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Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

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Description

Key Characters:

Growth Form: Erect or ascending perennial herbs 4–20 dm tall, sometimes somewhat woody toward base.

Stems: Stems few to many-branched, glabrous or sparsely strigillose.

Roots:

Leaves: Leaves simple. Opposite and decussate (pairs of leaves alternating 90 degrees between nodes). Blades lanceolate to oblanceolate, 3–10 cm long, 1–1.5 cm wide. Apex acute. Base attenuate. Surfaces finely scaberulous. Margins coarsely serrate. Veins prominent. Sessile. Stipules absent.

Flowers: Flowers numerous in several slender, terminal spikes 3–12 cm long, sparsely short–pilose throughout; Bracts subulate, shorter than to as long as the calyx; short–pilose. Flowers bisexual (perfect), alternate, nearly zygomorphic. Calyx sometimes purple, 1.5–2.3 mm long. Corolla usually blue, occasionally pale blue, lavender, or purple, ca. 2–3 mm long, 5-lobed, imbricate. Stamens 4, in 2 pairs, inserted on and included in corolla tube; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-lobed, 2-carpellate, 4-celled, placentation axile; ovules 1 per cell, attached laterally or near base, erect, anatropous; style 2-lobed, the lobes dissimilar, only 1 stigmatic.

Fruit: Schizocarps readily separating at maturity into 4 crustaceous nutlets; pericarp hard and dry. Seeds 1 per nutlet; oblong; triangular in cross section; endosperm essentially absent.

Ploidy: 2n = 28; 56

Habitat: Occurring in dry to wet; disturbed habitats.

Elevation Range: 10–2,280 m.

Historical Distribution

Images

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Synonyms (32)

Verbena affinis M.Martens & Galeotti (1844)
Verbena bonariensis (Misapplied)
Verbena bonariensis var. litoralis (Kunth) Hook. ex Müll.Berol. (1845)
Verbena brasiliensis var. subglabrata Moldenke (1950)
Verbena cajamarcensis Binder ex Molinari (2015)
Verbena caracasana Kunth (1818)
Verbena carolina var. glabra Moldenke (1959)
Verbena gentryi Moldenke (1941)
Verbena glabrata var. tenuispicata Moldenke (1967)
Verbena integrifolia Sessé & Moc. (1886)
Verbena integrifolia f. albiflora Moldenke (1968)
Verbena lanceolata Willd. ex Spreng. (1825)
Verbena litoralis f. albiflora (Moldenke) Moldenke (1979)
Verbena litoralis f. magnifolia Moldenke (1977)
Verbena litoralis var. albiflora Moldenke (1940)
Verbena litoralis var. caracasana (Kunth) Briq. (1904)
Verbena litoralis var. glabrior Benth. (1846)
Verbena litoralis var. leptostachya Schauer (1847)
Verbena litoralis var. litoralis
Verbena litoralis var. melanopotamica Hauman (1913)
Verbena litoralis var. portoricensis Moldenke (1982)
Verbena litoralis var. pycnostachya Schauer (1847)
Verbena litoralis var. subglabrata (Moldenke) N.O'Leary & Múlgura (2012)
Verbena longifolia M.Martens & Galeotti (1844)
Verbena longifolia f. albiflora Moldenke (1961)
Verbena longifolia var. pubescens Moldenke (1966)
Verbena minutiflora var. peruviana Moldenke (1981)
Verbena nudiflora Nutt. ex Turcz. (1863)
Verbena paucifolia Turcz. (1863)
Verbena sedula Moldenke (1955)
Verbena sedula var. darwinii Moldenke (1968)
Verbena sedula var. fournieri Moldenke (1969)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Kaho'olawe Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Notes

  • Description digitized by Ikaika Mendez
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • Has been used medicinally in Hawai'i especially as a mash applied to cuts and bruises and also to sprained and fractured areas.
  • in 1837, specific locality unknown

Bibliography

Name Published In: F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 2: 276 (1818)

