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Psilotum nudum

(L.) P.Beauv. (1805)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Psilotales Family: Psilotaceae Genus: Psilotum

‘o‘omoa, moa, moa nahele, pipi, upright whiskfern, whisk fern

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

  • Branched stem used in children's game called 'moa nahele' (cock fighting), winner would announce victory by crowing like a rooster (Krauss 1993:88). Spores used like talcum powder (Valier 1995:4).

PROPAGATION/CULTIVATION

  • Grows well under medium to high light in moist-dry soils, as well as in well-drained, uncut moss (Hoshizaki & Moran 2002:470).

Natural History

Statewide Status

Indigenous

Island Status

Ni'ihau(Incl. Lehua) Indigenous
Kuaihelani (Midway Atoll) Extirpated
Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Kaho'olawe Unknown
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Prodr. Aethéogam.: 112 (1805)

Other References

Palmer 2003:272 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst 2005:46 (DESCR); Kennedy et al. 2010:23 (Mi); Starr & Starr 2008/Caum 35 BISH (Mi†, 1923); Note: no BISH spms. from Ka, as of Mar 2025, retained based on Warren et al. (1994)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Psilotum nudum Makiki Preserved_Specimen Mann, H. 140 Oahu BISH 1864-01-01
2 Psilotum nudum Haupu Mountain, Mahaulepu, southern Kauai. Preserved_Specimen Epiphytic. Higgins, N.H. 146 Kauai BISH 1956-12-31
3 Psilotum nudum Oahu Preserved_Specimen Skottsberg, C.J.F. 242064 Oahu BISH 1924-08-17
4 Psilotum nudum N.E. slope Puu Olomana, Kailua rock crevices, aleurites forest Preserved_Specimen Fosberg, F.R. 10761 Oahu BISH 1935-01-20
5 Psilotum nudum Hilo terminal portion of the 1860-1 lava flow Preserved_Specimen Forbes, C.N. 552 Hawaii BISH 1915-05-17
6 Psilotum nudum Koolau Range, Waikane-Schofield Trail Preserved_Specimen Cowan, R.S. 66 Oahu BISH 1946-09-22
7 Psilotum nudum Waimea Forest; on rocks and trees. Preserved_Specimen Lyons, A.B. Hawaii BISH 1860-01-01
8 Psilotum nudum ridge to Puu Piei, Kahana Bay ground beneath dense Brassaia canopy; with Wikstroemia, Bobea, and Osmanthus Preserved_Specimen Takeuchi, W.N. 182 Oahu BISH 1984-05-26
9 Psilotum nudum Makaha Valley, Waianae Mts forest near stream on tree trunk Preserved_Specimen St.John, H. 11588 Oahu BISH 1932-04-03
10 Psilotum nudum ridge to Puu Piei, above Kahana Bay on ground beneath Brassaia; with Wikstroemia, Bobea, and Osmanthus Preserved_Specimen Takeuchi, W.N. 182 Oahu BISH 1984-05-26
11 Psilotum nudum Maakua - Papali Ridge, Koolau Mts, Hauula Preserved_Specimen Occurrence: common along open rocky ridge with scattered brush. Branchlets triangular. Fosberg, F.R. 29700 Oahu BISH 1948-03-21
12 Psilotum nudum Hilo forest Preserved_Specimen Newell, Brother M. Hawaii BISH 1910-01-01
13 Psilotum nudum Small gully, N.W. side Ulumawao, Kailua Steep brushy rock slide Preserved_Specimen Fosberg, F.R. 13562 Oahu BISH 1937-01-24
14 Psilotum nudum Molokai Preserved_Specimen Rock, J.F.C. 6178 Molokai BISH 1910-03-26
15 Psilotum nudum trail to Kalaupapa windswept cliffs on N coast; cracks in rock cliff faces Preserved_Specimen Crosby, M.R. 1667 Molokai BISH 1964-07-04
16 Psilotum nudum Koko Crater, Maunalua Rock crevices in north-facing cliffs. Preserved_Specimen Fosberg, F.R. 9140 Oahu BISH 1933-01-27
17 Psilotum nudum Honolulu, Hawaii Loa Ridge trail. Psidium cattleianum forest along summit trail, Growing at base of Psidium cattleianum with Carex meyenii, Miconia crenata. Preserved_Specimen Kennedy, B. 88 Oahu BISH 2019-04-26
18 Psilotum nudum East Maui, Waihoi Valley, 200' along transect 1 trail on Cheirodendron Preserved_Specimen Harrison, B. 172 Maui BISH 1972-08-04
19 Psilotum nudum Pohakea Pass, Waianae Mts, Honoaluili dry Lantana slope Preserved_Specimen Fosberg, F.R. 9507 Oahu BISH 1933-05-12
