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Dryopteris unidentata var. paleacea

(Hillebr.) Herat ex Fraser-Jenk. (1994)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Dryopteridaceae Genus: Dryopteris

‘akole

Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Check
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Bibliography

Name Published In: Thaiszia 4(1): 31 (1994)

Other References

Palmer 2003:150 (K, O, Mo, M, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Dryopteris unidentata var. paleacea W Maui, Lihau Montane Ohia Mixed Wet Forest. Near the summit of Lihau with Metrosideros, Coprosma, Broussaisia, Cibotium, Cyrtandra, Diplazium spp., Marattia, Microlepia and Dryopteris spp. Preserved_Specimen Welton, P. 985 Maui BISH 1991-07-09
2 Dryopteris unidentata var. paleacea Forest near Hilo Forest Preserved_Specimen Newell, C.L. Hawaii BISH 1910-01-01
3 Dryopteris unidentata var. paleacea head of Awehi Gulch, Kaohai forest Preserved_Specimen St.John, H. 18881 Lanai BISH 1938-04-15
4 Dryopteris unidentata var. paleacea Nature Conservancy Kamakou Preserve General Vegetation Monitoring Plot 1. 100 meters from Papaala Pali Open Metrosideros with Cibotium chamissoi, Cheirodendron, Labordia, Trematolobelia and Dubautia. Preserved_Specimen Higashino, P.K. 10094 Molokai BISH 1983-10-12
5 Dryopteris unidentata var. paleacea Keanakolu; Humuula; Hilo Forest Reserve; N. Hilo Dist Moist gulch in acadia koa woods. Preserved_Specimen Large tufts; rhizome subterranean. Frond 1. St.John, H. 18590 Hawaii BISH 1937-12-30
6 Dryopteris unidentata var. paleacea Waikolu Valley (transect 27, near USFWS station 5) In understory of Diospyros-Metrosideros forest, steep ridge on sidewalls of valley. Preserved_Specimen Kepler, A.K. Molokai BISH 1981-03-31
7 Dryopteris unidentata var. paleacea Slopes of Mauna Loa In wet forest. Preserved_Specimen Rock, J.F.C. 6 Hawaii BISH 1958-01-01
8 Dryopteris unidentata var. paleacea Kukuinui Ridge, Wailau Valley moist woods Preserved_Specimen St.John, H. 13287 Molokai BISH 1933-07-04
9 Dryopteris unidentata var. paleacea Kohala mountains, Puu Ahia area; northof Puu on drainage of Waipahoehoe stream. In Metrosideros-Cibotium wet forest with Cheirodendron, Psychotria, Myrsine, Melicope clusifolia, Melicope volcanica, Styphelia, Ilex, Clermontia waimeae, Clermontia calophylla, Trematolobelia, Broussaisia, Labordia acuminatum, Hedyotis terminalis. Preserved_Specimen Large erect fern 1-1.5 meters high, fronds 3-4 pinnate, stipes scales large, prominent, dense, deltate, distal stipe scales prominent, reddish brown to dark brown rachises and costae with numerous scales. Perlman, S. 17834 Hawaii BISH 2001-12-19
10 Dryopteris unidentata var. paleacea Kauai Waimea District. Kauaikinana headwaters below Pihea. Metrosideros-Cheirodendron-Dicranopteris montane wet, summer mesic forest with Polyscias waimeae, P. waialealae, Melicope clustifolia, M. puberula, Bobea brevipes, Scaevola procera, S. glabra, Elaeocarpus bifudus, Xylosma hawaiiense, Syzyglum sandwicensis, Psychotria marinian Preserved_Specimen Terrestrial rhizome erect, stipe and rachis covered in red-brown scales, observed with D. u. unidentata, uncommon. Wood, K.R. 15073 Kauai BISH 2014-09-11
