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Clermontia tuberculata

C.N.Forbes (1912)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Asterales Family: Campanulaceae Genus: Clermontia

‘ōhā [oha], ‘ōhā wai [oha wai], hāhā [haha]

Description

Key Characters:

Growth Form: Shrubs or trees, 4–5 m tall, terrestrial.

Stems: Stems branched repeatedly, spreading, light gray or light brown, glabrous or rarely pubescent, with numerous helically arranged leaf scars near apex, pith solid, latex white, viscous.

Roots:

Leaves: Leaves simple. Alternate. Blades oblanceolate or oblong, 9–20 cm long, 2.5–5.5 cm wide. Apex obtuse. Base cuneate or attenuate. Upper surfaces glabrous; lower surfaces glabrous or minutely muricate on the veins; lamina coriaceous; upper surfaces dark green; lower surfaces pale green. Margins callose–crenulate. Petioles 1–6 cm long, muricate. Stipules absent.

Flowers: Flowers in inflorescences 2–flowered or often 1–flowered by abortion, muricate; peduncle 0.5–1 cm long, spreading, bibracteate at apex; pedicels 3–5 cm long, spreading, bibracteolate at base. Flowers bisexual (perfect), epigynous, zygomorphic, resupinate, protandrous. Hypanthium 1.4–1.9 cm long, 1.4–1.9 cm in diameter, obconic or obovoid, muricate. Calyx lobes 3–9 mm long, 3–5 mm wide, triangular, shallowly triangular, or deltate, muricate; apex obtuse or acute. Corolla 5–7 cm long, bilabiate, dark rose, muricate; tube suberect, 1–2 cm long, 1.2–1.8 cm in diameter; lobes thick, fleshy, the dorsal two 3.8–5.2 cm long, 3–5 cm wide, suberect, the ventral three 2–3 cm long, 3–5 mm wide, spreading. Stamens 5, syngenesious, alternate with the corolla lobes, staminal column somewhat exserted, suberect or curved; anthers connate, dithecal, opening introrsely by longitudinal slits, dark purple, the 3 dorsal ones a little longer than the 2 ventral ones, the latter with tufts of stiff white trichomes at apex, otherwise glabrous, or sometimes pubescent along the sutures, rarely so on the surfaces, the tube 1.5–2 cm long, 5.5–7.5 mm in diameter, glabrous or sparingly pubescent; filaments connate above, free from the corolla, 4.1–5.8 cm long, magenta, glabrous. Pollen tricolporate, prolate, ellipsoidal. Ovary inferior, 2-carpellate, 2-loculed; ovule placentation axile, placentae large; style slender, terete (cylindrical), with a ring of stiff white hairs near apex; stigma 2-lobed, the lobes appressed and non–receptive as the style grows through the anther tube, pushing out pollen, after which the stigmas spread and become receptive.

Fruit: Berries unknown; presumably muricate. Seeds numerous; tiny; dark brown or black; minutely foveate–reticulate; shiny; with a straight; short to spatulate dicotyledonous embryo embedded in oily endosperm.

Ploidy:

Habitat: Windward slopes of Haleakalā; montane rain forests and cloud forests.

Elevation Range: 1,650–1,825 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Maui Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Occas. Pap. Bernice Pauahi Bishop Mus. 5: 8 (1912)