20 Psilotum nudum Manoa Arboretum, Haukulu Preserved_Specimen Russell, P. 2 Oahu BISH 1947-07-05
21 Psilotum nudum Kaali Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
22 Psilotum nudum Kauai Preserved_Specimen Forbes, C.N. Kauai BISH
23 Psilotum nudum Kaiholena Preserved_Specimen Munro, G.C. 454 Lanai BISH 1929-08-20
24 Psilotum nudum Hawaii Preserved_Specimen Degener, O. 5719 Hawaii BISH 1927-09-29
25 Psilotum nudum N.Kona, Mamaloah Hwy., near mile 28, Kaupulehu Preseve Preserved_Specimen Flynn, T.W. 1284 Hawaii BISH 1985-10-15
26 Psilotum nudum Preserved_Specimen BISH
27 Psilotum nudum Kamanaiki, Kalihi Preserved_Specimen Hume, E.P. 2 Oahu BISH 1930-10-15
28 Psilotum nudum Waikalaalaua Gulch, Waipio, Koolau Mts Preserved_Specimen Hosaka, E.Y. 194 Oahu BISH 1930-04-06
29 Psilotum nudum Lanihuli ridge trail Preserved_Specimen MacDaniels, L.H. 195 Oahu BISH 1927-01-23
30 Psilotum nudum Puu Hapapa, Waianae Mts Preserved_Specimen Krajina, V.J. 620401107 Oahu BISH 1962-04-01
31 Psilotum nudum Nuuanu-Pauoa ridge Preserved_Specimen Wilbur, R.L. 227 Oahu BISH 1947-09-28
32 Psilotum nudum Puu o Kaeha, Kawela Preserved_Specimen St.John, H. 12483 Molokai BISH 1932-12-23
33 Psilotum nudum Bryan's Mtn. House Trail at Grant Boundry Preserved_Specimen Krajina, V.J. 620520021 Oahu BISH 1962-05-20
34 Psilotum nudum Waimea, Na Pali-Kona forest reserve, Kumuwela road, near gate 5 Preserved_Specimen Chock, A.K. 1012 Kauai BISH 1953-08-23
35 Psilotum nudum Mt. Tantalus Preserved_Specimen Topping, D.L. 2698 Oahu BISH 1923-07-21
36 Psilotum nudum Pupukea, Koolau Range Preserved_Specimen Lyon, H.L. Oahu BISH 1931-07-24
37 Psilotum nudum Palehua/Palikea Trail Preserved_Specimen Bennett, T. 204 Oahu BISH 1982-03-11
38 Psilotum nudum 4.4 mi SE of Manuka State Park, along Hwy. 11 Preserved_Specimen Wagner, W.L. 4894 Hawaii BISH 1983-07-28
39 Psilotum nudum Along Tantalus Trail, Koolau Range, Oahu Preserved_Specimen Cowan, R.S. 226 Oahu BISH 1944-05-27
40 Psilotum nudum Wahiawa Preserved_Specimen Forbes, C.N. Oahu BISH 1909-08-05
41 Psilotum nudum Kilauea, Bird Park Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-08-09
42 Psilotum nudum Maui Preserved_Specimen Forbes, C.N. Maui BISH
43 Psilotum nudum Kaiholena Preserved_Specimen Munro, G.C. Lanai BISH 1913-09-01
44 Psilotum nudum Molokai Preserved_Specimen Stokes, J.F.G. Molokai BISH 1909-01-01
45 Psilotum nudum Olea, Puna Preserved_Specimen Pukui, M.K. Hawaii BISH 1931-01-01
46 Psilotum nudum Waimea Preserved_Specimen Pearsall, G.A. Oahu BISH
47 Psilotum nudum Waiawa Valley. Preserved_Specimen Lyon, H.L. Oahu BISH 1928-08-04
48 Psilotum nudum Lanai Preserved_Specimen Forbes, C.N. Lanai BISH 1913-06-01
49 Psilotum nudum Laie Ridge Preserved_Specimen Suehiro, A. Oahu BISH 1932-02-14
50 Psilotum nudum Kona, above Alahaka Bay, top of long Holua slide Preserved_Specimen Bryan Jr., E.H. Hawaii BISH 1957-01-25
51 Psilotum nudum S ridge of Kipapa Gulch, Waipio, Koolau Mts. Zone 2 Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-05-15
52 Psilotum nudum Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-06-06
53 Psilotum nudum Koolau Preserved_Specimen Rock, J.F.C. Oahu BISH 1908-08-01
54 Psilotum nudum Waiahole Ditch Trail Preserved_Specimen Meebold, A. Oahu BISH 1932-06-01
55 Psilotum nudum Oahu Preserved_Specimen Forbes, C.N. Oahu BISH
56 Psilotum nudum Makaha Valley, Kaala Range, Oahu Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
57 Psilotum nudum Kaiholena Preserved_Specimen Munro, G.C. Lanai BISH 1915-09-01
58 Psilotum nudum Kilauea, floor of Kilaueaiki Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-03
59 Psilotum nudum W Wailupe Preserved_Specimen Wong, C.C.Y. Oahu BISH 1951-06-21
60 Psilotum nudum unknown Preserved_Specimen Mann, H. BISH 1864-01-01
61 Psilotum nudum unknown Preserved_Specimen Mann, H. BISH 1864-01-01