11 Dryopteris unidentata var. paleacea Palikea. Southern Waianae mountains. Mesic forest dominated by Schinus terebinthifolius, Morella faya, and Psidium cattleianum with remnant native canopy intermixed. Native plants include Metrosideros polymorpha, Cheirodendron trigynum, Microlepia setosa, Dryopteris glabra var. nuda, Wikstroemia oahuensis, Cyanea membranaceae, Clermontia oblongifolia, Athyrium microphyllum, Diplazium sandwichinanum, Kadua affinis, Asplenium lobulatum, and many others. Preserved_Specimen Large fern 2 m tall, the stipe and rachis densely covered with broadly deltate scales, the scales easily falling away when disturbed. Frond 3 pinnate pinnatifid, sori exiindusiate. Costa and costules with abundant hairlike dark brown scales. Stipe and frond somewhat viscous and sticky. Thomas, M.K. 417 Oahu BISH 2022-07-29
12 Dryopteris unidentata var. paleacea Hanalei District. Hanakapiai, eastern branch, Hono O Na Pali NAR,. Metrosideros-Cheirodendron-Dicranopteris montane wet forest with Diplopterygium pinnatum, Dryopteris unidentata var. paleacea, Scaevola procera, Pritchardia perlmanii, Kadua affinis, Psychotria mariniana, P. hexandra, Syzygium sandwicensis, Melicope wawrana, Bro Preserved_Specimen Terrestrial fern, rhizome sub-erect 17 cm. long X 15 cm. wide.; occasional. Wood, K.R. 17163 Kauai BISH 2016-10-25
13 Dryopteris unidentata var. paleacea Kapulei Metrosideros, Dicranopteris, Sadleria pallida, Perrottetia sandwicensis Preserved_Specimen rhizome dark brown, sub-erect 20 cm long; stipe 1 m long, covered thickly at base with red-brown scales, thining upwards to rachis; lamina 125 cm long Wood, K.R. 7216 Molokai BISH 1998-02-24
14 Dryopteris unidentata var. paleacea W Maui, Lahaina District, on trail to Puu Kukui In closed canopy Ohia woods. Preserved_Specimen Palmer, D.D. 678 Maui BISH 1991-07-03
15 Dryopteris unidentata var. paleacea W Maui, Lihau Montane Ohia Mixed Wet Forest Preserved_Specimen Welton, P. 985 Maui BISH 1991-07-09
16 Dryopteris unidentata var. paleacea Kamakou Preserve. Hanalilolilo Trail in upper part of preserve Wet montane Metrosideros forest. Preserved_Specimen Large terrestrial fern Pratt, L.W. Molokai BISH 1982-05-27
17 Dryopteris unidentata var. paleacea Makawao District, eastern Honomanu Metrosieros-Cheirodendron-Dicranopteris wet forest with Perrottetia sandwicensis, Diplazium sandwichianum, Gunnera, Dryopteris wallichiana, Pteris excelsa, Sadleria spp, Asplenium spp, Scaevola chamissoniana, Laborida hirtella, Astelia menziesiana, Cl Preserved_Specimen terrestrial rhizome erect or sub-erect, scales glossy dark brown Wood, K.R. 15046 Maui BISH 2012-06-26
18 Dryopteris unidentata var. paleacea Keanakolu; Humuula; Hilo Forest Reserve; N. Hilo Dist Moist gulch in acadia koa woods. Preserved_Specimen Large tufts; rhizome subterranean. Frond 1. St.John, H. 18590 Hawaii BISH 1937-12-30
19 Dryopteris unidentata var. paleacea Palikea. Southern Waianae mountains. Mesic forest dominated by Schinus terebinthifolius, Morella faya, and Psidium cattleianum with remnant native canopy intermixed. Native plants include Metrosideros polymorpha, Cheirodendron trigynum, Microlepia setosa, Dryopteris glabra var. nuda, Wikstroemia oahuensis, Cyanea membranaceae, Clermontia oblongifolia, Athyrium microphyllum, Diplazium sandwichinanum, Kadua affinis, Asplenium lobulatum, and many others. Preserved_Specimen Large fern 2 m tall, the stipe and rachis densely covered with broadly deltate scales, the scales easily falling away when disturbed. Frond 3 pinnate pinnatifid, sori exiindusiate. Costa and costules with abundant hairlike dark brown scales. Stipe and frond somewhat viscous and sticky. Thomas, M.K. 417 Oahu BISH 2022-07-29