Other References

Wagner et al. 1990:436 (EM); Lammers 1991:30 (KEY, DESCR)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23966 Maui PTBG 5/28/2014
2 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN palmer, r. CM 1/4/1994
3 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros mix shrub montane wet forest. In rich, pristine forest. PRESERVED_SPECIMEN Steve Perlman 10512 Maui PTBG 3/18/1988
4 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation PRESERVED_SPECIMEN K. R. Wood 15644 Maui PTBG 8/22/2013
5 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN j. f. rock 12837 Maui US
6 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 1682 Maui PTBG 3/11/1992
7 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Maui US
8 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood & s. p. perlman 6644 Maui US 8/27/1997
9 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN j. f. rock s.n. Maui US
10 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman, s. gon, m. buck & a. engilis 10512 Maui US 3/18/1988
11 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Maui US
12 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Montane wet forest. PRESERVED_SPECIMEN s. p. perlman & h. l. oppenheimer 21725 Maui US 8/14/2009
13 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J. F. 12837 Maui US 3/12/2020
14 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Gon, S.; Buck, M.; Engilis, A. 10512 Maui US 3/18/1988
15 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Hank Oppenheimer H61320 Maui PTBG 6/20/2013
16 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood & S. P. Perlman 6644 Maui US 8/27/1997
17 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J. F. s.n. Maui US 3/11/2020
18 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN perlman, s.p. gon, s.; buck, m. collector number: 8093 Hawaii US 3/20/1988
19 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information along perennial stream. Vegetation: Metrosideros, Cheirodendron, Broussaisia, Dicranopteris, Melicope, Sadleria, Cibotium, Clermontia, Cyanea, Astelia, Peperomia, Athyrium, Kadua, Lycopodiella, Vaccinium, Labordia, Isachne, Eragrostis, Deschampsia, Pipturus, Pilea, Machaerina, Dryopteris, Coprosma, Psychotria, Cyrtandra, Prunella, Ageratina, Tibouchina, Plantago major, Rubus hawaiensis, Korthalsella, PRESERVED_SPECIMEN Oppenheimer, H.L. H51420 Maui BISH 5/28/2014
20 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation W/ Broussaisia arguta, Myrsine sandwicensis, Dubautia plantaginea, D. scabra, Kadua affinis, K. axillaris, Scaevola chamissoniana, Clermontia samuelii subsp. hanaensis, C. tuberculata, C. grandiflora PRESERVED_SPECIMEN wood, k.r. bustamente, k. collector number: 15644 Maui BISH
21 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 12837 Maui BISH 3/1/1912
22 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation W/ Broussaisia arguta, Myrsine sandwicensis, Dubautia plantaginea, D. scabra, Kadua affinis, K. axillaris, Scaevola chamissoniana, Clermontia samuelii subsp. hanaensis, C. tuberculata, C. grandiflora PRESERVED_SPECIMEN Wood, K.R. 15612 Maui BISH 8/19/2013
23 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Medeiros, A.C. 458 Maui BISH 7/10/1983
24 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information stream bank with Broussaisia, Metrosideros, Gouldia PRESERVED_SPECIMEN harrison, b. collector number: 426 Maui BISH 8/14/1973
25 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Vegetation: Metrosideros, Cheirodendron, Polyscias, Broussaisia, Dicranopteris, Diplopterygium, Sticherus, Machaerina, Cibotium, Dubautia, Trematolobelia, Pipturus, Sadleria, Dryopteris, Kadua, Coprosma, Phyllostegia, Labordia, Carex, Scaevola, Eragrostis, Amauropelta, Adenophorus, Lellingeria, Lepisorus, Oreogrammitis, Pilea, Selaginella, Peperomia, Juncus, Tibouchina, Hedychium, Paspalum, Sacciolepis, Erechtites. PRESERVED_SPECIMEN Oppenheimer, H.L. H81335 Maui BISH 8/21/2013
26 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH 3/1/1911
27 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P. 6644 Maui US 8/27/1997
28 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros montane wet forest. PRESERVED_SPECIMEN Steve Perlman 12507 Maui PTBG 1/22/1992