62 Psilotum nudum unknown Preserved_Specimen Ballieu, M. BISH 1875-01-01
63 Psilotum nudum unknown Preserved_Specimen Mann, H. BISH 1864-01-01
64 Psilotum nudum N.Kona, Puuwaawaa; along Huehue-Puuwaawaa pipe line, between water tanks 1-3 Preserved_Specimen Kondo, Y. 111.5 Hawaii BISH 1946-07-24
65 Psilotum nudum Waimea, Kokee, Na Pali-Kona forest reserve Preserved_Specimen St.John, H. 10718 Kauai BISH 1930-12-26
66 Psilotum nudum Coral plain between Ewa & Sisal Preserved_Specimen Forbes, C.N. 2354 Oahu BISH 1916-03-14
67 Psilotum nudum Waimea, Na Pali-Kona forest reserve, Kumuwela road, near gate 5 Preserved_Specimen Chock, A.K. 1012 Kauai BISH 1953-08-23
68 Psilotum nudum Koko Head Preserved_Specimen Shaw, G.W. 2064 Oahu BISH
69 Psilotum nudum Manoa Cliffs Trail, E side of Mt. Tantalus, Koolau Range Preserved_Specimen Ozaki, E.T. 1686 Oahu BISH 1951-07-20
70 Psilotum nudum Palolo-Waialae ridge Preserved_Specimen MacDaniels, L.H. 518 Oahu BISH 1927-01-27
71 Psilotum nudum Kau District. Preserved_Specimen Giffard, W.M. 360 Hawaii BISH 1928-07-01
72 Psilotum nudum Honopu, Awaawapuhi Trail. Preserved_Specimen Higgins, N.H. 123 Kauai BISH 1956-12-29
73 Psilotum nudum Waimano Valley Preserved_Specimen Topping, D.L. 2747 Oahu BISH 1924-02-17
74 Psilotum nudum Milolii ridge Preserved_Specimen Cranwell, L.M. 2921 Kauai BISH 1938-08-14
75 Psilotum nudum Kula, Maui Preserved_Specimen Degener, O. 5713 Maui BISH 1926-10-23
76 Psilotum nudum In nursery near Koko Head, Oahu, Hawaii Preserved_Specimen Buchanan, H. 572 Oahu BISH 1961-05-30
77 Psilotum nudum Kamalo Gulch Preserved_Specimen Forbes, C.N. 589 Molokai BISH 1912-09-01
78 Psilotum nudum Mountains near Koele Preserved_Specimen Forbes, C.N. 173 Lanai BISH 1913-06-01
79 Psilotum nudum S of Hilo Preserved_Specimen Krajina, V.J. 611228042 Hawaii BISH 1961-12-28
80 Psilotum nudum Koko Head Preserved_Specimen Shaw, G.W. 5839 Oahu BISH
81 Psilotum nudum Wahiawa gulch Preserved_Specimen Rock, J.F.C. 19 Oahu BISH 1908-07-01
82 Psilotum nudum Wailupe Valley Preserved_Specimen Forbes, C.N. 1577 Oahu BISH 1910-04-12
83 Psilotum nudum Ridge between Pahole and Kapuna Gulches Preserved_Specimen Welton, P. 663 Oahu BISH 1991-01-27
84 Psilotum nudum Koolau Mts, Paalaa Uka, near Poamoho Trail Preserved_Specimen Krajina, V.J. 6110 Oahu BISH 1961-01-01
85 Psilotum nudum Kawaiiki - Ditch trail Preserved_Specimen Woolford, H.A. 14 Oahu BISH 1955-10-09
86 Psilotum nudum East Maui, Hana Preserved_Specimen Handy, E.S. 334 Maui BISH 1933-06-06
87 Psilotum nudum Honolulu, Bishop Museum Campus, native Hawaiian garden. Preserved_Specimen Kennedy, B. 20179 Oahu BISH 2017-11-09
88 Psilotum nudum unknown Preserved_Specimen BISH
89 Psilotum nudum unknown Preserved_Specimen Hillebrand, W.B. BISH
90 Psilotum nudum Makiki Preserved_Specimen Mann, H. Oahu BISH 1864-01-01
91 Psilotum nudum East Maui, Hana District, Kipahulu Valley, W part of valley, Puu Palikea terrestrial Preserved_Specimen Lamoureux, C.H. 4056 Maui BISH 1967-08-10
92 Psilotum nudum Kawaiiki Valley, Kawailoa Mesic koa and ohia-lehua forest Preserved_Specimen Grady, G.L. 1045 Oahu BISH 1947-11-02
93 Psilotum nudum pali above peninsula W of Wailau Valley among brush on face Preserved_Specimen Fosberg, F.R. 9660 Molokai BISH 1954-07-04
94 Psilotum nudum Kahuku, Ocean View Estastes, near jct of Moana Dr. and Tiki Lane arid aa lava; at bases of Metrosideros trees Preserved_Specimen clumps Herbst, D.R. 5073 Hawaii BISH 1974-10-10
95 Psilotum nudum National Park, beginning of Mauna Loa scenic strip road, off trail to weather station bark of Metrosideros, shaded Preserved_Specimen epiphyte Herat, T. 620 Hawaii BISH 1974-06-19
96 Psilotum nudum Kaaliwai With Adiantum, in spring on cliff Preserved_Specimen Large tufts St.John, H. 23574 Niihau BISH 1949-03-29
97 Psilotum nudum North of Wilhelmina Rise, Oahu Exposed ridge Preserved_Specimen Degener, O. 3516 Oahu BISH 1926-04-24