20 Dryopteris unidentata var. paleacea Hanalei District. Hanakapiai, eastern branch, Hono O Na Pali NAR,. Metrosideros-Cheirodendron-Dicranopteris montane wet forest with Diplopterygium pinnatum, Dryopteris unidentata var. paleacea, Scaevola procera, Pritchardia perlmanii, Kadua affinis, Psychotria mariniana, P. hexandra, Syzygium sandwicensis, Melicope wawrana, Bro Preserved_Specimen Terrestrial fern, rhizome sub-erect 17 cm. long X 15 cm. wide.; occasional. Wood, K.R. 17163 Kauai BISH 2016-10-25
21 Dryopteris unidentata var. paleacea Palikea. Southern Waianae mountains. Mesic forest dominated by Schinus terebinthifolius, Morella faya, and Psidium cattleianum with remnant native canopy intermixed. Native plants include Metrosideros polymorpha, Cheirodendron trigynum, Microlepia setosa, Dryopteris glabra var. nuda, Wikstroemia oahuensis, Cyanea membranaceae, Clermontia oblongifolia, Athyrium microphyllum, Diplazium sandwichinanum, Kadua affinis, Asplenium lobulatum, and many others. Preserved_Specimen Large fern 2 m tall, the stipe and rachis densely covered with broadly deltate scales, the scales easily falling away when disturbed. Frond 3 pinnate pinnatifid, sori exiindusiate. Costa and costules with abundant hairlike dark brown scales. Stipe and frond somewhat viscous and sticky. Thomas, M.K. 417 Oahu BISH 2022-07-29
22 Dryopteris unidentata var. paleacea Kokee State Park, Kalalau Valley at 18.4 of Hwy 550 Steep, forested slope below rim of valley. Terrestrial. Preserved_Specimen 5 live fronds with several old stipes below. Growing with Dryopteris (Flynn 2110) and Diplazium under Metrosideros canopy. Flynn, T.W. 2111 Kauai BISH 1987-03-20
23 Dryopteris unidentata var. paleacea Kamakou Preserve. Hanalilolilo Trail in upper part of preserve Wet montane Metrosideros forest. Preserved_Specimen Large terrestrial fern Pratt, L.W. Molokai BISH 1982-05-27
24 Dryopteris unidentata var. paleacea Makawao District, eastern Honomanu Metrosideros-Cheirodendron-Dicranopteris wet forest with Perrottetia sandwicensis, Diplazium sandwichianum, Gunnera, Dryopteris wallichiana, Pteris excelsa, Sadleria spp, Asplenium spp, Scaevola chamissoniana, Labordia hirtella, Astelia menziesiana, Cl Preserved_Specimen terrestrial rhizome erect or sub-erect, scales glossy dark brown Wood, K.R. 15046 Maui BISH 2012-06-26
25 Dryopteris unidentata var. paleacea W Maui, Lihau Montane Ohia Mixed Wet Forest Preserved_Specimen Welton, P. 979 Maui BISH 1991-07-09
26 Dryopteris unidentata var. paleacea Ainapo to Puu Pili, Kapapala Kau Dist. dry Metrosideros woods Preserved_Specimen St.John, H. 18434 Hawaii BISH 1937-12-21
27 Dryopteris unidentata var. paleacea Hanalei District. Hanakapiai, eastern branch, Hono O Na Pali NAR,. Metrosideros-Cheirodendron-Dicranopteris montane wet forest with Diplopterygium pinnatum, Dryopteris unidentata var. paleacea, Scaevola procera, Pritchardia perlmanii, Kadua affinis, Psychotria mariniana, P. hexandra, Syzygium sandwicensis, Melicope wawrana, Bro Preserved_Specimen Terrestrial fern, rhizome sub-erect 17 cm. long X 15 cm. wide.; occasional. Wood, K.R. 17163 Kauai BISH 2016-10-25
