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Haplostachys haplostachya

(A.Gray) H.St.John (1973)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Lamiaceae Genus: Haplostachys

honohono

Description

Key Characters:

Growth Form: Erect perennial herbs.

Stems: Stems quadrangular, 3–6 dm long, densely white tomentose.

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly cordate, 8–17 cm long, 3–5.5 cm wide. Apex obtuse to acute. Upper surfaces densely puberulent, green, rugose; lower surfaces densely white tomentose; blades somewhat fleshy. Margins crenate. Petioles 2.5–5 cm long. Stipules absent.

Flowers: Flowers subsessile, in simple, often dense, terminal racemes, 2 flowers per verticillaster, 10–30 cm long, bracts oblong-lanceolate, usually 4–6 mm long, lowest pair up to 20 mm long. Calyx actinomorphic, cylindrical, closed at maturity or nearly so, 8–12 mm long, densely tomentose, 5-toothed, the teeth linear (2–)3–5 mm long, splitting longitudinally, usually persistent or sometimes only ventral 1/2 persistent. Corolla zygomorphic, bilabiate, white or sometimes tinged purple externally; tube narrowly funnelform, somewhat curved, ca. 14–22 mm long, upper lip 8–10 mm long, lower lip 10–14 mm long, the lobes suborbicular, middle lobe largest, margins strongly undulate-crisped; nectary disk often present at base of ovary, annular or developed on anterior side only. Stamens 4, both pairs of similar length or the upper pair slightly longer, ascending under the upper lip, included; filaments inserted near the throat; anthers dithecal; anther sacs divaricate, opening by longitudinal slits. Ovary superior, 2-carpellate, placentation basal-axile; ovules 1 per ovary lobe; style 1, included, shortly 2-lobed, the lobes clavate, pubescent in upper part; stigmatic only at the truncate apex, 1 lobe often rudimentary; stigmas 2.

Fruit: Nutlets 1-seeded; dark brown; ca. 3.5–4 mm long; truncate at both ends; apex sparsely tomentose; enclosed in the persistent closed calyx; pericarp thick and hard. Seeds with endosperm absent or scanty and oily.

Ploidy:

Habitat: Formerly known from barren ridges; Kaua'i; sandy isthmus and Kula; Maui; and ; Nohonaohae Cinder Cone; Pu'ukapele; and Waiki'i; Hawai'i. Presently known only from Kipukakālawamauna; Hawai'i.

Elevation Range: 990–2,135 m.

Historical Distribution

Uses and Culture

USES

Ethnobotanical Images

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Natural History

Statewide Status

Endemic

Island Status

Kaua'i Extirpated
Maui Extirpated
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: List & Summary Fl. Pl. Hawaiian Is.: 292 (1973)

