Cyperus hypochlorus

Hillebr. (1888)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Cyperaceae Genus: Cyperus

‘ahu‘awa [ahuawa]

Description

Key Characters:

Growth Form: Perennials with short, woody rhizomes clothed with dark reddish brown or purplish brown scales and their fibrous remnants.

Stems: culms stout, 50–130 cm tall, 5–16 mm in diameter near base, trigonous, smooth.

Roots:

Leaves: Leaves simple. Leaves several. Blades linear as long as or slightly longer than the culm, 3–15 mm wide. Blades coriaceous. Margins entire. Sheaths dark reddish brown. Stipules absent.

Flowers: Flowers in inflorescences umbelliform, compound or partially compound, moderately dense, 5–18 cm long, 7–25 cm wide, rays 6–12, spreading, 3–16 cm long, once branched, each branch with 2–4 secondary rays, spikes cylindrical to oblong–elliptic, 2.5–4 cm long, 1.2–2 cm wide, densely bearing numerous spikelets, bracteoles setaceous, shorter than to as long as the subtending lateral spikes; involucral bracts 5– 10, the lower ones much longer than the inflorescence and up to 12 mm wide; spikelets 3–8–flowered, widely spreading to slightly reflexed, linear-lanceoloid, subterete, slightly turgid, 6–16 mm long, 1.5– 2.5 mm wide, rachilla broadly winged; glumes green and often tinged pale yellowish brown, chartaceous with hyaline upper margins, oblong-lanceolate or oblong–elliptic, 3.5–5.5 mm long, 1.5–2.5 mm wide, laxly imbricate, suberect but those in upper part of spikelet weakly spreading, 7–nerved or 9–nerved, midnerve prolonged into a minute recurved mucro. Flowers bisexual (perfect). Corolla (petals) absent. Stamens 3.

Fruit: Achenes oblong–ellipsoid; trigonous; 1.7–2 mm long; ca. 0.8 mm wide. Seeds 1; seed coat thin; free from pericarp; embryo basal; endosperm abundant.

Ploidy:

Habitat: Occurring in mesic to wet forest.

Elevation Range: 350–1,280 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Specimens