Other References

Wagner et al. 1990:1325 (Mi, Ni, K, O, Mo, L, M, Ka, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
2 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
3 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
4 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
5 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
6 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
7 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
8 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
9 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
10 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
11 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
12 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
13 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
14 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
15 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
16 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
17 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
18 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
19 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
20 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
21 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
22 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
23 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
24 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
25 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
26 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
27 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
28 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
29 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
30 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
31 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
32 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
33 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
34 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
35 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
36 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
37 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
38 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
39 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
40 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
41 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
42 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
43 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
44 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
45 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
46 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
47 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
48 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
49 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
50 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
51 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
52 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
53 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
54 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
55 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
56 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
57 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
58 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
59 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
60 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
61 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
62 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
63 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
64 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
65 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
66 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
67 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
68 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
69 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
70 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
71 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
72 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
73 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
74 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
75 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
76 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
77 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
78 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
79 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
80 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
81 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
82 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
83 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
84 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
85 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
86 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
87 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
88 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
89 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
90 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
91 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
92 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
93 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
94 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
95 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
96 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
97 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
98 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
99 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
100 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
101 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
102 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
103 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
104 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
105 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
106 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
107 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
108 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
109 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
110 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
111 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
112 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
113 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
114 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
115 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
116 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
117 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
118 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
119 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
120 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
121 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
122 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
123 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
124 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
125 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
126 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
127 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
128 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
129 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
130 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
131 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
132 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
133 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
134 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
135 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
136 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
137 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
138 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
139 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
140 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
141 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
142 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
143 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
144 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
145 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
146 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
147 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
148 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
149 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
150 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
151 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
152 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
153 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
154 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
155 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
156 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
157 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
158 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
159 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
160 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
161 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
162 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
163 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
164 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
165 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
166 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
167 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
168 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
169 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
170 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
171 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
172 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
173 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
174 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
175 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
176 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
177 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
178 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
179 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
180 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
181 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
182 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
183 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
184 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
185 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
186 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
187 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
188 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
189 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
190 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
191 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
192 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
193 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
194 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
195 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
196 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
197 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
198 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
199 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
200 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
201 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
202 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
203 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
204 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
205 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
206 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
207 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
208 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
209 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
210 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
211 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
212 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
213 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
214 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
215 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
216 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
217 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
218 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
219 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
220 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
221 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
222 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
223 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
224 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
225 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
226 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
227 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
228 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
229 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
230 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
231 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
232 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
233 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
234 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
235 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
236 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
237 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
238 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
239 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
240 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
241 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
242 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
243 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
244 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
245 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
246 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
247 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
248 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
249 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
250 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
251 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
252 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
253 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
254 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
255 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
256 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
257 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
258 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
259 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
260 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
261 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
262 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
263 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
264 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
265 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
266 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
267 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
268 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
269 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
270 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
271 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
272 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
273 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
274 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
275 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
276 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
277 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
278 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
279 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
280 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
281 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
282 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
283 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
284 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
285 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
286 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
287 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
288 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
289 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
290 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
291 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
292 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
293 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
294 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
295 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
296 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
297 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
298 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
299 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
300 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
301 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
302 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
303 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
304 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
305 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
306 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
307 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
308 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
309 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
310 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
311 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
312 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
313 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
314 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
315 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
316 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
317 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
318 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
319 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
320 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
321 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
322 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
323 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
324 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
325 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
326 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
327 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
328 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
329 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
330 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
331 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
332 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
333 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
334 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
335 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
336 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
337 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
338 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
339 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
340 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
341 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
342 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
343 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
344 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
345 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
346 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
347 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
348 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
349 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
350 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
351 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
352 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
353 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
354 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
355 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
356 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
357 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
358 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
359 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
360 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
361 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
362 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
363 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
364 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
365 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
366 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
367 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
368 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
369 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
370 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
371 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
372 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
373 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
374 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
375 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
376 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
377 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
378 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
379 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
380 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
381 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
382 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
383 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
384 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
385 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
386 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
387 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
388 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
389 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
390 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
391 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
392 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
393 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
394 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
395 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
396 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
397 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
398 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
399 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
400 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
401 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
402 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
403 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
404 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
405 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
406 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
407 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
408 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
409 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
410 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
411 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
412 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
413 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
414 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
415 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
416 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
417 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
418 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
419 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
420 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
421 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
422 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
423 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
424 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
425 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
426 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
427 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
428 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
429 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
430 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
431 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
432 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
433 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
434 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
435 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
436 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
437 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
438 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
439 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
440 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
441 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
442 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
443 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
444 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
445 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
446 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
447 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
448 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
449 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
450 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
451 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
452 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
453 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
454 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
455 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
456 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
457 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
458 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
459 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
460 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
461 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
462 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
463 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
464 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
465 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
466 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
467 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
468 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
469 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
470 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
471 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
472 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
473 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
474 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
475 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
476 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
477 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
478 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
479 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
480 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
481 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
482 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
483 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
484 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
485 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
486 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
487 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
488 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
489 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
490 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
491 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
492 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
493 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
494 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
495 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
496 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
497 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
498 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
499 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
500 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
501 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
502 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
503 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
504 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
505 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
506 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
507 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
508 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
509 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
510 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
511 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
512 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
513 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
514 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
515 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
516 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
517 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
518 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
519 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
520 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
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