28 Dryopteris unidentata var. paleacea Nature Conservancy Kamakou Preserve General Vegetation Monitoring Plot 1. 100 meters from Papaala Pali Open Metrosideros with Cibotium chamissoi, Cheirodendron, Labordia, Trematolobelia and Dubautia. Preserved_Specimen Higashino, P.K. 10094 Molokai BISH 1983-10-12
29 Dryopteris unidentata var. paleacea Hanalei District, Honopu, NW of Kainamanu Acacia koa- Metrosideros polymorpha - Diranopteris linearis montane mesic with Bobea brevipes, Ilex anomala, Wikstroemia furcata, Claoxylon sandwicensis, Polyscias waimeae, Melicope anisata, M. barbigera, Kadua affinis, Psychotria greenwelliae, Planchonella sandwicensis, Do Preserved_Specimen rhizome erect 15 cm long, 10 cm wide, occasional Wood, K.R. 15157 Kauai BISH 2012-08-08
30 Dryopteris unidentata var. paleacea Kauai Waimea District. Kauaikinana headwaters below Pihea. Metrosideros-Cheirodendron-Dicranopteris montane wet, summer mesic forest with Polyscias waimeae, P. waialealae, Melicope clustifolia, M. puberula, Bobea brevipes, Scaevola procera, S. glabra, Elaeocarpus bifudus, Xylosma hawaiiense, Syzyglum sandwicensis, Psychotria marinian Preserved_Specimen Terrestrial rhizome erect, stipe and rachis covered in red-brown scales, observed with D. u. unidentata, uncommon. Wood, K.R. 15073 Kauai BISH 2014-09-11
31 Dryopteris unidentata var. paleacea W Maui, Lahaina District, up trail from Haelaau Cabin at transect station 4.5400 mesic forest Preserved_Specimen fern to 2 m Annable, C.R. 3860 Maui BISH 1998-03-05
32 Dryopteris unidentata var. paleacea Mountains. Preserved_Specimen Nelson, D. 12 Hawaii BISH 1779-01-01
33 Dryopteris unidentata var. paleacea Kapulei Metrosideros, Dicranopteris, Sadleria pallida, Perrottetia sandwicensis Preserved_Specimen rhizome dark brown, sub-erect 20 cm long; stipe 1 m long, covered thickly at base with red-brown scales, thining upwards to rachis; lamina 125 cm long Wood, K.R. 7216 Molokai BISH 1998-02-24
34 Dryopteris unidentata var. paleacea Keanakolu; Humuula; Hilo Forest Reserve; N. Hilo Dist Moist gulch in acadia koa woods. Preserved_Specimen Large tufts; rhizome subterranean. Frond 1. St.John, H. 18590 Hawaii BISH 1937-12-30
35 Dryopteris unidentata var. paleacea North Hilo District: Keanakolu; Humuula; Hilo Forest Reserve Moist gulch in acadia koa woods. Preserved_Specimen Large tufts; rhizome subterranean. Frond 1. St.John, H. 18590 Hawaii BISH 1937-12-30
36 Dryopteris unidentata var. paleacea Hanalei District, Honopu, NW of Kainamanu Acacia koa- Metrosideros polymorpha - Diranopteris linearis montane mesic with Bobea brevipes, Ilex anomala, Wikstroemia furcata, Claoxylon sandwicensis, Polyscias waimeae, Melicope anisata, M. barbigera, Kadua affinis, Psychotria greenwelliae, Planchonella sandwicensis, Do Preserved_Specimen rhizome erect 15 cm long, 10 cm wide, occasional Wood, K.R. 15157 Kauai BISH 2012-08-08
37 Dryopteris unidentata var. paleacea W Maui, Lahaina District, on trail to Puu Kukui In closed canopy Ohia woods. Preserved_Specimen Palmer, D.D. 678 Maui BISH 1991-07-03
38 Dryopteris unidentata var. paleacea Hanalei District. Hanakapiai, eastern branch, Hono O Na Pali NAR,. Metrosideros-Cheirodendron-Dicranopteris montane wet forest with Diplopterygium pinnatum, Dryopteris unidentata var. paleacea, Scaevola procera, Pritchardia perlmanii, Kadua affinis, Psychotria mariniana, P. hexandra, Syzygium sandwicensis, Melicope wawrana, Bro Preserved_Specimen Terrestrial fern, rhizome sub-erect 17 cm. long X 15 cm. wide.; occasional. Wood, K.R. 17163 Kauai BISH 2016-10-25