Bibliography

Name Published In: Fl. Hawaiian Isl.: 468 (1888)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Cyperus hypochlorus Hillebr. PRESERVED_SPECIMEN o. degener 8384 MU 6/14/1927
2 Cyperus hypochlorus Hillebr. KAUAI: Waimea District; Makaha Ridge and Valley; north rim just before facility; Eragrostis Lowland Dry Shrub & Acacia- Metrosideros Mesic Forest w/ Dodonaea, Bidens, Artemesia, & Antidesma, Alphitonia, Coprosma, Carex, Schiedea. Major Threats: Erigeron karvinskianus, Pluchia symphytifolia, in forested area threats were Lantana camara, Rubus rosifol- ius, Psidium guajava. Elev. 1800 ft. PRESERVED_SPECIMEN K. R. Wood 2229 Kauai PTBG 1/2/1993
3 Cyperus hypochlorus Hillebr. Hanaulaiki, upper Papalaoa, W. Maui. In Metrosideros - Styphelia shrubland. Understory in thick shrub. PRESERVED_SPECIMEN K. R. Wood 9182 Maui PTBG 11/14/2001
4 Cyperus hypochlorus Hillebr. Kawaiiki Ridge. South facing slope just below the ridge. Diverse Mesic Forest PRESERVED_SPECIMEN Natalia Tangalin 3110 Kauai PTBG 4/13/2012
5 Cyperus hypochlorus Hillebr. Waimea Canyon, just SE of Kukui peak. Acacia-Metrosideros lowland dry to mesic forest PRESERVED_SPECIMEN K. R. Wood 16351 Kauai PTBG 3/5/2015
6 Cyperus hypochlorus Hillebr. Puu Kaua Region, Lualualei, steep ridge just to south of Kaua. PRESERVED_SPECIMEN K. R. Wood 10627 Oahu PTBG 3/30/2004
7 Cyperus hypochlorus Hillebr. PRESERVED_SPECIMEN o. degener 8450 Laysan US 4/24/1928
8 Cyperus hypochlorus Hillebr. Makaha Valley. Diospyros-Acacia-Metrosideros montane mesic forest. PRESERVED_SPECIMEN K. R. Wood 15889 Kauai PTBG 3/20/2014
9 Cyperus hypochlorus Hillebr. Kuia Natural Area Reserve, north facing slopes. Mixed mesic forest. PRESERVED_SPECIMEN K. R. Wood 6270 Kauai PTBG 5/10/1997
10 Cyperus hypochlorus Hillebr. waimea, s kohala PRESERVED_SPECIMEN e. y. hosaka 2048 Hawaii US 5/21/1938
11 Cyperus hypochlorus Hillebr. Waimea, S Kohala PRESERVED_SPECIMEN Hosaka, E. Y. 2048 Hawaii US 5/21/1938
12 Cyperus hypochlorus Hillebr. pohakea gulch, w maui PRESERVED_SPECIMEN o. degener 8398 Maui US 7/11/1927
13 Cyperus hypochlorus Hillebr. Waimea PRESERVED_SPECIMEN Hitchcock, A. S. 14389 Hawaii US 8/26/2016
14 Cyperus hypochlorus Hillebr. e slope of puu kaua PRESERVED_SPECIMEN o. degener 17568 Oahu US 1/19/1936
15 Cyperus hypochlorus Hillebr. Kawaihapai. PRESERVED_SPECIMEN Degener, O. 12872 Oahu US 9/1/1940
16 Cyperus hypochlorus Hillebr. waimea PRESERVED_SPECIMEN a. s. hitchcock 14389 Hawaii US 8/26/1916
17 Cyperus hypochlorus Hillebr. 17 mi from kohala toward waimea PRESERVED_SPECIMEN o. degener 2224 Hawaii US 8/14/1926
18 Cyperus hypochlorus Hillebr. At upper end of Hanalilolilo pipe-line PRESERVED_SPECIMEN Degener, O. 8450 Molokai US 4/24/2028
19 Cyperus hypochlorus Hillebr. niaupala fishpond, kaluaaha salt marsh PRESERVED_SPECIMEN h. st. john, r. baxter & l. william 19953 Molokai US 1/2/1939
20 Cyperus hypochlorus Hillebr. at upper end of hanalilolilo pipe-line PRESERVED_SPECIMEN o. degener 8450 Molokai US 4/24/1928
21 Cyperus hypochlorus Hillebr. Along Pipe-Line Trail, Olinda PRESERVED_SPECIMEN Degener, O. 8384 Maui US 6/14/2027
22 Cyperus hypochlorus Hillebr. 17 mi from Kohala toward Waimea PRESERVED_SPECIMEN Degener, O. 2224 Hawaii US 8/14/2026
23 Cyperus hypochlorus Hillebr. along pipe-line trail, olinda PRESERVED_SPECIMEN o. degener 8384 Maui US 6/14/1927
24 Cyperus hypochlorus Hillebr. Along the Hanapepe River, near the falls PRESERVED_SPECIMEN Heller, A. A. 2466 Kauai US 1895-00-00
25 Cyperus hypochlorus Hillebr. 17 mi from kohala toward waimea PRESERVED_SPECIMEN o. degener 2225 Hawaii US 8/1/1926
26 Cyperus hypochlorus Hillebr. zig-zag trail up cliff, kealia, waianae mountains PRESERVED_SPECIMEN f. r. fosberg 12861 Oahu US 2/2/1936
27 Cyperus hypochlorus Hillebr. E slope of Puu Kaua PRESERVED_SPECIMEN Degener, O. 17568 Oahu US 1/19/1936
28 Cyperus hypochlorus Hillebr. along the hanapepe river, near the falls PRESERVED_SPECIMEN a. a. heller 2466 Kauai US 1895-07-02
29 Cyperus hypochlorus Hillebr. palehua, waianae range PRESERVED_SPECIMEN o. degener 17623 Oahu US 11/23/1935
30 Cyperus hypochlorus Hillebr. Palehua, Waianae Range PRESERVED_SPECIMEN Degener, O. 17623 Oahu US 11/23/1935
31 Cyperus hypochlorus Hillebr. Hanalei Dist., Kalalau Rim, NW facing aspect, below Pihea Peak PRESERVED_SPECIMEN Wood, K. R. 1133 Kauai US 8/13/1991
32 Cyperus hypochlorus Hillebr. kokee camp PRESERVED_SPECIMEN o. degener 2253 Kauai US 6/23/1926
33 Cyperus hypochlorus Hillebr. kawaihapai. PRESERVED_SPECIMEN o. degener 12872 Oahu US 9/1/1940
34 Cyperus hypochlorus Hillebr. along pipe-line trail, olinda PRESERVED_SPECIMEN o. degener 8385 Oahu US 6/22/1927