98 Psilotum nudum Pohakea Pass, Waianae Mts., Halona, Lualualei Forest Reserve tree crotch Preserved_Specimen tuft, 3' in diam. St.John, H. 13135 Oahu BISH 1933-05-12
99 Psilotum nudum E of Mapulehu Valley, near ridge W of Pukoo Valley trail on ridge Preserved_Specimen St.John, H. 12916 Molokai BISH 1932-12-29
100 Psilotum nudum Puna, 1/4 mile e of Kapoho Crater basalt crust in open field Preserved_Specimen St.John, H. 22296 Hawaii BISH 1946-12-22
101 Psilotum nudum S ridge, Kipapa Gulch, Waipio, Koolau Mts on tree trunk, moist woods Preserved_Specimen Hosaka, E.Y. 548 Oahu BISH 1932-05-15
102 Psilotum nudum Mauna Loa slope, partly shaded aa lava Preserved_Specimen Degener, O. 68 Hawaii BISH 1922-08-25
103 Psilotum nudum jct. of Kulani and 15.75 mi Rds., Waiakea, S. Hilo wet Metrosideros and Cibotium, on mossy trunks Preserved_Specimen St.John, H. 22369 Hawaii BISH 1946-12-24
104 Psilotum nudum Pahole Gulch, Waialua, Waianae Mts steep, well drained slope near crest in west section. Sunny; with Peperomia tetraphylla, Metrosideros, Styphelia, Acacia koa Preserved_Specimen Nagata, K.M. 1360 Oahu BISH 1976-07-27
105 Psilotum nudum Puna flow of 1840, near RR track Preserved_Specimen Forbes, C.N. 1060 Hawaii BISH 1915-07-07
106 Psilotum nudum Honolulu Forest Reseve, Manoa Cliffs Trail wet forest Preserved_Specimen terrestrial Lammers, T.G. 5326 Oahu BISH 1983-08-27
107 Psilotum nudum Maunapuluo, Hanakapiai, Napali Coast under tree Preserved_Specimen St.John, H. 10890 Kauai BISH 1930-12-30
108 Psilotum nudum Hanapepe river valley base of cliff, open slope Preserved_Specimen St.John, H. 13521 Kauai BISH 1933-12-23
109 Psilotum nudum W ridge of Kaukaua Gulch, Kipahulu summit, koa woods, on base of koa tree Preserved_Specimen St.John, H. 17811 Maui BISH 1936-12-28
110 Psilotum nudum Site 08 OS, Olaa, Olaa Forest Reserve. Ohia-treefern dieback. Primary vegetation: Metrosideros polymorpha, Coprosma spp., Ilex anomala, Gouldia sp., Cheirodendron trigynum, Psychotria hawaiiensis, Psidium cattleianum, Pritchardia sp., Soils are a deep, moderately well drained ash over an unknow substrate. Preserved_Specimen Adee, K.T. Hawaii BISH
111 Psilotum nudum ridge to Puu Piei, above Kahana Bay on ground beneath Schinus terebinthifolius Preserved_Specimen erect Takeuchi, W.N. 105 Oahu BISH 1984-03-11
112 Psilotum nudum N.W. ridge Puu Olomana, Kailua dry rocky ridge Preserved_Specimen Fosberg, F.R. 9034 Oahu BISH 1932-11-13
113 Psilotum nudum Hulopoe Gulch, Mahana on ground in thicket Preserved_Specimen St.John, H. 18749 Lanai BISH 1938-04-09
114 Psilotum nudum trail along Pauoa Ridge on trees Preserved_Specimen Crosby, M.R. 1352 Oahu BISH 1964-06-13
115 Psilotum nudum Aiea Trail, Kalauao-North Halawa, Koolau Mountains Dry forest. Preserved_Specimen epiphytic Higgins, N.H. 71 Oahu BISH 1956-01-08
116 Psilotum nudum East Maui, 0.9 mi E of Ulupalakua among aa lava with Bryophyllum and Cocculus in dry area Preserved_Specimen erect, shrubby, with green stems, yellow (when mature) spore cases Herbst, D.R. 852 Maui BISH 1968-01-04
117 Psilotum nudum Hauula, off the foot trail, at end of jeep trail around the ridge on cibotium trunk Preserved_Specimen Herat, T. 171 Oahu BISH 1972-08-16
118 Psilotum nudum Site 10 OS, Piihonua, Hilo Forest Reserve. Ohio-koa dieback. Primary vegetation: Acacia koa, Metrosideros polymorpha, Psidium cattleianum, Cheirodendron trigynum, Antidesma platyphyllum, Melastoma malabathricum and Clermontia sp. Soils are a deep, somewhat poorly drained ash over an unknown substrate. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-20
119 Psilotum nudum East Maui, Hana District, Haleakala, Kaupo Gap on old eroded lava flows Preserved_Specimen 7.5 tall Olson, G.E. 26 Maui BISH 1937-12-09
120 Psilotum nudum Maakua-Papali Ridge, Hauula, Koolau Mts. Open dry ridge. Preserved_Specimen Kitamura, F. Oahu BISH 1933-10-15