29 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Montane wet forest. PRESERVED_SPECIMEN Perlman, S. P.; Oppenheimer, H. L. 21725 Maui US 8/14/2009
30 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information with Broussaisia, Cheirodendron, Clermontia, Metrosideros PRESERVED_SPECIMEN harrison, b. collector number: 413 Maui BISH 8/13/1973
31 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN medeiros, a.c. collector number: 458 Maui BISH 7/10/1983
32 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information in shady wet forest. Vegetation: Metrosideros, Cheirodendron, Melicope, Leptecophylla, Coprosma, Broussaisia, Ilex, Smilax, Cibotium, Sadleria, Athyrium, Diplazium, Clermontia, Kadua, Dicranopteris, Sticherus, Diplopterygium, Astelia, Uncinia, Elaphoglossum, Asplenium, Dryopteris, Vaccinium, Labordia, Nertera, Carex, Adenophorus, Peperomia, Tibouchina, Juncus, Axonopus. PRESERVED_SPECIMEN Oppenheimer, H.L. H61218 Maui BISH 6/26/2012
33 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information forest at edge of large open sedge meadow; with Broussaisia, Coprosma PRESERVED_SPECIMEN harrison, b. collector number: 557 Maui BISH 7/7/1973
34 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation W/ Broussaisia arguta, Myrsine sandwicensis, Dubautia plantaginea, D. scabra, Kadua affinis, K. axillaris, Scaevola chamissoniana, Clermontia samuelii subsp. hanaensis, C. tuberculata, C. grandiflora PRESERVED_SPECIMEN Wood, K.R. 15644 Maui BISH 8/22/2013
35 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information In a wet gully. PRESERVED_SPECIMEN Hobdy, R.W. 3591 Maui BISH 6/16/1993
36 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information With Myrsine, Broussaisia, Cyrtandra, Metrosideros, and Cheirodendron. PRESERVED_SPECIMEN harrison, b. collector number: 289 Maui BISH 6/29/1973
37 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information stream bank with Broussaisia, Metrosideros, Gouldia PRESERVED_SPECIMEN Harrison, B. 426 Maui BISH 8/14/1973
38 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Maui BISH
39 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 201.m Maui BISH 7/1/1910
40 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation W/ Bourssaisia arguta, Myrsine sandwicensis, Dubautia plantaginea, D. scabra, Kadua affinis, K. axillaris, Scaevola chamissoniana, Clermontia samuelii subsp. hanaensis, C. tuberculata, C. grandiflora PRESERVED_SPECIMEN Wood, K.R. 15635 Maui BISH 8/21/2013
41 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest with Perrottetia sandwicensis, Diplazium sandwichianum, Gunnera, Dryopteris wallichiana, Pteris excelsa, Sadleria spp, Asplenium spp, Scaevola chamissoniana, Labordia hirtella, Astelia menziesiana, Cl PRESERVED_SPECIMEN wood, k.r. bily, p., oppenheimer, h., bustamente, k. collector number: 15051 Maui BISH 6/26/2012
42 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Woods PRESERVED_SPECIMEN Forbes, C.N. 201.M Maui BISH 7/1/1910
43 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Maui BISH 3/1/1911
44 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Montane native rainforest PRESERVED_SPECIMEN Carter, I. CW9 Maui BISH 6/6/2011
45 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation W/ Bourssaisia arguta, Myrsine sandwicensis, Dubautia plantaginea, D. scabra, Kadua affinis, K. axillaris, Scaevola chamissoniana, Clermontia samuelii subsp. hanaensis, C. tuberculata, C. grandiflora PRESERVED_SPECIMEN wood, k.r. oppenhiemer, h. collector number: 15635 Maui BISH
46 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information forest at edge of large open sedge meadow; with Broussaisia, Coprosma PRESERVED_SPECIMEN Harrison, B. 557 Maui BISH 7/7/1973
47 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information wet ohia forest with native trees, shrubs, treefern PRESERVED_SPECIMEN Higashino, P.K. 9301 Maui BISH 6/28/1980
48 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Dodonaea, weedy grasses PRESERVED_SPECIMEN harrison, b. collector number: 256 Oahu BISH 6/29/1973
49 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN harrison, b. collector number: 256 Oahu BISH 6/29/1973