Other References

Wagner et al. 1990:799 (K, M, H); Wilkes s.n. US 57479 (K†, 1840); Hank Oppenheimer pers. obs./Hillebrand s.n. BISH 53186 (M†, 1871)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood 553 Hawaii US 1/30/1991
2 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN D. H. Lorence 8782 Kauai US 11/12/2001
3 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN f. r. fosberg 60618 Hawaii US 9/10/1980
4 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Steve Bergfeld s.n. Hawaii PTBG 2/12/1991
5 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lorence, D. H. 8782 Kauai US 11/12/2001
6 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman 14328 US
7 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN f. r. fosberg 60607 Maui US 9/10/1980
8 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wilkes Explor. Exped. s.n. Kauai US 1838-00-00
9 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Several hundred plants along road, PRESERVED_SPECIMEN Wood, K. R.; Obata, J.; Bergfeld, S.; Stormont, B. 553 Hawaii US 1/30/1991
10 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Fosberg, F. R. 60607 Hawaii US 9/10/1980
11 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman & m. castilio 14328 Hawaii US 8/5/1994
12 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN d. h. lorence 8782 Kauai US 11/12/2001
13 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wilkes Explor. Exped. s.n. Maui US 1838-00-00
14 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN wilkes explor. exped. s.n. Maui US 1838-08-08
15 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Fosberg, F. R. 60618 Hawaii US 9/10/1980
16 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Castilio, M. 14328 Hawaii US 8/5/1994
17 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Sunny slopes of cinder cone. PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 511 Hawaii BISH 4/13/1943
18 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Dry forest; old aa lava vlow overrun by Pennisetum. Assoc. with Tagetes minuta, Myoporum sandwicense, Dodonaea, Sida, Helichrysum foetidum, etc. PRESERVED_SPECIMEN Gustafson, R. 1694 Hawaii BISH 5/21/1980
19 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information On extremely arid aa lava. PRESERVED_SPECIMEN degener, o. greenwell, a.; murashige, t. collector number: 19957 Hawaii BISH 8/25/1949
20 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C. N. 461.H Hawaii US 8/11/2020
21 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Ken Marr KM 209 Hawaii PTBG 11/24/1988
22 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Brackish pond back of sand beach. Growing in 1ft of water. PRESERVED_SPECIMEN wilkes explor. exped. s.n. Kauai BISH 1838-08-08
23 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information On extremely arid aa lava. PRESERVED_SPECIMEN Degener, O. 19957 Hawaii BISH 8/25/1949
24 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s.n. Maui BISH
25 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 461.h Maui BISH 8/8/1911
26 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Dry forest; old aa lava vlow overrun by Pennisetum. Assoc. with Tagetes minuta, Myoporum sandwicense, Dodonaea, Sida, Helichrysum foetidum, etc. PRESERVED_SPECIMEN gustafson, r. collector number: 1694 Hawaii BISH 5/21/1980
27 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information In Dodonaea viscosa mixed shrubland with Myoporum, Chenopodium oahuense, Sida fallax, Bidens menziesii, Eragrostis atropioides, Pennisetum setaceum. PRESERVED_SPECIMEN Perlman, S. 14328 Hawaii BISH 8/5/1994
28 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information ohia PRESERVED_SPECIMEN warshauer, f.r. herbst, d.r.; hohn, j. collector number: frw 2167 Kauai BISH 1/12/1979
29 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lorence, D.H. 8782 Kauai BISH 9/12/2001
30 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Open Euphorbia forest. PRESERVED_SPECIMEN Warshauer, F.R. FRW 2167 Hawaii BISH 1/12/1979
31 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Vegetation: Euphorbia forest. PRESERVED_SPECIMEN Lamoureux, C.H. 4941 Hawaii BISH 1/6/1977
32 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Vegetation: Euphorbia forest. PRESERVED_SPECIMEN lamoureux, c.h. collector number: 4941 Hawaii BISH 1/6/1977
33 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lorence, D.H. Collector Number: 8782 Kauai BISH 9/12/2001
34 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 461.H Hawaii BISH 8/1/1911
35 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN c. n. forbes 461.h F
36 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN e. j. rémy 394 ILL
37 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN j. f. c. rock 8350 NY 6/7/1910
38 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN o. degener 19957 NY 8/25/1949
39 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Slopes PRESERVED_SPECIMEN c. n. forbes 461h NY 8/7/1911
40 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN j. f. c. rock 8350 NY 6/7/1910
41 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Cultivated. PRESERVED_SPECIMEN David H. Lorence 8782 Kauai PTBG 9/12/2001
42 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information in open, intermediate-stature forest. PRESERVED_SPECIMEN k. r. wood, j. obata, s. bergfeld & b. stormont 553 Hawaii BISH 1/30/1991
43 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 8350 Hawaii BISH 6/1/1910
44 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN lorence, d.h. collector number: 8782 Kauai BISH 9/12/2001
45 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information In Dodonaea viscosa mixed shrubland with Myoporum, Chenopodium oahuense, Sida fallax, Bidens menziesii, Eragrostis atropioides, Pennisetum setaceum. PRESERVED_SPECIMEN perlman, s.p. castilio, m. collector number: 14328 Hawaii BISH 8/5/1994
46 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN collector number: 369 Maui BISH
47 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN 369 Maui BISH
48 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information Sunny slopes of cinder cone. PRESERVED_SPECIMEN Fagerlund, G.O. 511 Hawaii BISH 4/13/1943
49 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 8350 Hawaii BISH 6/1/1910
50 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information In Dodonaea viscosa mixed shrubland. PRESERVED_SPECIMEN Steve Perlman 14328 Hawaii PTBG 8/5/1994
51 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 553 Hawaii PTBG 1/30/1991
52 Haplostachys haplostachya (A.Gray) H.St.John Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN wilkes explor. exped. s.n. Kauai US 1838-08-08
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