Koolau Mts. Preserved_Specimen Ziegler, M.F. Oahu BISH 1987-05-28
66 Psidium cattleyanum Waianae Mts., Pohakea Pass Preserved_Specimen Ozaki, E.T. 1472 Oahu BISH 1955-04-17
67 Psidium cattleyanum Keaiwa Heiau State Rec. Area 10 mi NW of Honolulu Preserved_Specimen Little Jr., E.L. 31010 Oahu BISH 1975-08-05
68 Psidium cattleyanum beyond Rainbow Falls Preserved_Specimen Lyon, H.L. 1061 Hawaii BISH 1909-06-21
69 Psidium cattleyanum Iao Valley Preserved_Specimen Wagner, W.L. 6491 Maui BISH 1991-08-14
70 Psidium cattleyanum Waimea, Na Pali-Kona forest reserve, Mohihi road, Kawaikoi stream Preserved_Specimen Chock, A.K. 964 Kauai BISH 1953-08-17
71 Psidium cattleyanum Waipio, S ridge of Kipapa Gulch Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-05-15
72 Psidium cattleyanum Mt. Kaala NAR summit plateau, along ridge trail Preserved_Specimen Char, W.P. 30 Oahu BISH 1985-09-20
73 Psidium cattleyanum Waianae Mountains, Kamaileunu Ridge, a SW ridge separating Waianae Kai and Makaha Valleys Preserved_Specimen Wagner, W.L. 5097 Oahu BISH 1983-12-11
74 Psidium cattleyanum Kipapa Preserved_Specimen Yoshinaga, E.K. Oahu BISH 1929-11-10
75 Psidium cattleyanum Waipio, S ridge of Kipapa Gulch Preserved_Specimen Chang, I. Oahu BISH 1932-05-15
76 Psidium cattleyanum Waikane Preserved_Specimen 31 Oahu BISH 1963-09-02
77 Psidium cattleyanum Honolulu, Nuuanu Ave, near Wilder residence Preserved_Specimen Kerr, M. Oahu BISH 1945-05-31
78 Psidium cattleyanum Hilo Preserved_Specimen Faurie, A. 49 Hawaii BISH 1909-05-01
79 Psidium cattleyanum Pupukea, Koolau Mts Preserved_Specimen Stone, B.C. 3647 Oahu BISH 1961-09-17
80 Psidium cattleyanum Girl Scout Camp, 4 miles from Sunset Beach in N Koolau Mts Preserved_Specimen Buchanan, H. 578 Oahu BISH 1961-08-30
81 Psidium cattleyanum Wahiawa Bog (Kanaele Swamp); margin of swamp Preserved_Specimen Wagner, W.L. 5206 Kauai BISH 1983-12-22
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