121 Psilotum nudum Kamokuiki Valley on mossy bank with liverworts Preserved_Specimen Erect herb, Height: 30 cm Bryan Jr., E.H. 766 Oahu BISH 1933-04-13
122 Psilotum nudum Kaneohe, slopes above Haiku Plantation northern exposure Preserved_Specimen Stemmermann, R.L. 409 Oahu BISH 1973-07-23
123 Psilotum nudum Pupukea Ridge Kaolite clay; grassland, by a post. Preserved_Specimen epiphyte Lee, J. 3 Oahu BISH 1981-10-01
124 Psilotum nudum W slope of Manuahi Ridge, SE of Keawanui secondary forest of Psidium guajava, Schinus, Pandanus tectorius, Metrosideros and Aleurites, ground cover of Kalanchoe pinnata and Zingiber zerumbet; in fork of a tree Preserved_Specimen epiphyte Lorence, D.H. 5434 Molokai BISH 1987-08-01
125 Psilotum nudum W ridge of Honomuni moist woods Preserved_Specimen St.John, H. 25174 Molokai BISH 1953-12-23
126 Psilotum nudum S.Kona, SW slopes of Mauna Loa 1856 flow Preserved_Specimen Krajina, V.J. 611228053 Hawaii BISH 1961-12-28
127 Psilotum nudum Wainiha Valley, stream below Hono o Na Pali on decaying trunk of Hibiscus tiliaceus, in thicket Preserved_Specimen Wilson, K.A. 229 Kauai BISH 1952-12-31
128 Psilotum nudum Kapalama Valley, Koolau Range arid scrub, dry ridge, in crevice of large rock Preserved_Specimen Kuykendall, J.R. 15 Oahu BISH 1942-11-01
129 Psilotum nudum East Maui, Kahuai, Ulupalakua-Hana Rd., Mokulau, opposite Hui Aloha church E of Kaupo ancient heiau Preserved_Specimen Chock, A.K. 335 Maui BISH 1951-12-27
130 Psilotum nudum National Park, Kau, beginning of Halape trail deep crack Preserved_Specimen Herat, T. 818 Hawaii BISH 1974-06-29
131 Psilotum nudum Koolau Mts., Lanipo Trail rocks on steep, windswept, dry ridge; with grasses and small shrubs Preserved_Specimen Herbst, D.R. 1398 Oahu BISH 1969-04-11
132 Psilotum nudum East Maui, ridge E of Kaupo Gap, Kipahulu forest reserve grass and Styphelia Preserved_Specimen Higashino, P.K. 2573 Maui BISH 1976-06-30
133 Psilotum nudum Kokee, on trail to Lookout In moss on rotten log Preserved_Specimen Alexander, A.M. 5199 Kauai BISH 1947-03-08
134 Psilotum nudum slope above Kaimuki dry ground Preserved_Specimen MacDaniels, L.H. 106 Oahu BISH 1926-11-01
135 Psilotum nudum Maakua-Papali ridge, Kaipapau Forest Reserve, Haaula, Koolau Mts forest Preserved_Specimen Onouye, R. Oahu BISH 1933-10-15
136 Psilotum nudum Kalalau trail, Waiahuakua Valley, on Napali Coast crevices of rocks, dryer areas Preserved_Specimen Stone, B.C. 747 Kauai BISH 1955-06-17
137 Psilotum nudum Munro Trail, N end, 2.2 mi in from Hwy Near bases of planted Eucalyptus forest with Schinus and Psidium Preserved_Specimen small, in clumps. Stems angled; dicotomously branched. Leaves reduced to scales. Spore cases 3 lobed in upper leaf axils Spence, G. 334 Lanai BISH 1973-10-22
138 Psilotum nudum ~200 m down Hilina Pali Rd. from intersection with Chain of Craters Rd., plot 1 Metrosideros, native shrub, Andropogon Preserved_Specimen ~6 tall Newell, C.L. 416 Hawaii BISH 1966-08-15
139 Psilotum nudum Tantalus, Oahu Exposed slope Preserved_Specimen Degener, O. 2070 Oahu BISH 1926-03-20
140 Psilotum nudum Red Hill exposed rocks Preserved_Specimen Gillespie, J.W. 1056 Oahu BISH 1927-01-01
141 Psilotum nudum Olomana In Lantana shrubbery Preserved_Specimen Christophersen, E. Oahu BISH 1924-10-26
142 Psilotum nudum S. ridge of Kipapa Gulch, E of Puu Kamana, Waipio, Koolau Mts wooded, on mossy trunk Preserved_Specimen St.John, H. 11668 Oahu BISH 1932-05-15
143 Psilotum nudum Aiea ridge Preserved_Specimen epiphyte Kato, M. 1033 Oahu BISH 1998-08-29
144 Psilotum nudum Kilauea disintergrating lava Preserved_Specimen Degener, O. 1545 Hawaii BISH 1922-12-20
145 Psilotum nudum W.side Puu Kalina, Waianae Mts, Waianae Kai crevices in face of precipice Preserved_Specimen Fosberg, F.R. 9245 Oahu BISH 1933-03-12