35 Cyperus hypochlorus Hillebr. 17 mi from Kohala toward Waimea PRESERVED_SPECIMEN Degener, O. 2225 Hawaii US 8/1/2026
36 Cyperus hypochlorus Hillebr. Kokee Camp PRESERVED_SPECIMEN Degener, O. 2253 Kauai US 6/23/2026
37 Cyperus hypochlorus Hillebr. Niaupala Fishpond, Kaluaaha salt marsh PRESERVED_SPECIMEN St. John, H.; Baxter, R. W.; William, L. G. 19953 Molokai US 1/2/1939
38 Cyperus hypochlorus Hillebr. Along Pipe-Line Trail, Olinda PRESERVED_SPECIMEN Degener, O. 8385 Maui US 6/22/2027
39 Cyperus hypochlorus Hillebr. Waimea Drainage Basin, W side PRESERVED_SPECIMEN Forbes, C. N. 1008.K. Kauai US 1917-00-00
40 Cyperus hypochlorus Hillebr. Pohakea Gulch, W Maui PRESERVED_SPECIMEN Degener, O. 8398 Maui US 7/11/2027
41 Cyperus hypochlorus Hillebr. Waianae Kai. PRESERVED_SPECIMEN Wood, K. R. 4164 Oahu US 3/14/1995
42 Cyperus hypochlorus Hillebr. makaha ridge, off makaha rd In mesic forest. PRESERVED_SPECIMEN hobdy, r.w. collector number: 3587 Kauai BISH 3/23/1993
43 Cyperus hypochlorus Hillebr. hanalei dist., kalalau rim, nw facing aspect, below pihea peak PRESERVED_SPECIMEN k. r. wood 1133 Oahu US 8/13/1991
44 Cyperus hypochlorus Hillebr. Zig-zag trail up cliff, Kealia, Waianae Mountains PRESERVED_SPECIMEN Fosberg, F. R. 12861 Oahu US 2/2/1936
45 Cyperus hypochlorus Hillebr. Waimea District. Makaha, upper north fork, north facing slopes behind and east of kokio keokeo. Metrosideros-Acacia-Alphitonia montane mesic forest w/ Euphorbia atrococca, Myrsine lanaiensis, Dodonaea viscosa, Psychotria mariniana, Wikstroemia furcata, Diospyros sandwicensis, Kadua affinis, Coprosma waimeae, C. foliosa, Polyscias kavaiensis, Anti PRESERVED_SPECIMEN Wood, K.R. 16669 Kauai BISH 12/3/2015
46 Cyperus hypochlorus Hillebr. waianae kai. PRESERVED_SPECIMEN k. r. wood 4164 Oahu US 3/14/1995
47 Cyperus hypochlorus Hillebr. Waimea District. Kawaiiki Ridge. South facing slope just below the ridge. Diverse Mesic Forest W/Acacia koa w/Metrosideros with Zanthoxylum, Dodonea, Dubautia, Kadua affinis, Syzgium, or ridges. Open upper hillside w/Dicranopteris patches, open large Dianella patch w/ Cyperus and grasses. PRESERVED_SPECIMEN Tangalin, N. 3110 Kauai BISH 4/13/2012
48 Cyperus hypochlorus Hillebr. Keawapilau Gulch PRESERVED_SPECIMEN Welton, P. 631 Oahu BISH 1/21/1991
49 Cyperus hypochlorus Hillebr. waimea drainage basin, w side PRESERVED_SPECIMEN c. n. forbes 1008.k. Kauai US 7/3/1917
50 Cyperus hypochlorus Hillebr. Waimea District; Waialae Ridge and N side of ridge above falls Metrosideros-Acacia mixed mesic forest w/ Mariscus pennatiformis, Panicum nephelophilum, Diospyros, Charpentiera, Dianella, Dodonaea, Styphelia, Antidesma, Poa sandvicensis, Schiedea, Alsinidendron, endemic ferns are diverse in cool, shady gulchs. PRESERVED_SPECIMEN Wood, K.R. 2369 Kauai BISH 2/10/1993
51 Cyperus hypochlorus Hillebr. waimea district; waialae ridge and n side of ridge above falls Metrosideros-Acacia mixed mesic forest w/ Mariscus pennatiformis, Panicum nephelophilum, Diospyros, Charpentiera, Dianella, Dodonaea, Styphelia, Antidesma, Poa sandvicensis, Schiedea, Alsinidendron, endemic ferns are diverse in cool, shady gulchs. PRESERVED_SPECIMEN wood, k.r. perlman, s. collector number: 2369 Kauai BISH 2/10/1993
52 Cyperus hypochlorus Hillebr. kohua ridge north facing slopes above Awini Stream Metrosideros-Acacia montane mesic forest with Alphitonia pondersoa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena aurea, Polyscias kavaiensis, Planch PRESERVED_SPECIMEN wood, k.r. query, m. collector number: 14946 Kauai BISH 4/26/2012
53 Cyperus hypochlorus Hillebr. Makaha Ridge, off Makaha Rd In mesic forest. PRESERVED_SPECIMEN Hobdy, R.W. 3587 Kauai BISH 3/23/1993
54 Cyperus hypochlorus Hillebr. Ridgeline in vicinity of Puu Kawiwi, Waianae Kai Forest Reserve Occasional, under a low Acacia-Metrosideros canopy. PRESERVED_SPECIMEN Takeuchi, W.N. Waianae 17b Oahu BISH 12/11/1983
55 Cyperus hypochlorus Hillebr. keawapilau gulch PRESERVED_SPECIMEN welton, p. haus, b. collector number: 631 Oahu BISH 1/21/1991
56 Cyperus hypochlorus Hillebr. olinda, along pipe-line trail Uncommon along trail in lowest bog, forming tussocks ca. 30 cm across PRESERVED_SPECIMEN degener, o. 8384 DUKE 6/14/1927
57 Cyperus hypochlorus Hillebr. ridgeline in vicinity of puu kawiwi, waianae kai forest reserve Occasional, under a low Acacia-Metrosideros canopy. PRESERVED_SPECIMEN takeuchi, w.n. wagner, w.l.; yee, r.s.n.; obata, j. collector number: waianae_17b Oahu BISH 12/11/1983
58 Cyperus hypochlorus Hillebr. kokee, kauai PRESERVED_SPECIMEN morr lee; moir & barnum 5653 CM 11/22/1925