Description
Key Characters:
Growth Form:
Monoecious or apparently sometimes dioecious shrubs 0.3–1(–3) m tall.
Stems:
Stems erect or when more than 1 m long, usually scandent with watery sap, branching non–phyllanthoid, main stem leafy and bearing flowers.
Roots:
Leaves:
Leaves simple.
Alternate, distichous.
Blades variable in shape, lanceolate to elliptic, sometimes ovate or broadly elliptic, (9–)15–65(–85) mm long, (5–)10–30(–34) mm wide.
Apex acute to rounded.
Base rounded or sometimes slightly cordate.
Surfaces glabrous or with a few widely scattered hairs.
Margins entire.
Petioles 0–3(–4) mm long.
Stipules cordate, 1–2 mm long, deciduous.
Flowers:
Flowers in axillary cymes (if staminate) or sometimes solitary (if pistillate).
Flowers unisexual.
Calyx of staminate flowers with 6 lobes, ca. 1–1.3 mm long. Calyx of pistillate flower 6-lobed, ca. 3 mm long.
Corolla (petals) absent.
Stamens in staminate flowers 3; filaments distinct.
Ovary superior; styles distinct, deeply bifid.
Fruit:
Capsules green to brown; depressed–globose; 3–3.5 mm in diameter; explosively dehiscent.
Seeds angled; ca. 1 mm long.
Ploidy:
Habitat:
Mesic forest; often on steep slopes or ridge tops; sometimes in dry shrubland.
Elevation Range:
60–950 m.