146 Psilotum nudum 6 mi N of Manuka Park, Kau in cracks of lava blocks, at margins of dry Metrosideros polymorpha forest with Aleurites moluccana, Dodonaea viscosa, Styphelia tameiameiae, Sophora chrysophylla, Acacia koa Preserved_Specimen tufts, axes yellow-green Carlquist, S. 2067 Hawaii BISH 1966-07-26
147 Psilotum nudum Kamehameha Girl's School, Kapalama, Koolau Range side of a boulder Preserved_Specimen Hosaka, E.Y. 2427 Oahu BISH 1938-12-10
148 Psilotum nudum Kaluanui Flats, mauka shaded slope Preserved_Specimen Degener, O. 30109 Lanai BISH 1964-01-24
149 Psilotum nudum Kwaiiki rock face, Diospyros zone Preserved_Specimen Takeuchi, W.N. 3633 Oahu BISH 1987-07-18
150 Psilotum nudum East Maui, Haleakala near Ulupalakua in 1675 lava flow Preserved_Specimen Krajina, V.J. 620421003 Maui BISH 1992-02-01
151 Psilotum nudum Puuwaawaa, near the Hululhulu escarpment, T16 in aa cracks Preserved_Specimen terrestrial Takeuchi, W.N. 5776 Hawaii BISH 1989-12-26
152 Psilotum nudum Wailupe, Kului in deep shade, under waiawi tree Preserved_Specimen Egler, F.E. 3798 Oahu BISH 1937-06-23
153 Psilotum nudum Laaukahi, Koloa, NW facing slope, 3/4 mi N moist wooded gulch Preserved_Specimen on ground St.John, H. 23004 Kauai BISH 1947-12-24
154 Psilotum nudum National Park, Kipuka Ki on dry aa lava Preserved_Specimen Webster, G.L. 1710 Hawaii BISH 1948-06-16
155 Psilotum nudum NW end of Alakai Swamp, Na Pali-Kona Forest Reserve open ridge Preserved_Specimen St.John, H. 10745 Kauai BISH 1930-12-27
156 Psilotum nudum Hanalei, Wainiha River valley just W of Wainiha, along Power House Rd. ~ 300 m S of Hwy. 56 secondary wet forest and shrubland with Psidium cattleianum, Casuarina, Syzygium cumini; overgrown with Passiflora laurifolia, ground cover of alien grasses; in fork of Syzygium cumini Preserved_Specimen epiphyte, stems erect, bright green. Ripe sporangia yellow Lorence, D.H. 7667 Kauai BISH 1995-05-13
157 Psilotum nudum Kapua on weathered aa, with Lantana Preserved_Specimen 20 cm tall Bryan Jr., E.H. 751 Hawaii BISH 1932-11-24
158 Psilotum nudum Honolulu, 1940 Manoa Rd cement pot, in soil, full sun Preserved_Specimen Lau, J. 1499 Oahu BISH 1985-10-07
159 Psilotum nudum Koloa, Wahiawa Mt., E side along Kuai stream mature secondary forest with Psidium, Kukui, Metrosideros, and Freycinetia, in fork of Psidium Preserved_Specimen epiphyte forming dense clone; rhizome showing lateral branching Lorence, D.H. 5186 Kauai BISH 1987-04-11
160 Psilotum nudum Former Naval Air Station Barbers Point, Northern Trap & Skeet Range On bare scraped limestone. Under Pluchea. With Waltheria indica, Stachytarpheta jamaicensis, Leucaena leucocephala, Casuarina equisetifolia and Prosopis palida. Preserved_Specimen In dense clumps. Stems green turning yellow-orange at tips. BISH staff 41 Oahu BISH 2021-05-04
161 Psilotum nudum Puulani-Ahupua between rocks of ancient heiau Preserved_Specimen Wilson, K.A. 97 Maui BISH 1951-12-27
162 Psilotum nudum North Kohala District, Kalala ahupuaa, Lahikiola cinder cone in understory of Metrosideros woodland Preserved_Specimen Herb Bunch, P.M. 102 Hawaii BISH 1989-03-13
163 Psilotum nudum National Park, close to Chain of Craters road, NE of Luu Manu crater base of Metrosideros tree Preserved_Specimen Herat, T. 1020 Hawaii BISH 1974-07-15
164 Psilotum nudum Kau District, Ainapo to Puu Pili, Kapapala exposed basalt ledges Preserved_Specimen St.John, H. 18435 Hawaii BISH 1937-12-21
165 Psilotum nudum Maakua-Papali ridge, Hauula, Kaipapau forest reserve tree trunk near wet forest Preserved_Specimen small fern Yoshioka, T. Oahu BISH 1933-10-15
166 Psilotum nudum Peacock Flats In Eucalyptus Forest above campgrounds. Preserved_Specimen Welton, P. 69 Oahu BISH 1988-11-13
167 Psilotum nudum Makaha Valley, Waianae Mts rainforest on ridge, on tree trunk Preserved_Specimen St.John, H. 25118 Oahu BISH 1953-05-03