50 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Mossy tree trunk PRESERVED_SPECIMEN harrison, b. collector number: 288 Oahu BISH 6/29/1973
51 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Woods PRESERVED_SPECIMEN forbes, c.n. collector number: 201.m Maui BISH 7/1/1910
52 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information thicket with Broussaisia, Cheirodendron PRESERVED_SPECIMEN Harrison, B. 256 Maui BISH 6/29/1973
53 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest with Perrottetia sandwicensis, Diplazium sandwichianum, Gunnera, Dryopteris wallichiana, Pteris excelsa, Sadleria spp, Asplenium spp, Scaevola chamissoniana, Labordia hirtella, Astelia menziesiana, Cl PRESERVED_SPECIMEN Wood, K.R. 15051 Maui BISH 6/26/2012
54 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation W/ Broussaisia arguta, Myrsine sandwicensis, Dubautia plantaginea, D. scabra, Kadua affinis, K. axillaris, Scaevola chamissoniana, Clermontia samuelii subsp. hanaensis, C. tuberculata, C. grandiflora PRESERVED_SPECIMEN wood, k.r. bustamente, k. collector number: 15612 Maui BISH
55 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information In Metrosideros montane wet forest with Vaccinium, Dubautia, Rubus hawaiensis, Broussaisia, Styphelia, Clermontia, Coprosma, Psychotria, Melicope, Gunnera PRESERVED_SPECIMEN Perlman, S. 8093 Maui BISH 3/20/1988
56 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information In a wet gully. PRESERVED_SPECIMEN hobdy, r.w. evanson collector number: 3591 Maui BISH 6/16/1993
57 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 201.M Maui BISH 7/1/1910
58 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information wet ohia forest with native trees, shrubs, treefern PRESERVED_SPECIMEN higashino, p.k. holt, r.a. collector number: 9301 Maui BISH 6/28/1980
59 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Occasional in forest understory. Vegetation: Metrosideros, Chierodendron, Ilex, Coprosma, Melicope, Myrsine, Leptecophylla, Lobelia, Cyanea, Cibotium, Sadleria, Carex, Peperomia. PRESERVED_SPECIMEN Oppenheimer, H.L. H31702 Maui BISH 3/13/2017
60 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH
61 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Montane native rainforest PRESERVED_SPECIMEN carter, i. fay, k. collector number: cw9 Maui BISH 6/6/2011
62 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information thicket with Broussaisia, Cheirodendron, Vaccinium PRESERVED_SPECIMEN Harrison, B. 288 Maui BISH 6/29/1973
63 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 201.M Maui BISH 7/1/1910
64 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information with Broussaisia, Cheirodendron, Clermontia, Metrosideros PRESERVED_SPECIMEN Harrison, B. 413 Maui BISH 8/13/1973
65 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros- Cheirodendron trigynum- Dicranopteris linearis montane wet forest. PRESERVED_SPECIMEN Steve Perlman 21725 Maui PTBG 8/14/2009
66 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation. PRESERVED_SPECIMEN K. R. Wood 15612 Maui PTBG 8/19/2013
67 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris montane wet forest, dissected by riparian vegetation. PRESERVED_SPECIMEN K. R. Wood 15635 Maui PTBG 8/21/2013
68 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest with bogs. PRESERVED_SPECIMEN K. R. Wood 6830 Maui PTBG 10/6/1997
69 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros montane wet forest. PRESERVED_SPECIMEN Steve Perlman 8093 Maui PTBG 3/20/1988
70 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest. PRESERVED_SPECIMEN K. R. Wood 15051 Maui PTBG 6/26/2012
71 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information Metrosideros wet forest. PRESERVED_SPECIMEN K. R. Wood 6644 Maui PTBG 8/27/1997
72 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 201.m Maui BISH 7/1/1910
73 Clermontia tuberculata C.N.Forbes Contact Bishop Museum Botany department for locality information With Myrsine, Broussaisia, Cyrtandra, Metrosideros, and Cheirodendron. PRESERVED_SPECIMEN Harrison, B. 289 Maui BISH 6/29/1973
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