39 Dryopteris unidentata var. paleacea W Maui District, Hanaula Springs Trail Preserved_Specimen Large knobby caudex, stipe 3 feet, densely covered with orange curly scales, frond 3 feet, quadripinnatifid, sulcate to third dividion, sori naked. Hobdy, R.W. 1065 Maui BISH 1981-05-06
40 Dryopteris unidentata var. paleacea Slopes of Mauna Loa In wet forest. Preserved_Specimen Rock, J.F.C. 6 Hawaii BISH 1958-01-01
41 Dryopteris unidentata var. paleacea Kalanilehua, Olaa. Preserved_Specimen Continuation of apex. Giffard, W.M. 347 Hawaii BISH 1928-07-01
42 Dryopteris unidentata var. paleacea W Maui, Hanaula Springs trail, in lush gully Preserved_Specimen Fern 4 ft tall; stipe and bottom side of rachis clothed with broad curly scales; frond tripinnate to quadripinnatifid; side of rachis sulcate; sori naked, brown, in large compact clusters. Looks same as Hobdy 957. Hobdy, R.W. 975 Maui BISH 1981-01-13
43 Dryopteris unidentata var. paleacea W Maui, Hanaula Forest Preserved_Specimen Red curly scales to fourth division (costules); frond quadripinnatifid. Hobdy, R.W. 2095 Maui BISH 1984-05-16
44 Dryopteris unidentata var. paleacea East Maui, Kula slope of Haleakala Preserved_Specimen Frond breast high, dark green, veins pellucid. Forbes, C.N. 2166 Maui BISH 1920-04-07
45 Dryopteris unidentata var. paleacea W Maui, Hanaula Forest Preserved_Specimen Red curly scales to fourth division (costules); frond quadripinnatifid. Hobdy, R.W. 2095 Maui BISH 1984-05-16
46 Dryopteris unidentata var. paleacea W Maui District, Hanaula Springs Trail Preserved_Specimen Large knobby caudex, stipe 3 feet, densely covered with orange curly scales, frond 3 feet, quadripinnatifid, sulcate to third dividion, sori naked. Hobdy, R.W. 1065 Maui BISH 1981-05-06
47 Dryopteris unidentata var. paleacea Kalanilehua, Olaa. Preserved_Specimen Continuation of base. Giffard, W.M. 347 Hawaii BISH 1928-07-01
48 Dryopteris unidentata var. paleacea W Maui, Lahaina District, Puu Kukui Trail, in forests fringing the bogs at Violet Lake Preserved_Specimen Fern 3-4 ft tall with densely red-scaly stipe; rachis sulcate to last division on top; veins prominent both above and below; sori naked, sporangia dark brown, shiny. Hobdy, R.W. 957 Maui BISH 1980-12-11
49 Dryopteris unidentata var. paleacea Kalanilehua, Olaa. Preserved_Specimen Upper part of frond. Giffard, W.M. Hawaii BISH 1927-07-26
50 Dryopteris unidentata var. paleacea W Maui District, Along Eke trail Preserved_Specimen Hobdy, R.W. 1694 Maui BISH 1982-10-21
51 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Forbes, C.N. 915 BISH 1919-07-25
52 Dryopteris unidentata var. paleacea Kalanilehua, Olaa. Preserved_Specimen Lower half of frond. Giffard, W.M. Hawaii BISH 1927-07-26
53 Dryopteris unidentata var. paleacea Kalanilehua, Olaa. Preserved_Specimen Middle of frond. Giffard, W.M. Hawaii BISH 1927-07-26
54 Dryopteris unidentata var. paleacea Maui Preserved_Specimen Forbes, C.N. 1735 Maui BISH 1919-11-27
55 Dryopteris unidentata var. paleacea East Maui Preserved_Specimen Hobdy, R.W. 2082 Maui BISH 1984-05-01
56 Dryopteris unidentata var. paleacea Kalanilehua, Olaa. Preserved_Specimen Apex of frond. Giffard, W.M. 347 Hawaii BISH 1928-07-01