168 Psilotum nudum Pupukea, Honolulu, Hawaii 15 % slope, loam Preserved_Specimen veg type: Eucalyptus, ferns, rose apple. branches all flattened, no roots proper. dist: found in high forest. Aki, J. 33 Oahu BISH 1969-09-26
169 Psilotum nudum S. Kona; Kapua dry aa fields in the open Preserved_Specimen erect Russ, G.W. 98 Hawaii BISH 1982-08-01
170 Psilotum nudum In crevices of rocks (lava blocks). Along Waahila Ridge Trail (= Mt. Olympus Trail), between Manoa and Palolo Valleys, Koolau Mts in sunny, grassy area of ridge top. With Sphenomeris, Dicranopteris emarginata, Bidens, introduced weeds (Psidium, Lantana). Preserved_Specimen Carlquist, S. 1632 Oahu BISH 1964-11-29
171 Psilotum nudum Hanalei District. Kalalau Valley, southwest side (back) of valley. Diospyros mixed mesic forest. W/Diospyroas sandwicensis, D. hillebrandii, Antidesma, Bobea elatior, Planchonella sandwicensis, Metrosideros polymorpha var glaberrima, Psydrax odorata, Pisonia sandwicensis, Aleurites moluccana, Fre Preserved_Specimen Epiphyte about 8 feet up in an Antidesma. Fertile with sporanigia. Walsh, S. 640 Kauai BISH 2018-10-12
172 Psilotum nudum SW of Haena; 3/4 mile in on Na Pali Coast Trail in recesses in black lava, along coast; with Pandanus, Setaria Preserved_Specimen Henrickson, J.S. 3993 Kauai BISH 1969-07-31
173 Psilotum nudum Maunapuluo, Hanakapiai, Napali Coast open rocky slope Preserved_Specimen St.John, H. 10896 Kauai BISH 1930-12-30
174 Psilotum nudum Between Maakua and Papali Gulches, Hauula Forest Reserve, Koolau Mts dry ridge Preserved_Specimen Morley, H. 139 Oahu BISH 1933-10-15
175 Psilotum nudum Lualualei, Waianae dryland forest Preserved_Specimen Takeuchi, W.N. Oahu BISH 1992-09-01
176 Psilotum nudum Palolo-Waialaenui Ridge, Koolau Mts.; lowest saddle among boulders Preserved_Specimen Hartung, M. Oahu BISH 1936-08-24
177 Psilotum nudum W side of Nonou, Wailua moist wooded gulch Preserved_Specimen St.John, H. 24938 Kauai BISH 1952-12-31
178 Psilotum nudum Kawaiiki Ditch Trail, Kawailoa, Koolau Mts rain forest, on bank side of trail Preserved_Specimen Wilson, K.A. 3 Oahu BISH 1951-10-14
179 Psilotum nudum Kalaupapa; Kauhako crater, SE Preserved_Specimen erect terrestrial fern Wysong, M.L. 515 Molokai BISH 2004-12-07
180 Psilotum nudum Pupukea-Paumalu Forest Reserve, Kaunala Gulch on tree trunk Preserved_Specimen Hosaka, E.Y. 2581 Oahu BISH 1941-02-08
181 Psilotum nudum Kaiholena Preserved_Specimen Munro, G.C. 83 Lanai BISH 1913-09-01
182 Psilotum nudum Napali cliff trail beyound Haena rocks above Preserved_Specimen Alexander, A.M. 5273 Kauai BISH 1947-03-15
183 Psilotum nudum W of Sea Arch, ~1 mi E of Kaena Point, Pauau deep crevices on almost bare pahoehoe lava Preserved_Specimen erect, tips yellow Fosberg, F.R. 46014 Hawaii BISH 1965-03-19
184 Psilotum nudum 1st gulch W of Manini Gulch, Kaena on ledge Preserved_Specimen St.John, H. 22868 Oahu BISH 1947-11-11
185 Psilotum nudum Kapaka Valley, the first valley E of Mokouia Valley Wet ledge by spring Preserved_Specimen St.John, H. 22840 Niihau BISH 1947-08-16
186 Psilotum nudum E part of Kaluaaha plateau among rocks in woods Preserved_Specimen Degener, O. 5710 Molokai BISH 1928-07-03
187 Psilotum nudum N mauka of City in patch of decadent forest Preserved_Specimen Degener, O. 22015 Lanai BISH 1952-03-02
188 Psilotum nudum Halawa Valley dry rocky ridge Preserved_Specimen St.John, H. 12705 Molokai BISH 1932-12-27
189 Psilotum nudum S ridge Kipapa Gulch, Waipio. Koolau Mts on tree trunk Preserved_Specimen St.John, H. 9969 Oahu BISH 1929-10-29
190 Psilotum nudum S end of Kau desert, 2 mi from end of Hilina Pali Rd crack in lava Preserved_Specimen Iltis, H.H. 134 Hawaii BISH 1967-06-22
191 Psilotum nudum Mountain View; Kazumura Lava Tube on floor in dark Preserved_Specimen Stone, F.D. Hawaii BISH 1985-01-20
192 Psilotum nudum Hulopoe and Kaiholena Gulches, ridge between gulches in brush and grass at edge of eroded area Preserved_Specimen Fosberg, F.R. 12368 Lanai BISH 1935-11-28
193 Psilotum nudum Waimea, Mahanaloa Valley, Kopiwai trail on Perrottetia sandwicensis; about 10 ft. up. Preserved_Specimen epiphyte Doty, M.S. 1 Kauai BISH 1952-02-05
194 Psilotum nudum Tantalus under rocks and trees Preserved_Specimen Meebold, A. 8324 Oahu BISH 1930-02-01
195 Psilotum nudum Kamoamoa inland; in cracks, deep in shade Preserved_Specimen Stone, B.C. 2879 Hawaii BISH 1958-06-20
196 Psilotum nudum East Maui, Hana District, Waihoi Valley, S fork of Waiohonu on trunk of Metrosideros Preserved_Specimen Harrison, B. 106 Maui BISH 1972-07-27
197 Psilotum nudum Honolulu, 1940 Manoa Rd cement pot, in soil, full sun Preserved_Specimen Lau, J. 1499 Oahu BISH 1985-10-07
198 Psilotum nudum Kalihi Valley, Koolau Range between rocks; dry Lantana zone Preserved_Specimen Nishida, T. Oahu BISH 1943-12-09
199 Psilotum nudum Foster Garden Kaolite clay. Preserved_Specimen epiphyte Lee, J. 15 Oahu BISH 1991-11-20
200 Psilotum nudum East Maui, Manawainui exotic grasses, Aleurites and Psidium Preserved_Specimen Higashino, P.K. 2261 Maui BISH 1976-06-25
201 Psilotum nudum Kau, Kahulai 1887 flow, dry crevices of bare aa lava Preserved_Specimen St.John, H. 22422 Hawaii BISH 1946-12-27
202 Psilotum nudum E branch, E Makaleha Stream, Mokuleia, Waianae Range dry forest Preserved_Specimen terrestrial Hatheway, W.H. 425 Oahu BISH 1950-07-02
203 Psilotum nudum Pupukea Ridge, fence row of a pasture Slope: 10%, Soil: Kaolite Clay, veg type: tropical grassland Preserved_Specimen Herbacious (Fern Allies), Dist: state-wide, wet, moist place Fafai, L. 2 Oahu BISH 1981-10-01
204 Psilotum nudum Head of east side of Kamilonui Valley dry rocky ledges Preserved_Specimen Fosberg, F.R. 13607 Oahu BISH 1937-03-07
205 Psilotum nudum Hauula, Koolau Range, Maakua-Papali ridge in dense clusters of growth on ground Preserved_Specimen mature sporangia yellow, 3 celled Wilson, K.A. 148 Oahu BISH 1952-10-26
206 Psilotum nudum Keana Bishopa Kipuka, above Hilina Pali, National Park on Metrosideros trunk Preserved_Specimen epiphyte Fosberg, F.R. 56733 Hawaii BISH 1977-03-21
207 Psilotum nudum Kuliouou 2, Koolau Range mossy rock in woods Preserved_Specimen St.John, H. 26914 Oahu BISH 1978-11-26
208 Psilotum nudum lower Kipahulu Valley, Palakea Stream area between upper pasture and gauging station Preserved_Specimen Gagné, B.H. Maui BISH 1975-03-01
209 Psilotum nudum E side of Nuuanu Valley dry ridge; on rocks Preserved_Specimen Grether, D.F. 3900 Oahu BISH 1945-10-09
210 Psilotum nudum Kalauao - Waimalu Ridge, Koolau Mts Wet forest, back of trees Preserved_Specimen Fosberg, F.R. 9340 Oahu BISH 1933-03-29
211 Psilotum nudum Mokolii Islet, Kualoa In cactus thicket. Preserved_Specimen Fosberg, F.R. 13568 Oahu BISH 1937-01-31
212 Psilotum nudum U.S. Naval Communications Center, east of Wahiawa Preserved_Specimen Occurence: Rare, one clump seen, terrestrial in forest in deep, steep-walled wooded gulch. Fosberg, F.R. 65119 Oahu BISH 1985-10-28
213 Psilotum nudum by Visitor Center and main ruins, Puuhonua o Honaunau Park exotic vegetation; modified by ancient Polynesians Preserved_Specimen Higashino, P.K. 10271 Hawaii BISH 1984-04-08
214 Psilotum nudum Waikolu Valley below Puu Kaeo steep wall, wet forest, on mossy ground Preserved_Specimen St.John, H. 23471 Molokai BISH 1948-12-22
215 Psilotum nudum Nuuanu-Pauoa ridge introduced weeds and grasses Preserved_Specimen Grady, G.L. 1001 Oahu BISH 1947-09-28
216 Psilotum nudum Kawaiiki, Oahu. In clefts of cliff Preserved_Specimen Degener, O. 10323 Oahu BISH 1936-02-09
217 Psilotum nudum Hulopoe Gulch, head of, at Munro Trail Preserved_Specimen epiphytic Degener, O. 28517 Lanai BISH 1963-08-01
218 Psilotum nudum Palolo Valley Preserved_Specimen Garber, D.W. 39 Oahu BISH 1919-11-02
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