57 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Rock, J.F.C. BISH
58 Dryopteris unidentata var. paleacea Maui Preserved_Specimen Mann, H. 502 Maui BISH 1864-01-01
59 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Lyon, H.L. BISH
60 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Lyon, H.L. BISH
61 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Lyon, H.L. BISH
62 Dryopteris unidentata var. paleacea W Maui, Puu Kukui Preserved_Specimen Krajina, V.J. 620418056 Maui BISH 1962-04-18
63 Dryopteris unidentata var. paleacea Haleakala, Keanae Gap, Halehoku Preserved_Specimen Forbes, C.N. 1077 Maui BISH 1919-08-07
64 Dryopteris unidentata var. paleacea West Maui, Puu Kukui 20.8N 156.6W Preserved_Specimen Krajina, V.J. 620418056 Maui BISH 1962-04-18
65 Dryopteris unidentata var. paleacea Maui Preserved_Specimen Mann, H. 502 Maui BISH 1864-01-01
66 Dryopteris unidentata var. paleacea Ridge left side of Kipahulu Valley Preserved_Specimen Forbes, C.N. 1735 Maui BISH 1919-11-27
67 Dryopteris unidentata var. paleacea Kauai Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
68 Dryopteris unidentata var. paleacea Kilauea, Wright Rd Preserved_Specimen Horner, E. Hawaii BISH 1959-07-01
69 Dryopteris unidentata var. paleacea East Maui, Olinda. Preserved_Specimen Lyon, H.L. Maui BISH 1908-12-05
70 Dryopteris unidentata var. paleacea Kaholuamano Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
71 Dryopteris unidentata var. paleacea Kilauea, Wright Rd Preserved_Specimen Horner, E. Hawaii BISH 1959-07-01
72 Dryopteris unidentata var. paleacea Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-06-30
73 Dryopteris unidentata var. paleacea Kauai. Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
74 Dryopteris unidentata var. paleacea [Hawaiian Islands; U.S.A.; O`ahu; Hawai`i; Honolulu] Preserved_Specimen Lay, G.T. Oahu BISH 1826-01-01
75 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Forbes, C.N. 915 BISH 1919-07-25
76 Dryopteris unidentata var. paleacea East Maui, Olinda. Preserved_Specimen Lyon, H.L. Maui BISH 1908-12-05
77 Dryopteris unidentata var. paleacea unknown Preserved_Specimen St.John, H. 18595 BISH 1937-12-30
78 Dryopteris unidentata var. paleacea W Maui District, Along Eke trail Preserved_Specimen Hobdy, R.W. 1694 Maui BISH 1982-10-21
79 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Rock, J.F.C. BISH
80 Dryopteris unidentata var. paleacea unknown Preserved_Specimen Rock, J.F.C. BISH
81 Dryopteris unidentata var. paleacea Hanalei District. Hanakapiai, eastern branch, Hono O Na Pali NAR,. Metrosideros-Cheirodendron-Dicranopteris montane wet forest with Diplopterygium pinnatum, Dryopteris unidentata var. paleacea, Scaevola procera, Pritchardia perlmanii, Kadua affinis, Psychotria mariniana, P. hexandra, Syzygium sandwicensis, Melicope wawrana, Bro Preserved_Specimen Terrestrial fern, rhizome sub-erect 17 cm. long X 15 cm. wide.; occasional. Wood, K.R. 17163 Kauai BISH 2016-10-25
82 Dryopteris unidentata var. paleacea Oahu Preserved_Specimen Lay, G.T. Oahu BISH 1826-01-01
83 Dryopteris unidentata var. paleacea W Maui, Lihau Montane Ohia Mixed Wet Forest. Near the summit of Lihau with Metrosideros, Coprosma, Broussaisia, Cibotium, Cyrtandra, Diplazium spp., Marattia, Microlepia and Dryopteris spp. Preserved_Specimen Welton, P. 979 Maui BISH 1991-07-09
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